A survey of Pireneitega from Tajikistan (Agelenidae, Coelotinae).

Five new species of Pireneitega species from Tajikistan are described: Pireneitega zonsteinisp. n. (♂♀), Pireneitega muratovisp. n. (♀), Pireneitega tyuraisp. n. (♀), Pireneitega ramitensissp. n. (♀) and Pireneitega kovblyukisp. n. (♂). Pireneitega major (Kroneberg, 1875) is redescribed for the first time based on the lectotype designated here. DNA barcodes for the five new species are documented for future use and as proof of molecular differences between these species.

Introduction

is the largest subfamily of with more than 650 species distributed in the Holarctic and southeast Asia (World Spider Catalog 2016). Kishida, 1955 with 30 species distributed across the Palaearctic (World Spider Catalog 2016, Zhang et al. 2016) is one of the most species-rich genera of the subfamily. It is relatively well studied in comparison to other species-rich (and non-monophyletic) genera Blackwall, 1841 and Ovtchinnikov, 1999. The species PageBreakof found in Caucasus and Xinjiang were recently revised (Kovblyuk et al. 2013; Zhang et al. 2016) but the genus remains poorly studied in Central Asia. Of three species known from central Asia (Mikhailov 2013: (Ermolajev, 1927) (currently considered a junior synonym of (L. Koch, 1878)), (Charitonov, 1946) and (Kroneberg, 1875)), is known only from the original description and only from two very short descriptions supplied with sketchy figures. A short trip by the junior author to Tajikistan revealed five new morphospecies of , each separated by distinct genetic gaps. The goal of this paper is to provide descriptions of the new species (including records of their molecular markers) and a redescription of whose type locality lies in northern Tajikistan.

Material and methods

Specimens were examined and measured with a Leica M205C stereomicroscope. Images were captured with an Olympus C7070 wide zoom digital camera (7.1 megapixels) mounted on an Olympus SZX12 dissecting microscope. Epigynes and male palps were examined after dissection. Epigynes were cleared by boiling it in 10% KOH solution before taking photos of the dorsal view. All measurements are given in millimeters. was photographed and drawn using an MBS-9 stereomicroscope with Pro-MicroScancamera. Leg measurements are given as: total length (femur, patella + tibia, metatarsus, tarsus).

Terminology used for copulatory organ characters in the text and figure legends follows Wang (2002) with some modifications.

Abbreviations used in the text and figure legends are:

A

epigynal atrium

ALE

anterior lateral eye

AME

anterior median eye

AME-ALE

distance between AME and ALE

AME-AME

distance between AME and AME

ALE-PLE

distance between ALE and PLE

CD

copulatory ducts

CF

cymbial furrow

CO

conductor

d

dorsal

E

embolus

EB

embolic base

ET

epigynal teeth

FD

fertilization ducts

Fe

femur

H

epigynal hood

MA

median apophysis

Mt

metatarsus

p

prolateral

PA

patellar apophysis

Pa

patella

PLE

posterior lateral eye

PME

posterior median eye

PME-PLE

distance between PME and PLE

PME-PME

distance between PME and PME

R

receptacle

r

retrolateral

RTA

retrolateral tibial apophysis

ST

subtegulum

T

tegulum

Ta

tarsus

Ti

tibia

v

ventral

VTA

ventral tibial apophysis

DNA barcodes were obtained for future use: a partial fragment of the mitochondrial gene cytochrome oxidase subunit I (COI) was amplified and sequenced for five new species using Primers LCO1490-oono (5’-CWACAAAYCATARRGATATTGG-3’) (Folmer et al. 1994; Miller et al. 2010) and HCO2198-zz (5’-TAAACTTCCAGGTGACCAAAAAATCA-3’) (Folmer et al. 1994; Zhao and Li 2016). For additional information on extraction, amplification, and sequencing procedures, see Zhao et al. (2013). All sequences were blasted in GenBank; accession numbers are provided in Table 1.

Table 1.

Voucher specimen information.

Species	GenBank accession number	Sequence length	Collection localities
Pireneitega zonsteini sp. n.	KY024475	642bp	Env. of Dushanbe, Hissar Mt. Ridge 48th km of Varzob Hwy
Pireneitega muratovi sp. n.	KY024477	642bp	Env. of Dushanbe Hissar, Mt. Ridge 20th km of Varzob Hwy Gusgarf Vill.
Pireneitega tyurai sp. n.	KY024478	642bp	Khatlon Area Khovaling Distr., Obimazar River
Pireneitega ramitensis sp. n.	KY024476	642bp	Khatlon Area Hissar Mt. Range Ramit Reserve
Pireneitega kovblyuki sp. n.	KY024474	642bp	Tajikstan: Khatlon Area Dangara Distr Sanglogh

Holotypes and some paratypes will be deposited in the . Most paratypes are deposited in the in Beijing, China.

Zoological Museum of the Moscow State University

Institute of Zoology, Chinese Academy of Sciences

Taxonomy

Kishida, 1955

Kishida, 1955: 11. Type species

Brignoli, 1982: 347. Type species

Note.

was long considered a nomen nudum (Yaginuma, in Brignoli 1983: p. 468). Kishida (1955), in a general survey of , considered to have been described by himself in 1928, although he had no publications that year. The genus " Kishida, 1928 [Genotype: roscida (Koch, 1868)]" was considered among the tribe Kishida, 1928 (Kishida 1955: p. 11). Although eye pattern was mentioned in the key to the genera of "", Kishida (1955) did not provide a formal description of the genus. Brignoli (1982) described (type species Brignoli, 1978) from Turkey. Subsequently, Wang and Jäger (2007) revalidated with as a junior synonym.

Diagnosis.

The chelicerae in most species of (including the type species) have 3 promarginal and 3 retromarginal teeth; other coelotines have either 2 or 4 retromarginal teeth (Zhang et al. 2016). The females can be distinguished by the widely separated epigynal teeth, the large atrium with subparallel margins, and the broad copulatory ducts (Fig. 2A–B); other coelotines usually have a small atrium and copulatory ducts. The males can be distinguished by the absence of a dorsal “apophysis” on the conductor, the small RTA, and the distinct median apophysis (Fig. 1A–C); other coelotines usually have a broad dorsal apophysis on the conductor and a reduced or indistinct median apophysis.

Figure 2.

sp. n., female paratype and male holotype. A Epigyne, ventral B Vulva, dorsal C Male habitus, dorsal D Female habitus, dorsal E Female habitus, ventral. Scale bars equal for D, E.

Figure 1.

Male palp of sp. n., holotype. A Prolateral B Ventral C Retrolateral. Scale bar 0.1 mm.

Composition.

Thirty species of are known from Spain to Sakhalin (World Spider Catalog 2016; Mikhailov 2013). One species, , was known from Tajikistan before the current study.

sp. n.

http://zoobank.org/1AF265B6-AAB0-4974-A8A7-94A906F8FBBF

Figs 1 , 2 , 8

Figure 8.

Localities of species from Tajikistan. 1 sp. n. 2 sp. n. 3 sp. n. 4 sp. n. 5 sp. n. 6 .

Type material.

Holotype ♂ (ZMMU): Tajikstan, environs of Dushanbe, Hissar Mt. Range, 48th km of Varzob Hwy, S exposed slope with litter & under stones, , 1530 m, 7.05.2015 (Y.M. Marusik, M. Saidov). Paratypes: 1♂1♀ (IZCAS), same data as holotype.

Etymology.

The species is named after Sergei Zonstein (University of Tel-Aviv, Israel) a partner of the junior author in the expedition to Tajikistan; noun (name) in genitive case.

The male can be distinguished from all other species except (Wang et al., 1990) by having a broad conductor and thick patellar apophysis. From it is distinguished by the blunt tip of the patellar apophysis (vs a tapering tip in ) (Fig. 1; Wang et al. 1990: figs 13–15). The female can be distinguished from all other species except by having a nearly trapezoidal atrium, long copulatory ducts, and short receptacles. From it can be distinguished by its short epigynal teeth, about 0.5 times as long as length of the atrium (vs long epigynal teeth in , about as long as the length of the atrium) (Fig. 2; Charitonov 1946: fig. 4).

Description.

Male (holotype): Total length 8.90. Carapace 4.40 long, 3.50 wide. Abdomen 4.50 long, 2.80 wide. Eye sizes and interdistances: AME 0.15, ALE 0.20, PME 0.15, PLE 0.20; AME-AME 0.07, AME-ALE 0.06, PME-PME 0.15, PME-PLE 0.18. Leg measurements: I: 12.95 (3.50, 4.30, 3.15, 2.00); II: 12.25 (3.25, 4.00, 3.00, 2.00); III: 10.40 (3.15, 3.00, 3.25, 1.00); IV: 16.00 (4.50, 5.00, 4.25, 2.25). Carapace greenish, the radial grooves indistinct, with black lateral margins. Abdomen blackish, with yellow herringbone pattern.

Spination in male

Palp as in Fig. 1: patellar apophysis long, more than half length of tibia; tibia short, about 1/4 length of tarsus; VTA subequal to the tibial length, without pointed tip, extending beyond the tibia; RTA short, about 1/6 length of VTA; cymbial furrow long, more than half length of cymbium; conductor broad and with two spiraling loops; median apophysis broad and nearly triangular; embolus with broad base originating proximally on base of tegulum.

Female (paratype): Total length 10.0. Carapace 4.75 long, 3.65 wide. Abdomen 5.25 long, 3.45 wide. Eye sizes and interdistances: AME 0.20, ALE 0.25, PME 0.21, PLE 0.26; AME-AME 0.08, AME-ALE 0.05, PME-PME 0.17, PME-PLE 0.20. Leg measurements: I: 12.50 (3.75, 4.25, 2.75, 1.75); II: 11.75 (3.50, 4.00, 2.75, 1.50); III: 10.60 (3.00, 3.50, 2.60, 1.50); IV: 15.00 (4.25, 4.75, 4.00, 2.00). Carapace yellow. Abdomen black, with yellow spots and herringbone pattern.

Epigyne as in Fig. 2A–B: epigynal teeth narrow and relatively short (shorter than width of atrium); septum short with weakly sclerotized tip, about 0.3 times as long as wide; atrium with well delimited posterior margin, about 1.3 times longer than wide, about 4 times longer than septum, subequal to width of septum; copulatory opening hidden by anterior margin of atrium; receptacles long, about 2 times longer than wide, separated by 2.5 times their diameters; copulatory ducts with 3 parts, the basal part running from receptacle posteriorly (Bd), median part running anteriorly (Md), and terminal part (Td) running posteriorly and leading to copulatory opening; median part as wide as terminal and 2 times longer than basal part; median part 1.5 times longer than receptacle; median parts touching each other; hoods indistinct.

Spination in female

Distribution.

Known only from the type locality (Fig. 8).

http://zoobank.org/A01FC654-273B-4E50-A278-052B957FBA4B

Figs 3 , 8

Figure 3.

sp. n., female holotype. A Epigyne, ventral B Vulva, dorsal C Habitus, dorsal D Habitus, dorsal E Habitus, ventral view. Scale bars equal for C, D, E.

Holotype ♀ (ZMMU): Tajikstan: env of Dushanbe, Hissar Mt. Ridge, 20th km of Varzob Hwy, Gusgarf [Gushharf] Vill., N exposed slope with litter & cliffs, , 1750 m, 8.05.2015, Y. M. Marusik. Paratype: 1♀ (IZCAS), same data as holotype.

The species is named after Tajikistan zoologist Rustam Muratov (Dushanbe, Tajikistan) who was very helpful in organizing the expedition to Tajikistan; noun (name) in genitive case.

The female can be distinguished from all other species except , (Zhu & Wang, 1994), and by having narrow epigynal teeth and an elongate oval atrium. It can be distinguished from by the pointed tip of septum (vs blunt tip in ), from by the elongate oval receptacles (vs spiralled in ), and from by its short epigynal teeth, ca. 0.8 times as long as length of the atrium (vs long epigynal teeth in , about as long as the length of the atrium) (Figs 3, 7; Charitonov 1946: fig. 4; Zhu and Wang 1994: figs 5–6).

Figure 7.

Epigyne of , lectotype. A Ventral B Dorsal.

Male: unknown.

Female (holotype): Total length 9.94. Carapace 4.49 long, 3.05 wide. Abdomen 5.45 long, 2.90 wide. Eye sizes and interdistances: AME 0.18, ALE 0.23, PME 0.24, PLE 0.30; AME-AME 0.10, AME-ALE 0.05, PME-PME 0.15, PME-PLE 0.10. Leg measurements: I: 11.25 (3.25, 4.00, 2.50, 1.50); II: 10.30 (3.00, 3.50, 2.50, 1.30); III: 9.70 (2.75, 3.00, 2.65, 1.30); IV: 13.75 (3.75, 4.25, 4.00, 1.75). Carapace yellow, the radial grooves indistinct. Abdomen whitish-yellow, with green herringbone pattern.

Epigyne as in Fig. 3A–B: epigynal teeth narrow, their length equal to width of the narrowest part of the atrium; septum with well delimited tip, ca. 0.5 times as long as wide; copulatory opening distinct; atrium with well delimited posterior margin, about 1.4 times longer than wide, ca. 2 times longer than and 0.7 times as wide as septum; receptacles long, about 2.5 times as long as wide, bases of receptacles separated by 2 diameters; copulatory ducts with 3 parts, median part as long as receptacles, and anterior part slightly wider than receptacles; hoods indistinct.

Spination

Known only from the type locality (Fig. 8).

http://zoobank.org/B14F37A9-6A33-446F-80AF-2C65472362D3

Figs 4 , 8

Figure 4.

sp. n., female holotype. A Epigyne, ventral B Vulva, dorsal C Habitus, dorsal D Habitus, dorsal E Habitus, ventral. Scale bars equal for C, D, E.

Holotype ♀ (ZMMU): Tajikstan: Khatlon Area, Khovaling Distr., Obimazar River, Sultan-Mazar, clay cliffs, , 1854 m, 27.04.2015 (Y.M. Marusik). Paratypes: 4♀ (IZCAS), same data as holotype.

The species is named after Sergei V. Tyura (Magadan, Russia) a friend of the junior author; noun (name) in genitive case.

The female can be distinguished from all other species except (Wang et al., 1990) by having short receptacles and copulatory ducts. It can be distinguished from by the broad and long epigynal teeth, about 0.85 times as long as atrium (vs short and narrow epigynal teeth in , about 0.5 times as long as atrium) (Fig. 4A–B; Wang et al. 1990: figs 84–85).

Male: unknown.

Female (holotype): Total length 5.15. Carapace 2.15 long, 1.75 wide. Abdomen 3.00 long, 2.00 wide. Eye sizes and interdistances: AME 0.10, ALE 0.13, PME 0.15, PLE 0.15; AME-AME 0.05, AME-ALE 0.10, PME-PME 0.02, PME-PLE 0.04. Leg measurements: I: 6.20 (1.90, 2.25, 1.25, 0.80); II: 5.10 (1.60, 1.75, 1.00, 0.75); III: 4.80 (1.50, 1.60, 1.00, 0.70); IV: 7.05 (2.05, 2.50, 1.50, 1.00). Carapace yellow, with black lateral margins. Abdomen blackish, with yellow herringbone pattern.

Epigyne as in Fig. 4A–B: epigynal teeth long (nearly as long as atrium); septum with weakly sclerotized tip, about 0.5 times as long as wide; atrium with weakly sclerotized posterior margin, about 0.7 times as long as wide, about 1.8 times longer than and 0.7 times as wide as septum; copulatory opening hidden; receptacles large, ca. 2 times longer than wide; copulatory ducts with two parts, terminal parts (Tp) not touching each other, about 0.5 length of receptacles, basal parts (Bp) shorter than width of receptacle; hoods indistinct.

Spination

Known only from the type locality (Fig. 8).

http://zoobank.org/C74C6BAE-DE7C-4A95-A4A2-5E5BFC45C341

Figs 5 , 8

Figure 5.

sp. n., female holotype. A Epigyne, ventral B Vulva, dorsal C Habitus, dorsal D Habitus, ventral E Habitus, ventral. Scale bars equal for C, D, E.

Holotype ♀ (ZMMU): Tajikstan: Khatlon Area, Hissar Mt. Range, Ramit Reserve, , 1324 m, 1.05.2015 (Y.M. Marusik). Paratypes: 4♀ (IZCAS), 2♀ (ZMMU), same data as holotype.

The specific name is an adjective and refers to the type locality; adjective.

The female can be distinguished from all other species except sp. n., , and , by having an elongate oval atrium, narrow epigynal teeth, and long copulatory ducts. It can be distinguished from sp. n. and by the narrow tip of the copulatory ducts (vs round tip in sp. n. and ) and from and by the bent epigynal teeth (vs straight epigynal teeth in and ) (Figs 3, 5, 7; Charitonov 1946: fig. 4; Zhu and Wang 1994: figs 5–6).

Male: unknown.

Female (holotype): Total length 12.00. Carapace 4.50 long, 3.55 wide. Abdomen 7.50 long, 4.75 wide. Eye sizes and interdistances: AME 0.20, ALE 0.23, PME 0.25, PLE 0.20; AME-AME 0.10, AME-ALE 0.20, PME-PME 0.10, PME-PLE 0.23. Leg measurements: I: 14.05 (4.00, 4.75, 3.45, 1.85); II: 13.40 (3.90, 4.50, 3.25, 1.75); III: 13.00 (3.75, 4.25, 3.25, 1.75); IV: 16.55 (4.75, 5.40, 4.40, 2.00). Carapace yellowish, with brown lateral margins. Abdomen pale-yellow, with brown spots.

Epigyne as in Fig. 5A–B: epigynal teeth pale, hyaline, long and thin, about 0.9 times as long as receptacles; septum with weakly sclerotized tip, ca. 0.5 times as long as wide, nearly triangular; copulatory ducts distinct; atrium about 1.4 times longer than wide, with well delimited posterior margin, ca. 2.8 times longer than and about as wide as septum; receptacles large, about. 3 times longer than wide; receptacle bases separated by about 2 diameters; copulatory ducts with 3 parts, basal part about 2/3 of receptacle length, median part as long as receptacle, terminal part somewhat shorter than median part; hoods distinct.

Spination

Known only from the type locality (Fig. 8).

http://zoobank.org/25787234-B768-4EB3-B6B2-781E025AB5D4

Figs 6 , 8

Figure 6.

Male palp of sp. n., holotype. A Prolateral B Ventral C Retrolateral. Scale bar 0.1 mm.

Holotype ♂ (ZMMU): Tajikstan, Khatlon Area, Dangara Distr., Sanglogh (=Sanglok) Mt. Range above Shar-Shar Pass, , 1700–2060 m, 29.04.2015, (Y.M. Marusik). Paratypes: 3♂ (IZCAS), 2♂ (ZMMU), same data as holotype.

The specific name is a patronym in honour of the well known arachnologist and friend of the junior author Mykola M. Kovblyuk (Simferopol, Ukraine); noun (name) in genitive case.

The male can be distinguished from all other species except by having a hook-shaped conductor, and narrow cymbium. It can be distinguished from by the short cymbial furrow, ca. 1/10 length of cymbium (vs long cymbial furrow in , about 0.5 length of cymbium) (Fig. 6; Wang et al. 1990: figs 81–83).

Male (holotype): Total length 7.90. Carapace 4.00 long, 3.00 wide. Abdomen 3.90 long, 2.65 wide. Eye sizes and interdistances: AME 0.15, ALE 0.20, PME 0.18, PLE 0.19; AME-AME 0.08, AME-ALE 0.07, PME-PME 0.13, PME-PLE 0.15. Leg measurements: I: 10.90 (3.25, 4.05, 2.00, 1.60); II: 9.85 (3.00, 3.50, 2.00, PageBreak1.35); III: 8.60 (2.75, 2.50, 2.10, 1.25); IV: 12.55 (3.70, 3.75, 3.50, 1.60). Carapace yellow, the radial grooves indistinct. Abdomen pale, with yellow herringbone pattern.

Palp as in Fig. 6A–C: patellar apophysis absent; tibia long, ca. 0.5 length of cymbium; VTA short and wide, about 1/3 length of tibia, without pointed tip; RTA short, about 1/5 length of VTA, poorly visible; cymbium long, its tip as long as or longer than genital bulb; conductor short, with hook-shaped, partially looped tip, tip located distally from tegulum; median apophysis broad and nearly triangular; embolus with broad, nearly tongue-shaped base, beginning at 6:30 o’clock position.

Spination

Female: Unknown.

Known only from the type locality (Fig. 8).

(Kroneberg, 1875)

Figs 7 , 8

Kroneberg, 1875: 15, pl. 1, fig. 6 (♀); Charitonov, 1946: 20, fig. 5 (♀).

: Ovtchinnikov, 1988: 142 (transferred from

:

:

Material examined.

Lectotype ♀ (ZMMU) with label «Ta 3845 1♀ ZMMU [Зоомузей МГУ]» «Lectotypus» 2/VI; Аyчи дагана [Auchi dagana] Kroneberg, 1875», ca . Paralectotype: 1♀ (ZMMU) with 2 labels «Ta1059, 1, » «Туркестанская Учёная Экспедиция Имераторскаго Общества Любителей Естествознанiя. Федченко [Turkestan Scientific Expedition of the Emperor’s Society of Devotees of Natural Sciences. Fedchenko]» and « n. sp. Ta, No.1059, Кокандское ханство, Федченко [Kokand Khanate, Fedchenko]».

Comments.

The figures of presented by Schenkel (1936), Hu and Wu (1989), and Song et al. (1999; copied from Hu and Wu 1989) are of a species other than , the identity of which is currently unknown. All records of this unknown species are from Xinjiang, China.

This species is easily distinguished from other species of found in Tajikistan by its larger size (carapace length >6 mm vs <4.75) and having 5 spines on tarsus IV (vs other species with 0‒4). The epigyne of is most similar to that of sp. n. and sp. n. It can be distinguished from sp. n. by its shorter receptacles with length/width ratio of 2.3 (vs 2.6 in ), shape of copulatory ducts, and shorter teeth (cf. Figs 3A–B and 7A–B). can be separated from sp. n by its wider epigynal atrium and shorter, wider receptacles as well as by its shorter and wider copulatory ducts (cf. Figs 5A–B and 7A–B).

Male: unknown.

Female: Lectotype. Total length 16.7. Carapace 7.0 long, 5.0 wide, fovea 1.25 long. Leg measurements: I:19.75 (5.5, 2.5, 4.6, 4.65, 2.5); II: 18.6 (5.1, 2.5, 4.0, 4.5, 2.5); III: 17.2 (4.75, 2.2, 3.55, 4.6, 2.1); IV: 21.85 (5.75, 2.3, 5.0, 6.25, 2.55).

Spination

Paralectotype ♀. Total length: 11.0. Carapace 6.0 long, 4.0 wide. Epigyne 0.51 wide.

Epigyne as in Fig. 7: epigynal teeth pale, hyaline, long and thin; septum with weakly sclerotized tip, about 0.4 times as long as wide, subtriangular; atrium as long as wide; receptacles large, about 2.5 times longer than wide; receptacle bases separated by ca. 2 diameters; copulatory ducts with 2 parts, basal part as long as receptacle, terminal part somewhat shorter receptacle.

Known from the type series females only. Exact locality is known for the lectotype only: Auchi lies on the northern macroslope of the Turkestan Mt Range (Fig. 8).

sp. n. Spination in male

	Fe	Pt	Ti	Mt	Ta
I	3d 2p 1r	–	3-3v	3-3v	–
II	3d 1p 1r	–	2p 3-3v	2p 3-3v	–
III	3d 2p 2r	1p 1r	1d 2p 2r 3-3v	2p 2r 3-3v	–
IV	3d 2p 1r	1p 1r	2p 2r 3-3v	2p 2r 3-3v	–

sp. n. Spination in female

	Fe	Pt	Ti	Mt	Ta
I	3d 2p 1r	–	3-3v	3-3v	–
II	3d 1p 1r	–	1p 3-3v	1p 3-3v	–
III	3d 1p 2r	1p 1r	2p 2r 3-3v	2p 2r 3-3v	–
IV	3d 1p 1r	1p 1r	2p 2r 3-3v	1p 2r 3-3v	–

sp. n. Spination

	Fe	Pt	Ti	Mt	Ta
I	3d 2p 1r	–	3-3v	1p 3-3v	–
II	3d 3p 2r	–	2p 3-3v	3-3v	–
III	3d 3p 2r	1p 1r	2p 2r 3-3v	5p 4r 3-3v	1p 1r
IV	3d 1p 1r	1d 1p 1r	1d 2p 2r 3-3v	5p 5r 3-3v	2p 1r

sp. n. Spination

	Fe	Pt	Ti	Mt	Ta
I	3d 2p	–	3-3v	3-3v	–
II	3d 1p 1r	1p	2p 3-3v	1p 3-3v	–
III	3d 1p 1r	1p 1r	2p 2r 3-3v	5p 4r 3-3v	2p 1r
IV	2d 1p 1r	1p 1r	2p 2r 3-3v	5p 4r 3-3v	2p 1r

sp. n. Spination

	Fe	Pt	Ti	Mt	Ta
I	3d 2p 1r	–	1p 3-3v	1p 3-3v	–
II	3d 2p 2r	–	2p 3-3v	2p 3-3v	–
III	3d 3p 2r	1p 1r	2p 2r 3-3v	5p 4r 3-3v	2p 2r
IV	3d 2p 1r	1p 1r	2p 2r 3-3v	5p 4r 3-3v	2p 2r

sp. n. Spination

	Fe	Pt	Ti	Mt	Ta
I	3d 2p 1r	–	3-3v	3-3v	–
II	3d 3p 1r	1p	2p 3-3v	3p 3-3v	–
III	3d 2p 2r	1d1p 1r	1d 2p 2r 3-3v	5p 5r 3-3v	1p 1r
IV	3d 1p 1r	1p 1r	2p 2r 3-3v	5p 5r 3-3v	2p 1r

(Kroneberg, 1875) Spination

	Fe	Pt	Ti	Mt	Ta
I	3d 2p 2r	–	3-3v	3-3v 1vm	–
II	3d 3p 2r	–	2p 3-3v	1p 3-3v	–
III	3d 3p 2r	1p 1(0)r	2p 2r 3-3v	5p 4r 3-3v	2p 1-1v
IV	3d 2p 1r	1p 1r	2p 2r 3-3v	5p 4r 3-3v	2p 3r