The bees of the family Halictidae (Hymenoptera) described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko.

The type specimens of the family Halictidae, described by Ferdinand Morawitz from the collection of Aleksey Fedtschenko deposited in the Zoological Museum of the Moscow State University and in the Zoological Institute, Russian Academy of Sciences, St. Petersburg (Russia), are critically reviewed. Precise information with illustrations of types for 43 taxa is provided. Lectotypes are here designated for the following seven nominal taxa: Halictus aprilinus Morawitz, 1876, H. cingulatus Morawitz, 1876, H. laevinodis Morawitz, 1876, H. limbellus Morawitz, 1876, H. nasica Morawitz, 1876, H. rhynchites Morawitz, 1876 and H. vulgaris Morawitz, 1876. Yulia V. Astafurova, Maksim Yu. Proshchalykin.

Introduction

More than 140 years ago (1876), the second part of Ferdinand Morawitz’s critical study on the bees collected by the Aleksey Fedtschenko 1869–1871 Expeditions in “Turkestan” was published. In the prior volume, “ genuinae” (1875), Morawitz treated a total of 255 species of numerous genera, of which many species were described as new. In this second part, “” (1876), the remaining bees were dealt with, including the species of the difficult genera , and , totalling 183 species (Pesenko and Astafurova 2003). The species treatments are of a high professional standard, the localities are precisely documented (A. Fedtschenko 1871, O. Fedtschenko 1874, Baker 2004, Kuhlmann 2005, Dathe and Proshchalykin 2017) and the type series have been carefully conserved over a long period, generally in the collections of the Zoological Museum of the Moscow State University, Moscow, Russia (ZMMU) and in the Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia (ZISP). To this day, these remain some of the most important manuscripts on bees of this region.

The entomological literature uses various, often obscure, terms and names for Central Asian regions and countries. The term “Turkestan” has a particularly special use in entomology, widely adopted by Morawitz (1875, 1876), Dalla Torre (1896), Meade-Waldo (1923) and in other fundamental papers, regardless of its imprecise assignment to countries (Proshchalykin and Dathe 2018).

The territory of Central Asia, described as the “Western Regions” (Xi Yui) in Chinese sources, was referred to in the Russian and European historiography of the 18th and early 19th Centuries as Lesser Bukharia, as opposed to Greater Bukharia, where the Bukhara Khanate was situated. In Europe, these lands came to be referred to in the 18th and 19th Centuries as Turkestan, i.e. “the Land of Turks,” which was the original Iranian name for the territory east of Fergana and Bukhara where nomadic Turkic tribes roamed. Subsequently, when the Turkic tribes occupied the enormous territory from the Caspian Sea to Lop Nor, the name Turkestan acquired a new meaning, so broad that it was deemed necessary to distinguish such areas as Western—Bukhara, or Russian Turkestan – and Eastern or Chinese Turkestan (Murzayev 1957). According to current views, Fedtschenko’s “Turkestan” comprises the countries Uzbekistan, Tajikistan, Kyrgyzstan and southern parts of Kazakhstan.

The family is represented in Morawitz’s 1876 publication by four genera ( Schenck, 1866, Latreille, 1804, Latreille, 1804 and Latreille, 1804), comprising 73 species (Nos. 327–399). Only 30 species were previously known, while the remaining 43 were newly described ( – 2 species; – 36, – 2, – 2) (Table 1). Few of these taxa were mentioned in subsequent publications, remaining enigmatic for decades. Since the 1930s, V. Popov [ZISP] based his studies on the taxonomy and ecology of the Central Asian bee fauna using these collections. Popov involved other mellitologists in his research, so that specimens also reached specialists in other European museums. For example, P. Blüthgen took part in such work on and published several taxonomic papers, based on the Fedtschenko collection (Blüthgen 1931a, 1934a, 1955, Popov 1935).

Table 1.

The nominal taxa of Fedtschenko’s , described by F. Morawitz, and their current status.

N	Species name	Sex	Current status	Depositaries of types
1.	Halictus albitarsis	♀	Homonym	LT (ZMMU); PLT (ZISP/ZMMU)
2.	Halictus annulipes	♀	Valid	LT (ZMMU); PLT (ZISP/ZMMU)
3.	Halictus aprilinus	♀	Valid	LT, PLT (ZMMU)
4.	Halictus atomarius	♀	Valid	LT, PLT (ZISP)
5.	Halictus cariniventris	♂	Valid	LT, PLT (ZMMU)
6.	Halictus cingulatus	♀	Valid	LT (ZMMU); PLT (ZISP/ZMMU)
7.	Halictus croceipes	♀, ♂	Valid	LT (ZMMU); PLT (ZISP/ZMMU)
8.	Halictus desertorum	♀	Valid	LT (ZMMU)
9.	Halictus determinatus	♀	Homonym	LT (ZMMU); PLT (ZISP/ZMMU)
10.	Halictus equestris	♀	Valid	LT (ZMMU)
11.	Halictus ferghanicus	♂	Synonym	LT (ZMMU); PLT (ZISP)
12.	Halictus fucosus	♂	Synonym	HT (ZMMU)
13.	Halictus fulvitarsis	♂	Valid	HT (ZMMU)
14.	Halictus funerarius	♀	Valid	LT (ZISP); PLT (ZISP/ZMMU)
15.	Halictus fuscicollis	♀	Valid	LT (ZMMU); PLT (ZISP)
16.	Halictus hyalinipennis	♀, ♂	Valid	LT (ZISP); PLT (ZISP/ZMMU)
17.	Halictus laevinodis	♀	Valid	LT (ZISP); PLT (ZMMU)
18.	Halictus limbellus	♀	Valid	LT (ZMMU); PLT (ZISP/ZMMU)
19.	Halictus longirostris	♀, ♂	Valid	LT (ZMMU); PLT (ZISP/ZMMU)
20.	Halictus maculipes	♀	Valid	LT (ZMMU)
21.	Halictus melanarius	♂	Valid	HT (ZMMU)
22.	Halictus minor	♀	Valid	LT (ZISP); PLT (ZISP/ZMMU)
23.	Halictus modernus	♀	Valid	HT (ZMMU)
24.	Halictus nasica	♀, ♂	Valid	LT (ZMMU); PLT (ZISP/ZMMU)
25.	Halictus nigrilabris	♂	Valid	LT (ZMMU); PLT (ZISP)
26.	Halictus nigripes	♂	Homonym	LT (ZMMU); PLT (ZISP/ZMMU)
27.	Halictus obscuratus	♀	Valid	LT (ZMMU); PLT (ZISP/ZMMU)
28.	Halictus palustris	♀	Valid	LT (ZMMU); PLT (ZISP)
29.	Halictus pectoralis	♀	Homonym	HT (ZMMU)
30.	Halictus picipes	♀	Valid	LT (ZMMU); PLT (ZISP/ZMMU)
31.	Halictus rhynchites	♀, ♂	Valid	LT (ZMMU); PLT (ZISP/ZMMU)
32.	Halictus scutellaris	♀	Valid	LT (ZMMU); PLT (ZISP/ZMMU)
33.	Halictus sogdianus	♀	Synonym	LT, PLT (ZMMU)
34.	Halictus trifasciatus	♀	Homonym	LT (ZMMU)
35.	Halictus varipes	♀, ♂	Synonym	LT, PLT (ZMMU)
36.	Halictus vulgaris	♀	Synonym	LT (ZMMU); PLT (ZISP/ZMMU)
37.	Sphecodes nigripennis	♀	Synonym	LT, PLT (ZMMU)
38.	Sphecodes pectoralis	♀	Valid	LT, PLT (ZMMU)
39.	Sphecodes rufithorax	♀, ♂	Synonym	LT (ZMMU); PLT (ZISP/ZMMU)
40.	Nomia edentata	♀, ♂	Valid	LT (ZMMU); PLT (ZISP/ZMMU)
41.	Nomia rufescens	♀	Valid	LT (ZMMU)
42.	Nomioides parviceps	♂	Valid	NT (ZISP)
43.	Nomioides turanica	♀, ♂	Valid	LT, PLT (ZISP/ZMMU)

Comment. LT – lectotype, PLT – paralectotype/s, NT – neotype.

The second attempt to recognise the true type material of Fedtschenko’s was made by K. Warncke, a teacher in Dachau (Germany), who visited ZMMU from 26.03.1975 to 01.04.1975 (Dathe and Proshchalykin 2017). He worked his way through the drawers containing the Fedtschenko material and labelled specimens of all nominal taxa found there as “Lectotypen.” Thereby, certain specimens from some groups of bees (, , etc.) received a red label with the inscription “Lectotypus Warncke 1975.” However, of all Fedtschenko’s described by Morawitz, only five specimens received such labels: Morawitz, 1876, Morawitz, 1876, Morawitz, 1876, Morawitz, 1876 and Morawitz, 1876 (Figs 32A, 33B, 34E, 35F, 36A). All other type specimens were left without nomenclatural status labels.

Figure 32.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 2.0 mm (A, D), 1.0 mm (B, C).

Figure 33.

Morawitz, 1876, lectotype, female A head, frontal view B habitus, lateral view and labels C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 2.0 mm (B, D), 1.0 mm (A, C).

Figure 34.

Morawitz, 1876, lectotype, female A habitus, lateral view B head, frontal view C metasoma, dorsal view D vertex, dorso-lateral view E labels F mesosoma, dorsal view. Scale bars: 1.0 mm (A–C, F), 0.5 mm (D).

Figure 35.

Morawitz, 1876, lectotype, female A habitus, lateral view B head, frontal view C metapostnotum, dorsal view D metasoma, dorsal view E mesosoma, dorsal view F labels. Scale bars: 1.0 mm (A, D), 0.5 mm (B, C, E).

Figure 36.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm.

In his publication on the genus , which appeared seven years later (Warncke 1982), only the sex and locality for each lectotype is cited, for example: “ Mor. / ♀ Lectotypus / Samarkand.” The selected specimen is not further individually identified and data on syntypes and paralectotypes are completely missing from the text. Thus, of the twenty seven designated lectotypes, fifteen are either invalid or unnecessary (when the type series includes only the holotype) and require corrections by subsequent authors (Pesenko 1986a, Ebmer 1997, current publication).

In the 1980’s, Yu. Pesenko continued the study of in the Fedtschenko’s collection of the ZISP and ZMMU, designating lectotypes for seven of Morawitz’s nominal taxa in the genera , and (Pesenko 1983, 1984, 1986a).

As a part of a detailed types inventory of the ZISP collection, all primary types of , including seven species described by F. Morawitz from the collection of A. Fedtschenko, are being progressively photographed and catalogued (Astafurova and Proshchalykin 2018, 2019, 2020). The present paper is the first complete, illustrated summary of all species of the family , described by F. Morawitz from the collection of A. Fedtschenko, an invaluable reference for researchers across this region who otherwise could not easily assign names to these difficult bees.

Materials and methods

All of the material listed below was examined for this study. In the following list, the taxa are treated in alphabetical order of the names used in the original descriptions. Each entry includes the name of the taxon in its original combination, the complete reference to the original description of the species (including the original combination and spelling of the name and the author, year and page of the description) and a list of type specimens present in the collections of the ZMMU and ZISP. The data from each label are separated by two slashes (//) . Square brackets are used for English translations and when information is added to specimen label information (e.g. geographical coordinates) or published data (e.g. current name of an old place name; affiliation to a present-day country). Photographs were made using a combination of a stereomicroscope Olympus SZX10 and a digital camera (Olympus OM-D and Canon EOS70D).

Illustrations were obtained by montaging from an image series that covers different focal planes into a single in-focus image with the Helicon Focus 6. The final illustrations were post-processed for contrast and brightness using Adobe® Photoshop® software.

The classification and current species status for and follow Michener (2007) and Ascher and Pickering (2020), for follow Astafurova et al. (2018a, b, 2019), for follow Astafurova (2014) and for follow Pesenko (1983).

Taxonomy

List of species

Subfamily

Genus

Morawitz, 1876

06D27E1F-7F9E-54A9-B19F-6B73690F7A82

Figure 1

Figure 1.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm (A, D), 0.5 mm (B, C).

Morawitz, 1876: 217 (key to females), 246, ♀.

Type locality.

Samarkand (Uzbekistan).

Published (original) locality.

Uzbekistan: Samarkand, Tashkent.

Lectotype.

♀, designated by Warncke 1982: 117, // 21.[III.1869] // Самаркандъ [Uzbekistan, Samarkand, ] // Mor., [N]372 [handwritten by F. Morawitz] // Mor., ♀ = lectotype, det A.W. Ebmer 1993 // Syntypus Lectotypus [ZMMU].

Paralectotypes

(12 ♀). 6 ♀, 11.[III.1871], 24., 25.[V.1871] // Ташкентъ [Tashkent] // [N]372; 1 ♀, 23.[III.1871] // Ташкентъ [Tashkent] // [N]372 // Mor., F. Morawitz det. [handwritten by F. Morawitz] [ZMMU]; 1 ♀, , 11.[III.1871] // Ташкентъ [Tashkent] // Mor., Typ. [handwritten by F. Morawitz]; 3 ♀, 11.[III.1871] // Ташкентъ [Tashkent] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor. [handwritten by F. Morawitz]; 1 ♀, 11.[III.1871] // Ташкентъ [Tashkent] // Paralectotypus Mor., design. Warncke, [19]82 Astafurova> [ZISP].

Current status.

(Dalla Torre, 1896), replacement name for Morawitz, 1876 (nec Schenck, 1853, nec Cresson, 1872).

Remarks.

The secondary designation of the lectotype by Ebmer 1995: 585 is unnecessary.

Description of male. Ebmer 1995: 585.

Distribution.

Kazakhstan, Turkmenistan, Uzbekistan, Afghanistan (Ebmer 1995).

Morawitz, 1876

3DD5E1FF-FF28-5BBA-885B-0EDB0E436896

Figure 2

Figure 2.

Morawitz, 1876, lectotype, female A habitus, lateral view B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view E labels. Scale bars: 1.0 mm (A), 0.5 mm (B–E).

Morawitz, 1876: 217 (key), 221, ♀.

Karatyube Mt., 15 km S Samarkand (Uzbekistan).

Uzbekistan: Samarkand, Karatyube Mt. (= 15 km S Samarkand).

designated by Warncke 1982: 81, ♀, 20[.V.1869] // Зеравшан.[ская] дол.[ина] [Uzbekistan, Zeravshan River valley, Karatyube Mt., ] // Mor., [N]332 [handwritten by F. Morawitz] // F. Morawitz det. 18.7.5. // Lectotypus Mor. 1876, design. Warncke, 1982 Astafurova> [ZMMU].

(2 ♀). 1 ♀, 17. [V.1869] // Самаркандъ [Uzbekistan, Samarkand] // [N]332 // F. Morawitz det. 1875 [ZMMU]; 1 ♀, 20[.V.1869] // Зеравшан.[ская] дол.[ина] [Zeravshan River valley] // Mor. Typ. [handwritten by F. Morawitz] // Paralectotypus Mor., 1876, design. Warncke, 1982 Astafurova> [ZISP].

(Morawitz, 1876).

Description of male. Kohl 1905: 238, as Vachal (synonymised by Warncke 1975: 89).

Bulgaria, Armenia, Turkey, Russia (North Caucasus), Afghanistan, Iran, Tajikistan, Uzbekistan, Kazakhstan, Pakistan (Astafurova and Proshchalykin 2017, Ascher and Pickering 2020).

Morawitz, 1876

B05FBBBF-88E9-58A6-9110-D1C63147BFBB

Figure 3

Figure 3.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm (A), 0.5 mm (B–D).

Morawitz, 1876: 216 (key), 228, ♀.

Kattakurgan (Uzbekistan).

Uzbekistan: Katty-Kurgan.

Lectotype (designated here).

♀, , 28. [IV.1869] [Uzbekistan, Kattakurgan, ] // ♀ // n. sp., ♀, F. Morawitz det., typus [handwritten by F. Morawitz] // Lectotypus Mor., design. Astafurova et Proshchalykin, 2020 [ZMMU].

Paralectotypes.

3 ♀, 28. [IV.1869] // Каттыкурганъ [Kattakurgan] // [N]343 // Paralectotypus Mor., design. Astafurova et Proshchalykin, 2020 [ZMMU].

(Morawitz, 1876).

Warncke (1982: 76) did not designate the lectotype, but only wrote “Typus Mus. Moskau”.

Description of male. Blüthgen 1925a: 119, as (synonymised by Blüthgen 1931a: 212).

Southern Kazakhstan, Turkmenistan, Uzbekistan, Mongolia, China (Xinjiang) (Pesenko 2007, Murao et al. 2017).

Morawitz, 1876

7D96A7D5-B70D-5CC6-824D-E5ADF1DA2B43

Figure (see Astafurova and Proshchalykin 2018: 11, figs 7a–e).

Morawitz, 1876: 218 (key), 254, ♀.

Tashkent (Uzbekistan).

Uzbekistan: Tashkent.

♀, designated by Ebmer 1985: 290, 8.[VIII.1870] // Ташкентъ [Uzbekistan, Tashkent, ] // Mor. [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] / / Syntypus // (Mor.), ♀, Lectotypus, det. A.W. Ebmer 1985 // Lectotypus [ZISP].

Paralectotype.

1 ♀, 8.[VIII.1870] // Ташкентъ [Tashkent] // Mor. [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] [ZISP].

( (Morawitz, 1876) (subspecies status according to Ebmer 1988b: 667).

Description of male. Bytinski-Salz and Ebmer 1974: 211, as Ebmer 1974 (synonymised by Ebmer 1985: 290).

Egypt, Palestine, Syria, Jordan, Israel, Turkey, Iran, Central Asia (Bytinski-Salz and Ebmer 1974, Warncke 1982, Pesenko 2007).

Morawitz, 1876

85C6AF01-01DF-5091-AD67-DF398449A07A

Figure 4

Figure 4.

Morawitz, 1876, lectotype, male A habitus, lateral view B metasoma, dorsal view C head, frontal view D mesosoma, dorsal view E labels. Scale bars: 1.0 mm.

Morawitz, 1876: 220 (key), 226, ♂.

Osh (Kyrgyzstan).

Uzbekistan: Samarkand, Dzhyuzak [=Jizzakh], Sokh; Kyrgyzstan: Osh.

♂, designated by Blüthgen 1955: 19, 1.[VIII.1871] // Ошъ [Kyrgyzstan, Osh, ] // Mor., [N]341 [handwritten by F. Morawitz] // Lectotypus Mor., design. Blüthgen Astafurova> [ZMMU].

(4 ♂). 2 ♂, 4., 7.[VII.1869] // Самарканд [Samarkand] // [N]341; 2 ♂, 18., 22.[VII.1870] // Джюзакъ [Dzhyuzak] // [N]341 // Paralectotypus , design. Blüthgen Astafurova> [ZMMU].

Morawitz, 1876 (subspecies status according to Ebmer 1988b: 578).

The lectotype designation by Warncke (1982: 138) is unnecessary.

Description of female: Strand 1921: 314, as (synonymised by Ebmer 1988b: 578).

Europe (except north), Russia (European part, except north), Turkey, Israel, Iran, Afghanistan, Pakistan, Central Asia, Mongolia, North China (Pesenko 2006a, Astafurova and Proshchalykin 2017).

Morawitz, 1876

ED601DFE-909A-5859-AE9F-EB0E15954C64

Figure 5

Figure 5.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm.

Morawitz, 1876: 218 (key), 245, ♀.

Samarkand (Uzbekistan).

Uzbekistan: Samarkand, Dzham [near Samarkand], Aksay [near Samarkand].

♀, 18.[III.1869] // Самаркандъ [Uzbekistan, Samarkand, ] // Mor., [N]371 [handwritten by F. Morawitz] // Lectotypus Mor., design. Astafurova et Proshchalykin, 2020 [ZMMU].

(22 ♀). 11 ♀, 27., 28. [II. 1869], 18., 21. [V. 1869] // Самаркандъ [Samarkand] // [N]371; 1 ♀, 12.[V.1969] // Заравшан.[ская] дол.[ина] [Zeravshan River valley, Dzham] // [N]370; 1 ♀, 16.[V.1969] // Заравшан.[ская] дол.[ина] [Zeravshan River valley, Aksay] // [N]370 [ZMMU]; 1 ♀, the same labels, but 21.[V.1869] [ZMMU]; 1 ♀, // 18.[V.1869] // Самаркандъ [Samarkand] // Mor. Typ. [handwritten by F. Morawitz]; 1 ♀, 18.[III.1869] // Самаркандъ [Samarkand] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♀, 20.[III.1869] // Самаркандъ [Samarkand] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor., ♀, CoType, F. Morawitz det. [handwritten by F. Morawitz]; 1 ♀, 27.[III.1869] // Самаркандъ [Samarkand] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♀, 16.[III.1869] // Самаркандъ [Samarkand] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor., ♀, F. Morawitz det. [handwritten by F. Morawitz]; 1 ♀, 30.[III.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley, Samarkand] // Mor. [handwritten by F. Morawitz]; 2 ♀, 12.[III.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley, Samarkand] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Mor., design. Astafurova et Proshchalykin, 2020 [ZISP].

(Morawitz, 1876).

Male unknown.

The lectotype designation by Warncke (1982: 116) is invalid because he labelled none of the eleven females from Samarkand deposited in ZMMU.

South Kazakhstan, Central Asia, Pakistan (Ebmer 1995, Murao et al. 2017).

Morawitz, 1876

0C3828D5-D7F1-5756-9507-F7DD0D8AE2DF

Figure 6

Figure 6.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm (A, D), 0.5 mm (B, C).

Morawitz, 1876: 217 (key to ♀), 220 (key to ♂), 224, ♀, ♂.

Yeri, Sughd Province (Tajikistan).

Kyrgyzstan: Taka; Uzbekistan: Karatyube [Mt., 15 km S Samarkand], Dzham Gorge, Urgut; Tajikistan: Iori.

♀, designated by Ebmer 1997: 956, // 1.[VI.1869] // Заравш.[анская] дол.[ина] [Tajikistan, Zeravshan River valley, Iori (= Yeri), ] // Mor. Typ [handwritten by F. Morawitz] // Mor. Blüthgen det. 1935 // Lectotype // (Mor.), ♀, Lectotypus, det. A.W. Ebmer 1974 [ZMMU].

(6 ♀). 1 ♀, 20.[V.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley, Karatyube] // Mor., [N]338 [handwritten by F. Morawitz] // Mor. Blüthgen det. 1935; 3 ♀, 8.[VIII.1871] // Така [Taka] // [N]338; 1 ♀, 13.[V.1869] // Джамское ущ. [Dzham Gorge] // [N]338 [ZMMU]; 1 ♀, 1.[VI.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley, Iori] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor., Blüthgen det. 1935 // Paralectotypus Mor., design. Ebmer 1997 Astafurova> [ZISP].

(Morawitz, 1876).

The lectotype designation by Warncke (1982: 106) is invalid because there are two females in ZMMU from the Zeravshan River valley, neither with Warncke’s lectotype label.

Turkey, Iran, Afghanistan, Central Asia, Kazakhstan (Ebmer 1997, Murao et al. 2017).

Morawitz, 1876

D1065512-A6E2-5834-99CF-F239AE07EA09

Figure 7

Figure 7.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm (A), 0.5 mm (B–D).

Morawitz, 1876: 217 (key), 228, ♀.

Kattakurgan (Uzbekistan).

Uzbekistan: Katty-Kurgan.

♀, designated by Warncke 1982: 107, 28.[IV.1869] // Катты-Курганъ [Uzbekistan, Katty-Kurgan (=Kattakurgan), ] // Mor., [N]344 [handwritten by F. Morawitz] // Lectotypus Mor., design. Warncke Astafurova> [ZMMU].

Morawitz, 1876.

Description of male. Blüthgen 1929: 84, figs 9a, b.

Turkey, southern Kazakhstan, Turkmenistan, Uzbekistan, Pakistan (Blüthgen 1929, Ebmer 1988a).

Morawitz, 1876

23A7E390-6AFD-58F8-8C57-54164BF4725B

Figure 8

Figure 8.

Morawitz, 1876, lectotype, female A habitus, lateral view B metasoma, dorsal view C mesosoma, dorsal view D head, frontal view E labels. Scale bars: 1.0 mm.

Morawitz, 1876: 217 (key), 233, ♀.

30 km SSE Samarkand, Sangu-dzhuman Pass (Uzbekistan).

Uzbekistan: “On the road to Sangy-Dzhuman [pass] and the Kulbasy Mountain”.

♀, designated by Pesenko 1984: 20, 25.[V.1869] // Сангы-Джуманъ [Uzbekistan, Sangu-dzhuman Pass, Zeravshan Ridge, 30 km SSE Samarkand, ] // Mor., [N]351 [handwritten by F. Morawitz] // Lectotypus Mor., ♀, design. Pesenko [1]981 [ZMMU].

(6 ♀). 3 ♀, the same label as in the lectotype; 1 ♀, 25.[V.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley] // [N]351 [ZMMU]; 2 ♀, Сангы-Джуманъ [Sangu-Dzhuman] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // F. Mor., ♀ [handwritten by F. Morawitz] // Paralectotypus Mor., ♀, design. Pesenko [1]981 [ZISP].

Dalla Torre, 1896, replacement name for Morawitz, 1876 (nec Walker, 1871).

Description of male. Ebmer 1980: 471, Fig. 1.

A rare montane Central Asian species. Northern Afghanistan, eastern Uzbekistan, Tajikistan and Kyrgyzstan (Pesenko 2005a).

Morawitz, 1876

769169D6-863A-5767-B503-0384BF866B38

Figure 9

Figure 9.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm.

Morawitz, 1876: 217 (key), 242, ♀.

Khozyay-Dun (Uzbekistan).

Uzbekistan: Khodzhaduk [= Khozyay-Dun].

♀, designated by Warncke 1982: 105, 21.[V.1869] // Заравшан.[ская] дол.[ина] [Uzbekistan, Zeravshan River valley, Khozyay-Dun, ] // Mor., [N]366 [handwritten by F. Morawitz] // Lectotypus Mor., design. Warncke [19]82 [ZMMU].

(Morawitz, 1876).

Description of male. Morawitz 1876: 219 (key), 243, as (synonymised by Blüthgen 1926: 391).

According to the Fedchenko list (Baker 2004: 243), the locality given in Warncke’s lectotype designation is in Uzbekistan, not Tajikistan.

Turkey, Uzbekistan, Tajikistan, Kyrgyzstan, south-eastern Kazakhstan (Pesenko 1986a, Ascher and Pickering 2020).

Morawitz, 1876

BD5F0E37-23E8-58FA-908F-A3558314BEE3

Figure 10

Figure 10.

Morawitz, 1876, lectotype, male A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm.

Morawitz, 1876: 219 (key), 243, ♂.

Shakhimardan (Uzbekistan).

Uzbekistan: near Shakhimardan.

♂, designated by Warncke 1982: 105, // 6.[VII.1871] // Шагимарданъ [Shakhimardan in the Uzbek enclave in the territory of Kyrgyzstan, Alai Ridge; ] // Mor., [N]367 [handwritten by F. Morawitz] // Lectotypus Mor., ♂, design. Warncke [19]82 [ZMMU].

(2 ♂). 1 ♂, // 2.[VII.1871] // Шагимарданъ [Shagimardan] // Mor., Typ. [handwritten by F. Morawitz]; 1 ♂, 2.[VII.1871] // Шагимарданъ [Shagimardan] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // M. [handwritten by F. Morawitz] // Mor., Pesenko det., 1985 // Paralectotypus, Mor., design. Warncke, [19]82 [ZISP].

(Morawitz, 1876) (synonymised by Blüthgen 1926: 391).

See Morawitz, 1876.

Morawitz, 1876

E2FF2321-642E-53B7-92A4-441223EDEA8C

Figure 11

Figure 11.

Morawitz, 1876, holotype, female A habitus, lateral view and labels B metasoma, dorsal view C head, frontal view D mesosoma, dorsal view. Scale bars: 1.0 mm.

Morawitz, 1876: 219 (key), 230, ♂.

30 km SE Kozhatogai, Turkistan Province (Kazakhstan).

Uzbekistan: steppe between Tashkent and Syrdarya River.

Holotype.

♂, 18.[V.1871] // Степь м.[ежду] С.[ыр] д.[арьей] и Т.[ашкентом] [Kazakhstan, Turkistan Province, steppe between Syrdarya River and Tashkent, 30 km SE Kozhatogai, ] // Mor., [N]345 [handwritten by F. Morawitz] // Evers. v. Mor., ♂, P. Blüthgen det. // Holotypus [ZMMU].

(Eversmann, 1852) (synonymised by Blüthgen 1922: 47).

The lectotype designation of Warncke (1982: 148) is unnecessary as the species was described from a single male that was directly written about by Morawitz (1876: 231).

North Africa, South and East Europe, Russia (south to Urals Mountains on the east), Caucasus, Turkey, Near East, Iraq, Iran, Afghanistan, Central Asia, Kazakhstan, Mongolia, China (Astafurova and Proshchalykin 2017).

Morawitz, 1876

D33FA76A-9B57-5EF7-8896-ED609504A6FA

Figure 12

Figure 12.

Morawitz, 1876, holotype, female A habitus, lateral view and labels B metasoma, dorsal view C head, frontal view D mesosoma, dorsal view. Scale bars: 1.0 mm.

Morawitz, 1876: 219 (key), 239, ♂.

Khodzha-Chiburgan River (Tajikistan/Kyrgyzstan, not Uzbekistan as was indicated in Warncke 1982: 91).

Tajikistan/Kyrgyzstan: Khodzha-Chiburgan River.

♂, 26.[VI.1871] // Чибурганъ [Tajikistan/Kyrgyzstan: Khodzha-Chiburgan River (near Vorukh, ≈ ) // Mor., [N]361 [handwritten by F. Morawitz] // Holotypus [ZMMU].

(Morawitz, 1876).

Description of male. Blüthgen 1934b: 147.

The lectotype designation by Warncke (1982: 91) is unnecessary as the species was described from a single male that was directly written about by Morawitz (1876: 240).

Tajikistan, Kyrgyzstan, Kazakhstan (Morawitz 1876, Pesenko 1986a, Murao et al. 2017).

Morawitz, 1876

BB3C364F-3185-508E-84E4-99096206C174

Figure (see Astafurova and Proshchalykin 2020: 414, figs 11a–e).

Morawitz, 1876: 217 (key), 235, ♀.

30 km SSE Samarkand, Sangu-dzhuman Pass (Uzbekistan).

Uzbekistan: Sangu Dzhuman.

♀, designated by Pesenko 1984: 21, // Сангы Джуманъ [Uzbekistan, Sangydzhuman Pass, 30 km SSE Samarkand, ] // 25.[V.1869] // Mor. Typ. [handwritten by F. Morawitz] // Lectotypus Mor., ♀, design. Pesenko [1]981 // Zoological Institute St. Petersburg INS_HYM_0000159 [ZISP].

(8 ♀). 8 ♀, the same label as in lectotype // Paralectotypus Mor. design. Pesenko [1]981 [3 ♀ – ZISP; 5 ♀ – ZMMU].

Morawitz, 1876.

A rare montane Central Asian species: Kazakhstan, Uzbekistan, Tajikistan, Western Kyrgyzstan, Iran, north-eastern Afghanistan and north-western China (Pesenko 2005a).

Morawitz, 1876

871A8FE5-8CA9-5E23-9138-F676B77806EF

Figure 13

Figure 13.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 0.5 mm.

Morawitz, 1876: 217 (key), 229, ♀.

50 km NW Chardara, Kyzylkum Desert (Turkistan Province, Kazakhstan).

Kazakhstan: “Kyzyl-Kum Steppe, near Baybek”.

♀, designated by Warncke 1982: 137, 30.[IV.1871] // Кизилъкумъ [Kazakhstan, Baybek Well, Kyzylkum Desert, ca. 50 km NW Chardara, ≈ ] // Mor., [N]345 [handwritten by F. Morawitz] // Mor., ♀, Lectotypus, design. A.W. Ebmer 1994 // Lectotypus Mor., design. Warncke [ZMMU].

1 ♀, Kisilkum [handwritten by F. Morawitz] // Mor. Typ. [handwritten by F. Morawitz] // Paralectotypus , labelled by Yu. Astafurova [ZISP].

Morawitz, 1876.

Description of male. Morawitz 1894: 68, as (synonymised by Blüthgen 1931a: 214).

Southern Kazakhstan, Turkmenistan, Iran, China (Xinjiang) (Morawitz 1876, 1894, Murao et al. 2017, Ascher and Pickering 2020).

Morawitz, 1876

F8F496CE-73D6-5D1A-897C-ADA4523E76EF

Figure (see Astafurova and Proshchalykin 2018: 23, figs 21a–e).

Morawitz, 1876: 218 (key to females), 220 (key to males), 253–254, ♀, ♂.

Tashkent (Uzbekistan).

Kazakhstan: Chardara, steppe between Tashkent and Syr-Darya; Uzbekistan: Dzhamsk Gorge, Dzhizmansk Gorge, Ulus, Dzham, Urgut, Keles, Samarkand, Soch, Shakhimardan, Uch-Kurgan; Kyrgyzstan: Alay, Osh, Gulsha, Taka.

♀, designated by Astafurova and Proshchalykin 2018: 22, Ташкентъ [Uzbekistan, Tashkent, ] // F. Mor., ♀ [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Lectotypus Morawitz, 1876, ♀, design. Astafurova & Proshchalykin 2018 [ZISP].

(29 ♀, 10 ♂). 1 ♀, Шагимарданъ [Shagimardan] // 3.[VII.1871] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♂, Така [Taka] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♀, 1 ♂, Сохъ [Sokh] // 29. [29.VI.1871] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♂, Уч-Курганъ [Uch- Kurgan] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 2 ♀, Ташкентъ [Tashkent] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] [ZISP]; 1 ♂, Ошъ [Osh] // 1.[VIII.1871]; 1 ♂, Самаркандъ [Samarkand] // 4.[VII.1869] // [N]384; 3 ♂, Самаркандъ [Samarkand] // 7.[VII.1869]; 2 ♀, Самаркандъ [Samarkand] // 3., 21.[III.1869]; 16 ♀, Ташкентъ [Tashkent] // 26., 27.[III.1871], 21., 23.[V.1871] and 1.,3.,5.,10.[VI.1871]; 1 ♀, Чардара [Chardara] // 27.[IV.1871]; 1 ♀, Учь-Курганъ [Uch-Kurgan] // 15.[VII.1871]; 2 ♂, Шагимарданъ [Shagimardan] // 2.[VII.1871] 5 ♀, Заравшан.[ская] дол.[ина] [Zeravshan River valley], 3., 11., 18., 23.[III.1871] // Paralectotypus Morawitz, 1876, design. Astafurova & Proshchalykin 2018 [ZMMU].

(Morawitz, 1876).

Iran, Afghanistan, Kazakhstan, Uzbekistan, Kyrgyzstan, Tajikistan (Morawitz 1876, Ebmer 1974, Warncke 1982, Murao et al. 2017).

Morawitz, 1876

7C938E0B-F404-5BEB-BC69-F48764E583B4

Figure (see Astafurova and Proshchalykin 2018: 24, figs 23a–e).

Morawitz, 1876: 218 (key to females), 248, ♀.

30 km SSE Samarkand, Sangu-dzhuman Pass (Uzbekistan).

Uzbekistan: Sangy Dzhuman.

♀, // Сангы Джуманъ [Uzbekistan, Sangy-dzhuman Pass, 30 km SSE Samarkand, Zeravshan Ridge, ] // 25.[III.1869] // Mor., Typ. [handwritten by F. Morawitz] // Lectotype , design. Astafurova et Proshchalykin, 2020 [ZISP].

(2 ♀). 1 ♀, 25.[III.1869] // Сангы Джуманъ [Sangy Dzhuman] // Mor. [handwritten by F. Morawitz]; 1 ♀ 25.[III.1869] // Сангы Джуманъ [Sangy Dzhuman] // [N]375 // Paralectotype , design. Astafurova et Proshchalykin, 2020 [ZMMU].

(Morawitz, 1876).

Astafurova and Proshchalykin (2018: 23) indicated the lectotype specimen as «Holotype», but Morawitz (1876) did not directly indicate a single specimen and two specimens from the type series are deposited in ZMMU.

Description of male. Blüthgen, 1934b: 154, Fig. 3.

Kazahkstan, Uzbekistan, Tajikistan, Kyrghyzstan, Afghanistan (Morawitz 1876, Blüthgen in Popov 1935, Ebmer 1980, Murao et al. 2017).

Morawitz, 1876

0CB23044-6369-5D2D-A43E-C7004EE64026

Figure 14

Figure 14.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm (A), 0.5 mm (B–D).

Morawitz, 1876: 218 (key), 249, ♀.

Samarkand (Uzbekistan).

Uzbekistan: Samarkand; Tajikistan: Peti.

♀, 5.[IV.1869] // Самаркандъ [Uzbekistan, Samarkand, ] // Mor., [N]377 [handwritten by F. Morawitz] // Lectotypus Mor., Astafurova et Proshchalykin, 2020 [ZMMU].

(3 ♀). 1 ♀, 5.[IV.1869] // Самаркандъ [Samarkand] // [N]377 // Typus [ZMMU]; 1 ♀, // 5.[IV.1869] // Самаркандъ [Samarkand] // F. Mor., Typ. [handwritten by F. Morawitz]; 1 ♀, Самаркандъ [Samarkand] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor., Typ. [handwritten by F. Morawitz] // Paralectotypus Mor., Astafurova et Proshchalykin, 2020 [ZISP].

(Morawitz, 1876).

Description of male. Blüthgen 1930: 763.

The lectotype designation by Warncke (1982: 69) is invalid because he labelled no female of the two females from Samarkand deposited in ZMMU.

Central and eastern Europe, Turkey, Caucasus, Russia (North Caucasus), Israel, Iran, Afghanistan, Tajikistan, Uzbekistan, Kazakhstan, China (Gansu) (Pesenko 2007, Astafurova and Proshchalykin 2017, Murao et al. 2017).

Morawitz, 1876

FA1223E9-2585-5D0A-A360-4B8E58FC0ADC

Figure 15

Figure 15.

Morawitz, 1876, lectotype, male A head, frontal view B habitus, lateral view C mesosoma, dorsal view D metasoma, dorsal view E labels. Scale bars: 1.0 mm (A, B), 0.5 mm (C, D).

Morawitz, 1876: 216 (key to females), 219 (key to males), 236, ♀, ♂.

Shakhimardan (Uzbekistan).

Uzbekistan: on the road to Sangy-dzhuman Pass, Shakhimardan; Tajikistan: Peti.

♂, designated by Warncke 1982: 80, 3.[VII.1871] // Шагимарданъ [Shakhimardan in the Uzbek enclave in the territory of Kyrgyzstan, Alai Ridge; ] // Mor., [N]356 [handwritten by F. Morawitz] // Lectotypus Mor., design. Warncke [19]82 Astafurova> [ZMMU].

(2 ♀). 1 ♀, 25.[V.1869] // Сангы Джуманъ [Sangy Dzhuman] // [N]356; 1 ♀, 12.[VII.1870] // Фанъ [Fan] // [N]356 [ZMMU]; 1 ♀, 2.[VII.1871] // Шагимарданъ [Shagimardan] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor. [handwritten by F. Morawitz] // Paralectotypus Mor., design. Warncke < identical red labels on each paralectotype specimen, labelled by Yu. Astafurova> [ZISP].

(Morawitz, 1876).

Greece, Israel, Lebanon, Turkey, Caucasus, Iran, Afghanistan, Central Asia, Kazakhstan, China (Xinjiang) (Murao et al. 2017, Niu et al. 2020).

Morawitz, 1876

B576028E-08FD-52B4-A80E-8B044BCC73C2

Figure 16

Figure 16.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm (A), 0.5 mm (B–D).

Morawitz, 1876: 218 (key to females), 247, ♀.

Sokh District [Uzbekistan].

Kekh [Sokh District, Uzbekistan].

♀ designated by Warncke 1982: 68, // 27.[VI.1871] // Сохъ [Uzbekistan, Sokh District, ≈ ] // Mor., [N]373 [handwritten by F. Morawitz] // Lectotypus Mor., design. Warncke Astafurova> [ZMMU].

(Morawitz, 1876).

Male unknown.

Turkey, Turkmenistan, Tajikistan, Uzbekistan, Iran, Afghanistan (Morawitz 1876, Ebmer 1984, 1986, Warncke 1982, Ascher and Pickering 2020).

Morawitz, 1876

13F40084-10DA-56C1-A4F2-92B7254C56F6

Figure 17

Figure 17.

Morawitz, 1876, holotype, male A view habitus, lateral view and labels B head, frontal C genitalia, dorsal view D mesosoma, dorsal view E metasoma, dorsal view. Scale bars: 1.0 mm (A, B, D, E), 0.5 mm (C).

Morawitz, 1876: 241, ♂.

Shakhimardan (Uzbekistan).

Near Shakhimardan.

♂, 9.[VII.1871] // Шагимарданъ [Shakhimardan in the Uzbek enclave in the territory of Kyrgyzstan, Alai Ridge; ] // Mor., [N]364 [handwritten by F. Morawitz] // (Mor.) syn: (Mor.) det. A.W. Ebmer 1979 // Holotypus [ZMMU].

(Morawitz, 1876) (subspecies status according to Ebmer 1998: 382).

Description of female: Ebmer 1980: 493, Figs 12 and 13, as Ebmer, 1980 (synonymised by Ebmer 1998: 382).

The lectotype designations by Ebmer (1980: 495) and by Warncke (1982: 91) were unnecessary as the species was described from a single male that was directly written about by Morawitz (1876: 241).

Kazakhstan, Uzbekistan, Tajikistan, Kyrgyzstan, Afghanistan, Mongolia (Hovd) (Pesenko 2006b).

Morawitz, 1876

B9D20F93-7D49-5662-B92C-95BD9EA40A1A

Figure (see Astafurova and Proshchalykin 2020: 418, figs 15a–e).

Morawitz, 1876: 217 (key), 233, ♀.

30 km SSE Samarkand, Sangu-dzhuman Pass (Uzbekistan).

Uzbekistan: Gus [near Urgut], Sangy-Dzhuman; Tajikistan: Pyandzhikent.

♀, designated by Pesenko 1984: 23, // Сангы Джуманъ [Uzbekistan, Sangy-dzhuman Pass, 30 km SSE Samarkand, ] // 25.[V.1869] // Mor. [handwritten by F. Morawitz] // Lectotypus Mor., design. Pesenko, [1]981, ♀ // Zoological Institute St. Petersburg INS_HYM_0000164 [ZISP].

(3 ♀). 1 ♀, Сангы Джуманъ [Sangy Dzhuman] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor. [handwritten by F. Morawitz] // Syn.: jarkandensis Strand, ♀ // Paralectotypus Mor., design. Pesenko, [1]981, ♀ [ZISP]; 1 ♀, 25.[V.1869] // Сангы Джуманъ [Sangy Dzhuman] // Mor., [N]352 [handwritten by F. Morawitz]; 1 ♀, 24.[V.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley, Gus] // [N]352 // Paralectotypus Mor., design. Pesenko, [1]981, ♀ [ZMMU].

Morawitz, 1876.

Description of male. Blüthgen 1936: 295, fig. 11.

Azerbaijan, Afghanistan, Iran, Kazakhstan, Central Asia, Altai, Pakistan, north-western and northern China, northern India (Pesenko 2005a, b).

Morawitz, 1876

2CE15736-D68B-502E-9EAE-6318C69E3B1F

Figure 18

Figure 18.

Morawitz, 1876, holotype, female A habitus, lateral view B head, frontal view C mesosoma, dorsal view D labels E metasoma, dorsal view. Scale bars: 1.0 mm.

Morawitz, 1876: 217 (key), 235, ♀.

Samarkand (Uzbekistan).

Uzbekistan: near Samarkand.

♀, 5.[VII.1870] // Самаркандъ [Uzbekistan, Samarkand, ] // Mor., [N]354 [handwritten by F. Morawitz] // Holotypus [ZMMU].

Morawitz, 1876.

The lectotype designation of Warncke (1982: 147) is unnecessary as the species was described from a single female that was directly written about by Morawitz (1876: 235).

Description of male. Ebmer 1984: 315, figs 3–5.

Turkmenistan, Uzbekistan, Kyrgyzstan, Afghanistan, Pakistan (Pesenko 2005a).

Morawitz, 1876

D85BC9E9-13A7-5DF7-BC2E-5BDE6246F0E3

Figure 19

Figure 19.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm (A, D), 0.5 mm (B, C).

Morawitz, 1876: 216 (key to females), 219 (key to males), 229, ♀, ♂.

Samarkand (Uzbekistan).

Southern Kazakhstan: Kysyl-Kum [desert] near draw-well Chakany; Uzbekistan: steppe between Katty-Kurgan and Ulus, Samarkand, Murzarabat, Chinaz, Sokh.

♀, 9.[VI.1869] // Заравшан.[ская] дол.[ина] [Uzbekistan, Zeravshan River valley, near Samarkand, ] // Mor., [N]346 [handwritten by F. Morawitz] // Lectotypus Mor., design. Astafurova et Proshchalykin, 2020 [ZMMU].

(28 ♀, 44 ♂). 14 ♀, the same label as in the lectotype; 11 ♀, 2 ♂, 9., 13.[VI.1869], 4., 7.[VII.1869] // Самаркандъ [Samarkand] // [N]346; 12 ♂, 24.[VII.1870], 29.[VIII.1870] // Мурзарабадъ [Murzarabad] // [N]346; 1 ♀, 9 ♂, 28.[IV.1871] // Кизилъкумъ [Kizilkum] //[N]346; 17 ♂, 25.[VII.1870] // Чиназъ [Chinaz] // [N]346 [ZMMU]; 1 ♀, 28.[IV.1871] // Кизилъкумъ [Kizilkum] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor. [handwritten by F. Morawitz]; 1 ♀, // 9.[VI.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley, near Samarkand] // Mor. Typ. [handwritten by F. Morawitz]; 3 ♂, 28.[VI.1871] // Сохъ [Sokh] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♂, 9.[VI.1869] // Самаркандъ [Samarkand] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Mor., design. Astafurova et Proshchalykin, 2020 [ZISP].

Morawitz, 1876.

The lectotype designation by Warncke (1982: 138) is invalid because he labelled none of the 15 females from “valle Serafshan” deposited in ZMMU.

Morocco, Kazakhstan, Uzbekistan, Turkmenistan, Iran, Afghanistan, Pakistan (Murao et al. 2017, Ascher and Pickering 2020).

Morawitz, 1876

6F363A18-5778-5D9F-A346-BA56996CDC92

Figure 20

Figure 20.

Morawitz, 1876, lectotype, male A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D antennae, dorsal view E metasoma, dorsal view. Scale bars: 1.0 mm.

Morawitz, 1876: 249, ♂.

Sarafschan River valley, Yeri (Tajikistan).

Tajikistan: valley Sarafschan between Iori and Dashty-Kazy.

♂, designated by Warncke 1982: 91, // 31.[V.1869] // Заравшан.[ская] дол.[ина] [Tajikistan, Zeravshan River valley, near Iori (= Yeri), ] // Mor., [N]378 [handwritten by F. Morawitz] // Lectotypus Mor. design. Warncke [1]982 [ZMMU].

1 ♂, Заравшан.[ская] дол.[ина] [Zeravshan River valley] // F. Morawitz [handwritten by F. Morawitz] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Mor., design. Warncke [1]982 // Zoological Institute St. Petersburg, INS_HYM 0000076 [ZISP].

(Morawitz, 1876).

Description of female. Blüthgen 1931b: 336, as (synonymised by Ebmer 1978: 39).

Iran, Afghanistan, Turkmenistan, Uzbekistan, Tajikistan, Kyrgyzstan (Blüthgen 1931b, Pesenko 1986a).

Morawitz, 1876

24CFCD7C-6EAB-522D-8E03-B15E80836F2A

Figure 21

Figure 21.

Morawitz, 1876, lectotype, male A habitus, lateral view B metasoma, dorsal view C head, frontal view D mesosoma, dorsal view E labels. Scale bars: 1.0 mm (A), 0.5 mm (B–D).

Morawitz, 1876: 251, ♂.

Karazuk, vicinity of Shakhimardan (Uzbekistan).

Tajikistan: Iori George, Iskander River; Uzbek enclave in Kyrgyzstan: Karakazuk; Kyrgyzstan: Alay.

♂, designated by Blüthgen 1934a: 302, 11.[VII.1871] // Каразукъ [Uzbekistan, Karazuk, vicinity of Shakhimardan ] // Mor., [N]380 [handwritten by F. Morawitz] // Mor., ♂, lecto-holotype, Blüthgen det. 1933 // Lectotypus Mor., design. Blüthgen [19]34 labelled by Yu. Astafurova [ZMMU].

(3 ♂). 1 ♂, 23.[VII.1871] // Алай [Alay] // [N]380 // Mor. ♂, Lecto-Paratype, Blüthgen det., 1933 [ZMMU]; 1 ♂, // 23.[VII.1871] // Алай [Alay] // Mor. Typ., [N]380 [handwritten by F. Morawitz]; 1 ♂, 22.[VII.1871] // Алай [Alay] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor. [handwritten by F. Morawitz] // Paralectotypus Mor., design. Blüthgen Astafurova> [ZISP].

(Dalla Torre, 1896), replacement name for Morawitz, 1876 (nec Lepeletier, 1841).

The specimens from Iskander and Iori George in the Morawitz type series are the holotype and paratypes of Blüthgen, 1934.

Description of female: Vachal 1902: 229, as (synonymised by Blüthgen 1934a: 301).

Uzbekistan, Kyrgyzstan, Kazakhstan, Tajikistan, Afghanistan, China (Xinjiang) (Morawitz 1876, Murao et al. 2017, Ascher and Pickering 2020).

Morawitz, 1876

EBA6D4CC-95C9-543D-957F-98FFF03A1131

Figure 22

Figure 22.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B metasoma, dorsal view C mesosoma, dorsal view D head, frontal view E. Scale bars: 1.0 mm.

Morawitz, 1876: 218 (key), 245, ♀.

Sangy-dzhuman Pass, 30 km SSE Samarkand [Uzbekistan].

Uzbekistan: Samarkand, Aksay; Tajikistan: Iori Gorge, Varzaminor [=Ayni], Sangy-Dzhuman Pass.

♀, designated by Warncke 1982: 116, 25.[V.1869] // Сангы Джуманъ [Uzbekistan, Sangy-dzhuman Pass, 30 km SSE Samarkand, ] // Mor., [N]370 [handwritten by F. Morawitz] // Lectotypus Mor., design. Warncke [19]82 Astafurova> [ZMMU].

(7 ♀). 1 ♀, 3.[IV.1869] // Самаркандъ [Samarkand] // [N]370; 1 ♀, 27.[II.1869] // Самаркандъ [Samarkand] // [N]370 // [handwritten by F. Morawitz]; 1 ♀, [7.VI.1870] // Варзаминоръ [Varzaminor] // [N]370; 1 ♀, 16.[V.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley, Aksay] // N[370]; 1 ♀, 2.[VI.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley, Iori Gorge] // N[370]; [ZMMU]; 1 ♀, // 25.[V.1869] // Сангы Джуманъ [Sangy Dzhuman] // Mor., Typ. [handwritten by F. Morawitz]; 1 ♀, Сангы Джуманъ [Sangy Dzhuman] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor.[handwritten by F. Morawitz]; 1 ♀, Заравшан.[ская] дол.[ина] [Zeravshan River valley] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // F. Morawitz, ♀ [handwritten by F. Morawitz] // Paralectotypus Mor., design. Warncke Astafurova> [ZISP].

(Morawitz, 1876).

Description of male. Blüthgen 1923a: 277.

Europe (except North), Cyprus, Azerbaijan, Russia (North Caucasus), Turkey, Syria, Jordan, Israel, Iran, Afghanistan, Central Asia, Kazakhstan (Astafurova and Proshchalykin 2017).

Morawitz, 1876

F01DBEB8-81F4-50B8-A2AE-D44EEF813413

Figure 23

Figure 23.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm.

Morawitz, 1876: 217 (key), 234, ♀.

Iskanderkul Lake (Tajikistan).

Tajikistan: “near Iskander-Kul Lake”.

♀, designated by Warncke 1982: 147, // 15.[VI.1870] // Искандеръ [Tajikistan, Iskanderkul Lake, Hissar Ridge, ] // Mor., [N]353 [handwritten by F. Morawitz] // Lectotypus Mor., ♀, design. Warncke [1]982 , labelled by Yu. Pesenko [ZMMU].

1 ♀, // 15.[VI.1870] // Искандеръ [Iskander] // Mor., Typ. [handwritten by F. Morawitz] // Paralectotypus Mor., ♀, design. Pesenko [1]981 [ZISP].

Morawitz, 1876.

Description of male. Blüthgen 1936: 291.

Kazakhstan, Uzbekistan, Tajikistan, Kyrgyzstan, China (Xinjiang) (Pesenko 1986b, Murao et al. 2017).

Morawitz, 1876

303C41DE-B945-5FF1-877F-EFB22CE12547

Figure 24

Figure 24.

Morawitz, 1876, holotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm (A), 0.5 mm (B–D).

Morawitz, 1876: 218 (key), 251, ♀.

Gulcha (Kyrgyzstan).

Kyrgyzstan: Gulsha [Gulcha].

♀, 10.[VIII.1871] // Гульша [Kyrgyzstan, Gulcha, ] // Mor., [N]381 [handwritten by F. Morawitz] // Holotype Mor., 1876 [ZMMU].

(Dalla Torre, 1896), replacement name for Morawitz, 1876 (nec Smith, 1853) (subspecies status according to Ebmer 1997: 932).

Description of male. Blüthgen 1923a: 271, as (synonymised by Warncke 1975: 96).

The lectotype designation by Warncke (1982: 106) is unnecessary as the species was described from a single female that was directly written about by Morawitz (1876: 251).

Egypt, Turkey, Near East, Iran, Azerbaijan, Central Asia, Mongolia (Hovd), China (Xinijang) (Ebmer 1997, Pesenko 2007).

Morawitz, 1876

49D01D9B-6A20-5422-B4C4-404DB7903E8E

Figure 25

Figure 25.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C mesosoma, dorsal view D metasoma, dorsal view. Scale bars: 1.0 mm (A, C, D), 0.5 mm (B).

Morawitz, 1876: 218 (key), 244, ♀.

Zeravshan River valley (Tajikistan).

between Panjakent and Iori (Tajikistan).

♀, designated by Pesenko 1986a: 138, 30.[V.1869] // Заравш.[анская] дол.[ина] [Tajikistan, Zeravshan River valley, between Panjakent and Iori (= Yeri)] // Mor., [N]369 [handwritten by F. Morawitz] // Lectotype Mor., ♀, design. Pesenko [1]985 [ZMMU].

(3 ♀). 1 ♀, the same label as in the lectotype [ZMMU]; 1 ♀, , the same label as in the lectotype; 1 ♀, Заравшан.[ская] дол.[ина] [Zeravshan River valley] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor., ♀ [handwritten by F. Morawitz] // Paralectotypus Mor., design. Pesenko [1]985 [ZISP].

(Morawitz, 1876).

The lectotype designation by Warncke (1982: 111) is invalid because he labelled neither of the two females from “valle Serafshan” deposited in ZMMU.

Description of male. Blüthgen in Popov 1935: 362.

Israel, Turkey, Iraq, Iran, Afghanistan, Turkmenistan, Uzbekistan, Tajikistan (Pesenko 1986a, Ascher and Pickering 2020).

Morawitz, 1876

18BB406B-2C39-53C6-93DD-036BF75BDC62

Figure 26

Figure 26.

Morawitz, 1876, lectotype, male A habitus, lateral view and labels B metasoma, dorsal view C mesosoma, dorsal view D antenna. Scale bars: 1.0 mm (A), 0.5 mm (B–D).

Morawitz, 1876: 217 (key to females), 220 (key to males), 222, ♀, ♂.

Shakhimardan (Uzbekistan).

Usbekistan: Khodzha-Chiburgan gorge, near Shakhimardan Usbekistan]; Kyrgyzstan: Alay, Kichi-alay.

♂, 7.[VII.1871] // Шагимарданъ [Shakhimardan in the Uzbek enclave in the territory of Kyrgyzstan, Alai Ridge, ] // [N]334 // Lectotypus Mor., design. Astafurova et Proshchalykin, 2020 [ZMMU].

(8 ♀, 5 ♂). 1 ♂, the same label as in a lectotype // Mor., [N]334 [handwritten by F. Morawitz]; 1 ♀, 2 ♂, 26.[VI.1871] // Чибурганъ [Chiburgan] // [N]334; 1 ♀, 21.[VI.1871] // Чибурганъ [Chiburgan] // [N]334; 2 ♀, 28.[VII.1871] // Кичи-Алай [Kichi-Alay] // [N]334 [ZMMU]; 2 ♀, 1 ♂, Чибурганъ [Chiburgan] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // F. Mor. [handwritten by F. Morawitz]; 1 ♂, Кчи-Алай [Kchi-Alay] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // F. Mor. [handwritten by F. Morawitz]; 1 ♀, < golden circle> // 22.[VII.1871] // Алай [Alay] // Mor., Typ. [handwritten by F. Morawitz]; 1 ♀, 22.[VII.1871] // Алай [Alay] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor. [handwritten by F. Morawitz] // Paralectotypus Mor., design. Astafurova et Proshchalykin, 2020 [ZISP].

(Morawitz, 1876).

The lectotype designation by Warncke (1982: 81) is invalid because he labelled neither of the two females from “Shakhimardan” deposited in ZMMU.

Turkey, Afghanistan, southern Kazakhstan, Turkmenistan, Uzbekistan, Kyrgyzstan (Ebmer 1995, Murao et al. 2017).

Morawitz, 1876

5F934AAA-0C72-5C79-B6AC-56423DAD631B

Figure 27

Figure 27.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B metasoma, dorsal C view head, frontal view D mesosoma, dorsal view. Scale bars: 1.0 mm.

Morawitz, 1876: 218 (key), 238, ♀.

Bairkum (Chimkent Province, Kazakhstan).

Kazakhstan: Bayrakum [= Bairkum]; Tajikistan: Pendzhikent, Iori.

♀, designated by Pesenko 1986a: 140, 4.[V.1871] // Байракумъ [Kazakhstan, Chimkent Province, Bairkum, Syr-Darya River, ] // Mor., [N]359 [handwritten by F. Morawitz] // Lectotypus Mor., ♀, design. Pesenko [1]985 [ZMMU].

(5 ♀). 1 ♀, the same label as in the lectotype; 1 ♀, 30.[V.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley, Iori] // N[359] [ZMMU]; 1 ♀, Байракумъ [Bayrakum] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // F. Mor., ♀ [handwritten by F. Morawitz]; 2 ♀, 4.[V.1871] // Байракумъ [Bayrakum] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus Mor., design. Pesenko [1]985 [ZISP].

(Morawitz, 1876).

Description of male. Blüthgen 1929: 53, as .

The lectotype designation by Warncke (1982: 111) is invalid because he labelled neither of the two females from “Bayrakum” deposited in ZMMU.

Southern Kazakhstan, Turkmenistan, Tajikistan, Kyrgyzstan, China (Xinnjiang) (Pesenko 1986a, Murao et al. 2017).

Morawitz, 1876

534DA223-0245-5C8D-9836-BBBE2E30E5B8

Figures 28

Figure 28.

Morawitz, 1876, lectotype, female A habitus, lateral view and labels B head, frontal view C metasoma, dorsal view D mesosoma, dorsal view. Scale bars: 1.0 mm (A), 0.5 mm (B–D).

Morawitz, 1876: 216 (key), 227, ♀.

Samarkand (Uzbekistan).

Uzbekistan: Samarkand, Dshyuzak [=Jizzakh], Iskander [River]; Kyrgyzstan: Osh.

♀, designated by Blüthgen 1934a: 303, 7.[VII.1870] // Самаркандъ [Uzbekistan, Samarkand, ] // Mor., [N]342 [handwritten by F. Morawitz] // Mor., ♀, Lecto-Holotype, Blüthgen det. 1931 // Lecto-Type // Lectotypus Mor., design. Blüthgen 1934 [ZMMU].

(5 ♀). 1 ♀, 21.[VI.1870] // Искандеръ [Iskander] // [N]342; 1 ♀, 2.[VIII. 1871] // Ошъ [Osh] // [N]342; 2 ♀, 4.[VII.1869] // Самаркандъ [Samarkand] // [N]342; 1 ♀, 18.[VII.1870] // Джюзакъ [Dzhyuzak] // [N]342 // Paralectotype Mor. design. Blüthgen Astafurova> [ZMMU].

Morawitz, 1874 (synonymised by Ebmer 1988b: 576).

Russia (European part, North Caucasus, Crimea), Cyprus, Turkey, Iran, Afghanistan, Central Asia, Mongolia, north-western China (Astafurova and Proshchalykin 2017).

Morawitz, 1876

FDA0BA5F-415E-5762-AF30-281E7AD30236

Figure 29

Figure 29.

Morawitz, 1876, lectotype, female A habitus, lateral view B labels C metasoma, dorsal view D head, frontal view E mesosoma, dorsal view. Scale bars: 2.0 mm (A), 1.0 mm (C–E).

Morawitz, 1876: 218 (key), 240, ♀.

Bairkum (Chimkent Province, Kazakhstan).

Kazakhstan: Bayrakum.

♀, designated by Warncke 1982: 90, // 4.[V.1871] // Байракумъ [Kazakhstan, Chimkent Province, Bairkum, Syr-Darya River, ] // Mor., [N]362 [handwritten by F. Morawitz] // Lectotypus Mor., design. Warncke [19]82 [ZMMU].

Ebmer, 1972, replacement name for Morawitz, 1876 (nec Schenck, 1853).

Male unknown.

Southern Kazakhstan (Morawitz 1876, Mutao et al. 2017). The record from Azerbaijan (Aliyev et al. 2007) is doubtful and needs checking.

Morawitz, 1876

06CD3366-2926-552D-8046-EEA89F2366B4

Figure 30

Figure 30.

Morawitz, 1876, lectotype, female A head, frontal view B habitus, lateral view and labels C metasoma, dorsal view D mesosoma, dorsal view. Scale bars: 1.0 mm (B), 0.5 mm (A, C, D).

Morawitz, 1876: 217 (key to females), 220 (key to males), 223, ♀, ♂.

Jizzakh (Uzbekistan).

Uzbekistan: Katty-Kurgan [= Kattakurgan], Dzhyuzak [= Jizzakh], Karatyube [= Karatepa near Samarkand], Urgut, Sangy-Dzhuman; Kyrgyzstan: near Osh.

♀, designated by Blüthgen 1955: 17, 19.[VII.1870] // Джюзакъ [Uzbekistan, Dzhyuzak (= Jizzakh), ] // [N]337 // Mor., ♀, Lecto-Holotype, P. Blüthgen det. // Typus // Lectotypus Mor., 1876, design. Blüthgen, 1955 [ZMMU].

(1 ♀, 2 ♂). 1 ♂, the same labels as in the lectotype // Mor. ♂, lecto-Paratype, Blüthgen det.; 1 ♂, the same label, but 14.[VII.1870] // Mor. ♂, lecto-Holotype, Blüthgen det.; 1 ♀, 20. [VI.1869] // Катты-Курганъ [Katty-Kurgan] // Mor. ♀, lecto-Paratype, Blüthgen det. // Paralectotype Mor., design. Blüthgen 1955 Astafurova> [ZMMU].

Smith, 1853 (synonymised by Sakagami and Ebmer 1987: 326).

Southern Palaearctic and Oriental Regions. North Africa, Israel, Arabian Peninsula, Turkey, Central Asia, Iraq, Iran, Afghanistan, Pakistan, Mongolia, north-western China, India, Nepal, Myanma,Thailand, Sri Lanka (Pesenko 2006a).

Morawitz, 1876

FD313813-96D6-5A74-A2FE-EE46362F4513

Figure 31

Figure 31.

Morawitz, 1876, lectotype, female A head, frontal view B habitus, lateral view and labels C metasoma, dorsal view D mesosoma, dorsal view. Scale bars: 1.0 mm (B, C), 0.5 mm (A, D).

Morawitz, 1876: 218 (key), 250, ♀.

Samarkand (Uzbekistan).

Uzbekistan: Tashkent, Samarkand, Katty-Kurgan [Kattakurgan].

♀, 3.[III.1869] [Uzbekistan, Samarkand, // [sic!] Mor., [N]379 [handwritten by F. Morawitz] // Lectotypus Mor., design. Astafurova et Proshchalykin, 2020 [ZMMU].

(257 ♀). 26 ♀, 4., 20., 23., 30.[III.1869], 3., 11., 19.[IV.1869] // Самаркндъ [Samarkand]; 5 ♀, 28.[IV.1869] // Каттыкурганъ [Kattykurgan]; 222♀, 10., 11., 24., 26., 27., 28., [II.1871], 23., 24. [III.1871], 1., 2., 3., 5., 8., 10., 11.[IV.1871] // Ташкентъ [Tashkent] [ZMMU]; 1 ♀, 3.[IV.1871] // Ташкентъ [Tashkent] // к. Ф. Моравица // Mor. [handwritten by F. Morawitz]; 1 ♀, 5.[IV.1871] // Ташкентъ [Tashkent] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 2 ♀, 8.[IV.1871] // Ташкентъ [Tashkent] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz]; 1 ♀, 1.[IV.1871] // Ташкентъ [Tashkent] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Paralectotypus, Mor., design. Astafurova et Proshchalykin [ZISP].

(Brullé, 1832) (synonymised by Blüthgen 1926: 391).

Europe (except North), North Africa, Caucasus, Russia (East of European part, North Caucasus), Turkey, Syria, Jordan, Israel, Iraq, Iran, Afghanistan, Pakistan, Central Asia, Kazakhstan, north India, Nepal (Astafurova and Proshchalykin 2017).

The lectotype designation by Warncke (1982: 116) is invalid because he labelled none of the 222 females from Tashkent deposited in ZMMU.

Genus Latreille, 1804

Morawitz, 1876

2E2C1C94-0E05-5CEF-86BE-FD06A326124C

Figure 32

Morawitz, 1876: 257, ♀.

Shardara District of Turkistan Province (Kazakhstan).

Kazakhstan: coasts of Kosaral Lake; Uzbekistan: Khodzhaduk [= Khozyay-Dun].

♀, designation by Warncke 1992: 30, 24.IV.1871 [the original blue data label was damaged] // Косаралъ [Kazakhstan, “Kosaral Lake”, Shardara (= Chardara) District of Turkistan (= South-Kazakhstan) Province, ≈ ] // Mor. [handwritten by F. Morawitz] // Lectotypus, Warncke 1975 [ZMMU].

1 ♀, 21.[V.1869] // Заравшан.[ская] дол.[ина] [Zeravshan River valley] // Paralectotype Mor., design. Warncke Astafurova> [ZMMU].

(Linnaeus 1758) (synonymised by Blüthgen 1923b: 510).

North Africa, Europe (north to 63°), Israel, Jordan, Russia (east to Yakutia), Turkey, Iran, Pakistan, Central Asia, Kazakhstan, Mongolia, NW China, India (Warncke 1992, Bogusch and Straka 2012, Astafurova et al. 2019).

Morawitz, 1876

168597F0-F09A-5EC4-B44F-55A8A2EA0C4E

Figure 33

Morawitz, 1876: 256, ♀.

Shardara District of Turkistan Province (Kazakhstan).

Kazakhstan: coasts of Kosaral Lake; Kyzylkum [desert] near Chakany Well.

♀, designation by Warncke 1992: 24, 24.[IV.1871] // Косаралъ [Kazakhstan, “Kosaral Lake”, Shardara (= Chardara) District of Turkistan (= South-Kazakhstan) Province], ≈ // Mor. [handwritten by F. Morawitz] // Lectotypus, Warncke 1975 [ZMMU].

1 ♀, 28.[VI.1871] // Кызылъкумъ [Kyzylkum] // Paralectotype Mor., design. Warncke Astafurova> [ZMMU].

Morawitz, 1876.

Description of male. Meyer 1919: 126, as sensu Meyer (non Hagens 1882) (see Blüthgen 1924: 475).

South Kazakhstan, Central Asia, China (Gansu, Xinjiang) (Astafurova et al. 2018a, b, 2020).

Morawitz, 1876

871E77F6-9DDB-5017-9259-074246B9574D

Figure 34

Morawitz, 1876: 255, ♀, ♂.

Bairkum (Chimkent Province, Kazakhstan).

Kazakhstan: Bayrakum [Bairkum]; steppe between Syr-Darya River and Tashkent.

♀, designated by Warncke 1992: 24, 17. [V.1871] // Байракумъ [Kazakhstan, Chimkent Province, Bairkum, Syr-Darya River, ] // F. Moraw., ♀ [handwritten by F. Morawitz] // Lectotypus Warncke, 1975 [ZMMU].

(2 ♀, 1 ♂). 1 ♀, Байракумъ [Bairkum] // кол.[лекция] Ф. Моравица [Collection of F. Morawitz] // F. Moraw., ♀ [handwritten by F. Morawitz] // F. Morawitz det., Typ.; 1 ♀, 20.[V.1871] // Степь м.[ежду] С.[ыр] д.[арьей] и Т.[ашкентом] [Steppe between Syrdarya River and Tashkent] // Mor., ♂ [handwritten by F. Morawitz] // Paralectotypus Mor., design. Warncke, [19]92 Astafurova> [ZMMU]; 1 ♀, the same labels [ZISP].

Lepeletier, 1825 (synonymised by Warncke 1992: 24).

North Africa, the Arabian Peninsula, Israel, Jordan, South Europe, Russia (south of European part), Turkey, Caucasus, Iran, Pakistan, Central Asia, Kazakhstan, NW China (Astafurova et al. 2019).

Genus

Morawitz, 1876

D6C3CB5F-DCAE-5E9E-8BD0-AAAB4FB031EA

Figure 35

Morawitz, 1876: 259, ♀, ♂.

Jizzakh (Uzbekistan).

Uzbekistan: Samarkand, Dzhyuzak.

♀, designated by Warncke 1976: 104, 20.[VII.1871] // Джюзакъ [Uzbekistan, Dzhyuzak (= Jizzakh), ] // Mor. [handwritten by F. Morawitz] // Lectotypus, Warncke 1975 [ZMMU].

(2 ♂). 1 ♂, the same labels as in the lectotype // Paralectotypus, Mor., design. Warncke [ZISP]; 1 ♂, 8.[VII.1869] // Самаркандъ [Samarkand] // Paralectotypus Mor., design. Warncke [ZMMU].

(Morawitz, 1876).

North Africa, Saudi Arabia, Turkey, Azerbaijan, Kazakhstan, Central Asia, Iraq, Iran, Afghanistan, Pakistan, India (Astafurova 2014).

Morawitz, 1876

65869E10-D5BC-5910-AE38-7623BFCCFE47

Figure 36

Morawitz, 1876: 261, ♀.

Zeravshan River valley (Uzbekistan).

Uzbekistan: “Aykul Lake” in Zeravshan River valley.

♀, designated by Warncke 1976: 106, 5.[VIII.1869] // Заравш[анская]. дол.[ина] [Uzbekistan, Zeravshan River valley, Aykul Lake near Chelek, ] // Mor. // Lectotypus, Warncke 1975 [ZMMU].

(Morawitz, 1876).

Description of male. Morawitz 1893: 79.

Turkey, Central Asia, Kazakhstan (Astafurova 2014).

Genus

Morawitz, 1876

A04B3F23-E067-5F3E-8AE2-0316930DAE54

Figure (see Astafurova and Proshchalykin 2019: 56, figs 51a–e)

Morawitz, 1876: 215, ♂.

Bairamali (Turkmenistan).

Uzbekistan: Samarkand.

♂, окр.[естности] Самарканда [Uzbekistan, Samarkand, ], 13.VI[1869], lost (also see Pesenko 1983: 168–170).

Neotype.

♂, designated by Pesenko 1983: 168, Байрам-Али Закасп.[ийcкая] обл.[асть] [Turkmenistan, Maryi Province, Bairamali, ], 14.VII.[1]928, В. Гуссаковский [V. Gussakovskij leg.] // m., ♂, Blüthgen det. 1931 // Neotypus Mor., ♂, design. Pesenko [1]980 [handwritten by Yu. Pesenko] // Zoological Institute St. Petersburg, INS_HYM_0000142 [ZISP].

Morawitz, 1876.

Description of female: Blüthgen 1925b: 45, as (synonymised by Blüthgen 1934c: 253).

Asia Minor, Afghanistan, Armenia, Central Asia (Astafurova and Proshchalykin 2019).

Morawitz, 1876

12CF6959-1988-5CDD-8935-CB84BD38C483

Figure (see Astafurova and Proshchalykin 2019: 68, figs 62a–e).

Morawitz, 1876: 214, ♀, ♂.

Samarkand (Uzbekistan).

Tajikistan: Murzarabat; Uzbekistan: Sokh, Samarkand.

♂, designated by Pesenko 1983: 174, 5.[VII.1870] // Самаркандъ [Uzbekistan, Samarkand, ] // Mor., Typ. [handwritten by F. Morawitz] // Lectotypus Mor., ♂, design. Pesenko, 1976 // Zoological Institute St. Petersburg, INS_HYM_0000131 [ZISP].

(5 ♂). 1 ♂, 28.[VI.1871] // Сохъ [Sokh] // n. sp. F. Morawitz det.; 1 ♂, 8.[VII.1870] // Самаркандъ [Samarkand] // Mor. [handwritten by F. Morawitz] [ZMMU]; 3 ♂, Сохъ [Sokh] // к.[оллекция] Ф. Моравица [Collection of F. Morawitz] // Mor. // Paralectotypus ♂ Mor., design. Pesenko, 1976 [ZISP].

Morawitz, 1876.

North Africa, Central Asia, Iran, Pakistan (Astafurova and Proshchalykin 2019).