A cybercatalogue of American sand fly types (Diptera, Psychodidae, Phlebotominae) deposited at the Natural History Museum, London.

BACKGROUND: Sand flies (Diptera, Psychodidae, Phlebotominae) are biting flies involved in the transmission of pathogens, including the protozoan parasite Leishmania amongst human and non-human animals (Rangel and Lainson 2009). NEW INFORMATION: A total of 60 species of American Phlebotominae (Diptera: Psychodidae), distributed amongst 16 genera were studied. A checklist of the primary and secondary type specimens held at the Natural History Museum, London (NHMUK), is given and 968 photographs of the specimens and their labels are made available on a Scratchpads website http://phlebotominaenhmtypes.myspecies.info.

Introduction

There are approximately 1,000 valid described species of sand flies in the world, of which 530 are known to occur in the Americas (Shimabukuro et al. 2017).

The aim of this work is to provide a checklist of types of the Neotropical and Nearctic regions held in the collections of the Natural History Museum, London (NHMUK formerly BMNH), together with electronic links to, scans of whole slides, plus detail images of the taxonomically informative morphological structures. This work updates the American portion of the 1990 checklist of types deposited at the NHMUK (Townsend and Lane 1990).

History

Owing to the small size and morphological homogeneity of phlebotomine sand flies, few species were described before the turn of the 20th century. Indeed little progress in their taxonomy was made until Adler and Theodor (1926) and Theodor (1932) drew attention to the utility of various internal structures (cibarium & spermathecae) for species differentiation. Currently, taxonomically valid species descriptions of phlebotomine sand flies still rely on adult morphology. Although newer molecular techniques are increasingly used for identifying species (Zapata et al. 2012), they have yet to be formally used as characters in species descriptions. Type specimens are confined to the adult stage, with few taxa described in the immature stages.

Significant donations to the NHMUK collection have come from: (i) Graham Bell Fairchild who worked on the Panamanian fauna in the 1950s and 60s; (ii) a number of early types described by Robert Newstead in the 1920s when he was a Professor at the Liverpool School of Tropical Medicine; (iii) and most significantly, the collection of Oskar Theodor, which the museum purchased in 1981 after his retirement from the University of Jerusalem. The NHMUK has also absorbed the specimen collections of the London School of Tropical Medicine and Hygiene (LSHTM).

The main body of the NHMUK collection was assembled by Dr D.J. Lewis who, though not a member of staff, was based at the museum for 30 years (1956-1986) whilst in the employ of the Medical Research Council in London. All his material was mounted in Berlese media which is not a permanent mountant. There are many examples of deteriorating Lewis slides in the NHMUK collection (Fig. 1), as well as instances where Lewis remounted other authors' type specimens into Berlese. His distinctive handwriting and messy slides, often bearing multiple specimens, are instantly recognisable.

Figure 1.

Example of an R. Newstead type remounted into Berlese by D. J. Lewis, showing the typical darkening of the mountant as it deteriorates.

Taxonomic treatment

taxonomy has a long history of differences in opinion amongst experts and various conflicting classification schemes. The current situation is no exception, with both classification of Young and Duncan (1994), which is based on the previous work of Lewis et al. (1977) and that of Galati (2003b) in use at the same time, for the New World fauna. For a comparison of how these two classifications relate to each other see Fig. 2.

Currently, the NHMUK collections are arranged according to Young and Duncan (1994), but we have used the Galati (2003b) naming systems here to reflect the current knowledge of the group.

In their guide to identification and distribution, Young and Duncan (1994) adopt the classification of Lewis et al. (1977) with only minor changes. They make it clear that their arrangement of taxa does not represent a phylogenetic analysis of the group and, aside from a few comments in the notes, give no information on the relationships between their subgeneric groups. It is clear that their groupings have been arrived at through a phenetic review of morphological characters with little focus on identifying any common ancestry between groups.

Lewis et al. (1977) proposed their classification at a time when those working on sand flies were faced with the choice between two family names, several subfamily names and, for some species, three possible generic names. Their aim was to create a stable general framework with flexible subdivisions into which, recent and future ideas on the evolutionary relationships within the group, could be assimilated. They present a review of the early literature on the group and, with reference to the International Code of Zoological Nomenclature (ICZN), proposed the use of the family name Newman 1834 and the subfamily name Rondani 1840, proposals that have been more or less universally accepted. Their proposal for the use of just five genera (, , , and ) now seems less successful.

At that time, 1977, around 600 sand fly species had been described, with 290 of these in the American genus (Young and Duncan 1994). The current checklist for American sand flies alone (Shimabukuro et al. 2017) stands at 530 and use of the Lewis et al. (1977) concept of the genus would place 495 of these taxa into a single genus. It now seems likely that, by placing the emphasis on a stable and simple classification over one that best represented the evolutionary history of the group, Lewis et al. (1977) created a number of polyphyletic groups, particularly their genus .

The classification of Galati (1995) and Galati (2003a) is more complex with the genus , as defined in Lewis et al. (1977) and Young and Duncan (1994), split into 23 genera and 22 sub-generic groupings and has been increasingly utilised by authors working on the American fauna.

The study by Galati (1995) represented the first comprehensive attempt to perform a phylogenetic analysis of the American , comprising a cladistic analysis of 88 adult morphology characters, using the subfamily as the principal outgroup. The classification has since been updated and represents our current best understanding of the evolutionary relationships within this important group (Galati 2003a, Galati 2003b).

There are some nomenclatural issues with the Galati classification as it stands at the present moment (see Shimabukuro et al. 2017). Within the classification a number of family group and genus group names are attributed to Artemiev from his 1991 work (Artemiev 1991). In this work, Artemiev does not accompany any of his new names with a description that states in words characters that are purported to differentiate the taxa, thus all his new names do not comply with ICZN Article 13.1.1 http://www.iczn.org/iczn/index.jsp and should be considered nomina nuda. Likewise, there are a large number of family group and genus group names in the classification attributed to Galati (1995), where she also fails to accompany any of her new names with a description that can be used to distinguish them. Galati lists the characters used, gives a polarised transformation series and maps the characters and taxa on to a cladogram, but this is not sufficient under the code to render the new names from her 1995 work available. In 2003, Galati updated her 1995 classification (Galati 2003b, Galati 2003a), this 2003 work contains extensive identification keys for all groups and any unavailable names should take authorship and date from this later act of establishment under ICZN articles 50 and 21.

Materials and methods

Checklist

The list contains information compiled from the examination of Nearctic and Neotropical species of sand flies from the Collection at the NHMUK.

Species are listed alphabetically, with genus and subgenus designations according to Shimabukuro et al. 2017. Links to the scratchpad entry and images for each specimen are given in the eventRemarks section of the checklist.

Readers familiar with the classification of Young and Duncan (1994) should refer to the scratchpads website http://phlebotominaenhmtypes.myspecies.info where the specimens have been arranged according to both classifications.

We have excluded from this catalogue two NHMUK specimens bearing “type” labels, for which we can find no record of the given species name in the literature. One labelled as the holotype of (author Feliciangeli), the other labelled as a paratype of L. patiñoi (authors Mangabeira & Galindo). It seems probable that these are unpublished manuscript names and are therefore unavailable under article 8 of the ICZN.

Specimen Images

Photographs were taken in the Sackler Biodiversity Imaging Laboratory (SBIL), at the NHMUK, with an Olympus BX63-CBH microscope equipped with a DP73 camera and differential interference contrast (DIC) illumination. The images were edited in the Cellsens software and stacked images were produced by HeliconFocus 6 software.

Whole slide scans were generated using a SatScan instrument at x5 magnification and images were cropped and processed using the Inselect software, version 0.1.35, available from the Natural History Museum https://naturalhistorymuseum.github.io/inselect.

Data resources

Bibliography

Most of the original species descriptions are still under copyright to their original publisher. The bibliographical and historical database of scientific articles on leishmaniases and sand flies hosted by the Laboratoire de Parasitologie at the University of Montpellier is an extremely useful resource http://www.leishpub.univ-montp1.fr.

The specimen images are available on the NHMUK data portal: https://doi.org/10.5519/0051226.

The checklist, specimen data and specimen images are available on a Scratchpads website http://phlebotominaenhmtypes.myspecies.info.

Checklists

Fairchild & Hertig, 1952

Fairchild & Hertig, 1952 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722073; sex: Female; Taxon: scientificName: ; acceptedNameUsage: ; Location: country: Panama; stateProvince: Colón; locality: Mojinga swamp; Event: eventDate: 09-05-51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722074; sex: Male; Taxon: scientificName: ; acceptedNameUsage: ; Location: country: Panama; stateProvince: Panamá; locality: Rio Majé, Charco el Toro; Event: eventDate: 03-22-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Distribution

Colombia, Costa Rica, Ecuador, Nicaragua, Panama

Notes

Synonym of (Rodríguez, 1950) Rodríguez 1950. Synonymised by Fairchild and Hertig 1958b.

Fairchild & Hertig, 1952

Fairchild & Hertig, 1952 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721992; sex: Female; Taxon: scientificName: ; Location: country: Panama; stateProvince: Colón; locality: Mojinga swamp, nr. Gatún; Event: eventDate: 08-21-51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721993; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Chiriquí; municipality: Barú; locality: Puerto Armuelles; Event: eventDate: 06-09-51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Belize, Bolivia, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama

Valid species in .

Rozeboom, 1942

Rozeboom, 1942 (

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722068; sex: Female; Taxon: scientificName: ; Location: country: Venezuela; stateProvince: Bolívar; municipality: Gran Sabana; locality: Gran Sabana; Event: year: 1941; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722120; Taxon: scientificName: ; Location: country: Venezuela; stateProvince: Bolívar; municipality: Gran Sobana; locality: Gran Sobana; Event: year: 1941; month: 12; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722121; sex: Female; Taxon: scientificName: ; Location: country: Venezuela; stateProvince: Bolívar; municipality: Gran Sobana; locality: Gran Sobana; Event: year: 1941; month: 12; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722122; sex: Female; Taxon: scientificName: ; Location: country: Venezuela; stateProvince: Bolívar; municipality: Gran Sobana; locality: Gran Sobana; Event: year: 1941; month: 12; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722123; Taxon: scientificName: ; Location: country: Venezuela; stateProvince: Bolívar; municipality: Gran Sobana; locality: Gran Sobana; Event: year: 1941; month: 12; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Brazil, Costa Rica, French Guiana, Panama, Peru, Venezuela

Valid species in .

Feliciangeli, Ordoñez & Manzanilla, 1984

Feliciangeli, Ordoñez & Manzanilla, 1984 (

Type status: Holotype. Occurrence: catalogNumber: BMNHE1722088; sex: Male; Taxon: scientificName: ; Location: country: Venezuela; stateProvince: Trujillo; locality: Loma Abajo; Event: eventDate: 06-01-70; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Venezuela

Valid species in .

Mangabeira, 1942

Mangabeira, 1942 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722081; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Belém; locality: Aurá, em buraco de tatu.; Event: year: 1940; month: 8; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Brazil, Colombia, Peru, Surinam, Venezuela

Valid species in .

Fairchild & Hertig, 1951

Fairchild & Hertig, 1951 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1251325; sex: Female; Taxon: scientificName: ; Location: country: Panama; stateProvince: Chiriquí; locality: La Victoria, Cerro Jefe; Event: eventDate: 08-29-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Allotype. Occurrence: catalogNumber: BMNHE1251326; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Chiriquí; locality: La Victoria, Cerro Jefe; Event: eventDate: 08-30-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Belize, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Honduras, Mexico, Nicaragua, Panama, Surinam

Valid species in .

Fairchild & Hertig, 1961

Fairchild & Hertig, 1961 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722010; sex: Female; Taxon: scientificName: ; Location: country: Panama; stateProvince: Chiriquí; locality: El Volcan; Event: eventDate: 03-23-54; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Panama

Valid species in .

Larrousse, 1920

Larrousse, 1920 (

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722011; sex: Female; Taxon: scientificName: ; Location: country: Brazil; locality: Albuquerque; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722043; sex: Male; Taxon: scientificName: ; Location: country: Brazil; locality: Albaquerque; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Argentina, Bolivia, Brazil

Valid species in .

Lewis, 1967

Lewis, 1967 (

Type status: Holotype. Occurrence: catalogNumber: BMNHE1722028; sex: Male; Taxon: scientificName: ; Location: country: Cayman Islands; stateProvince: Cayman Brac; locality: Bamboo Bay; Event: eventDate: 12-18-64; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722050; sex: Female; Taxon: scientificName: ; Location: country: Cayman Islands; stateProvince: Cayman Brac; locality: Bamboo Bay; Event: eventDate: 18/12/64; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722051; sex: Female; Taxon: scientificName: ; Location: country: Cayman Islands; stateProvince: Cayman Brac; locality: Bamboo Bay; Event: eventDate: 18/12/64; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722106; sex: Female; Taxon: scientificName: ; Location: country: Cayman Islands; stateProvince: Cayman Brac; locality: Stake Bay; Event: eventDate: 12-16-64; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722107; sex: Male; Taxon: scientificName: ; Location: country: Cayman Islands; stateProvince: Cayman Brac; locality: Bamboo Bay; Event: eventDate: 12-18-64; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722108; sex: Male; Taxon: scientificName: ; Location: country: Cayman Islands; stateProvince: Cayman Brac; locality: Stake Bay; Event: eventDate: 12-16-64; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722109; sex: Male; Taxon: scientificName: ; Location: country: Cayman Islands; stateProvince: Cayman Brac; locality: Stake Bay; Event: eventDate: 12-16-64; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722110; sex: Female; Taxon: scientificName: ; Location: country: Cayman Islands; stateProvince: Cayman Brac; locality: Bamboo Bay; Event: eventDate: 12-18-64; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722111; sex: Male; Taxon: scientificName: ; Location: country: Cayman Islands; stateProvince: Cayman Brac; locality: Bamboo Bay; Event: eventDate: 12-18-64; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722112; sex: Female; Taxon: scientificName: ; Location: country: Cayman Islands; stateProvince: Cayman Brac; locality: Stake Bay; Event: eventDate: 12-16-64; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722113; sex: Male; Taxon: scientificName: ; Location: country: Cayman Islands; stateProvince: Cayman Brac; locality: Bamboo Bay; Event: eventDate: 12-18-64; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Cayman Islands

Valid species in .

Fairchild & Trapido, 1950

Fairchild & Trapido, 1950 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722029; sex: Female; Taxon: scientificName: ; Location: country: Haiti; stateProvince: Nord; locality: La Voute, 4.8 miles from Cap-Haïtien on Limbe Rd.; Event: eventDate: 06-09-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722030; sex: Male; Taxon: scientificName: ; Location: country: Haiti; stateProvince: Nord; municipality: le Cap-Haïtien; locality: Milot, le Cap-Haïtien; Event: eventDate: 06-09-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722031; sex: Female; Taxon: scientificName: ; Location: country: Dominican Republic; stateProvince: Hato Mayor; municipality: Sabana de la Mar; locality: Cacao plantation 6 km South of Sabana de la Mar; Event: eventDate: 06-12-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722032; sex: Female; Taxon: scientificName: ; Location: country: Haiti; stateProvince: Nord; municipality: le Cap-Haïtien; locality: Ruins of Pauline Bonaparte's palace East of town; Event: eventDate: 06-09-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722033; sex: Male; Taxon: scientificName: ; Location: country: Haiti; stateProvince: Nord; municipality: le Cap-Haïtien; locality: Ruins of Pauline Bonaparte's palace East of town; Event: eventDate: 06-09-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722034; sex: Male; Taxon: scientificName: ; Location: country: Dominican Republic; stateProvince: Santiago; locality: Santiago area; Event: eventDate: 06-14-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Dominican Republic, Haiti

Valid species in .

Fairchild & Trapido, 1950

Fairchild & Trapido, 1950 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722037; sex: Male; Taxon: scientificName: ; Location: country: Jamaica; stateProvince: Kingston; locality: Rockfort, St Andrew parish, Kingston; Event: eventDate: 05-20-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Jamaica

Valid species in .

Fairchild & Hertig, 1948

Fairchild & Hertig, 1948 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722035; sex: Female; Taxon: scientificName: ; Location: country: Mexico; stateProvince: México; municipality: Zumpango; locality: Zumpango; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722036; sex: Female; Taxon: scientificName: ; Location: country: Guatemala; stateProvince: Escuintla; municipality: San José; locality: San José; Event: eventDate: 06-03-45; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Belize, Guatemala, Mexico

Valid species in .

Fairchild & Hertig, 1948

Fairchild & Hertig, 1948 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722040; sex: Female; Taxon: scientificName: ; Location: country: Puerto Rico; stateProvince: Lares; locality: Lares; Event: eventDate: 07-03-47; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Puerto Rico

Valid species in .

Mangabeira, 1942

Mangabeira, 1942 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721985; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Belém; locality: Asura; Event: year: 1940; month: 8; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Brazil, Peru

Valid species in .

Fairchild & Trapido, 1950

Fairchild & Trapido, 1950 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722041; sex: Female; Taxon: scientificName: ; Location: country: Cuba; stateProvince: Camagüey; locality: Camagüey; Event: eventDate: 05-30-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722042; sex: Male; Taxon: scientificName: ; Location: country: Cuba; stateProvince: Pinar del Río; municipality: Viñales; locality: San Vincente, Viñales; Event: eventDate: 06-04-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722114; sex: Female; Taxon: scientificName: ; Location: country: Cuba; stateProvince: Camagüey; locality: spelling on slide "Camaguay"; Event: eventDate: 05-30-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722115; Taxon: scientificName: ; Location: country: Cuba; stateProvince: Matanzas; locality: Ceiba Mocha; Event: eventDate: 06-02-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722116; sex: Male; Taxon: scientificName: ; Location: country: Cuba; stateProvince: Camagüey; locality: spelling on slide "Camaguay"; Event: eventDate: 05-30-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Cuba, United States of America

Valid species in .

Fairchild & Hertig, 1961

Fairchild & Hertig, 1961 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1251321; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Colón; locality: Juan Mina; Event: eventDate: 11-29-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1251322; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Panamá; locality: Rio Majé, Charco el Toro; Event: eventDate: 03-22-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1251323; sex: Female; Taxon: scientificName: ; Location: country: Panama; locality: Canal zone, Camp Piña; Event: eventDate: 03-06-54; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1251324; sex: Female; Taxon: scientificName: ; Location: country: Panama; stateProvince: Colón; locality: Mojinga swamp; Event: eventDate: 07-24-54; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Belize, Brazil, Colombia, Costa Rica, Ecuador, Mexico, Nicaragua, Panama, Venezuela

Valid species in .

Vargas & Díaz-Nájera, 1953

Vargas & Díaz-Nájera, 1953 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722142; sex: Female; Taxon: scientificName: ; Location: country: Mexico; stateProvince: Chiapas; locality: Finca El Vergel; Event: eventDate: 05-28-35; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722143; sex: Female; Taxon: scientificName: ; Location: country: Mexico; stateProvince: Chiapas; locality: Finca El Vergel; Event: eventDate: 05-28-35; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Belize, Guatemala, Mexico

Valid species in .

Fairchild & Trapido, 1950

Fairchild & Trapido, 1950 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722070; sex: Female; Taxon: scientificName: ; Location: country: Jamaica; stateProvince: Manchester; locality: Melrose Hill, nr. Williams Field; Event: eventDate: 05-25-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722071; sex: Male; Taxon: scientificName: ; Location: country: Jamaica; stateProvince: Manchester; locality: Melrose Hill, nr. Williams Field; Event: eventDate: 05-25-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722117; sex: Female; Taxon: scientificName: ; Location: country: Jamaica; stateProvince: Manchester; locality: Melrose Hill, nr. Williams Field.; Event: eventDate: 05-25-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722118; sex: Male; Taxon: scientificName: ; Location: country: Jamaica; stateProvince: Saint Catherine; locality: nr. Springvale; Event: eventDate: 05-21-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722119; Taxon: scientificName: ; Location: country: Jamaica; stateProvince: Manchester; locality: Manchester area; Event: eventDate: 05-25-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Jamaica

Valid species in .

Fairchild & Hertig, 1952

Fairchild & Hertig, 1952 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722075; sex: Female; Taxon: scientificName: ; Location: country: Panama; stateProvince: Panamá; locality: Pacora; Event: eventDate: 08-13-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722076; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Panamá; locality: Pacora; Event: eventDate: 08-13-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Brazil, Colombia, Costa Rica, Ecuador, Panama, Venezuela

Valid species in .

Nuñez-Tovar, 1924

Nuñez-Tovar, 1924 (

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722089; sex: Male; Taxon: scientificName: ; Location: country: Venezuela; Event: year: 1922; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722091; sex: Male; Taxon: scientificName: ; Location: country: Venezuela; Event: year: 1922; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Colombia, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Peru, Venezuela

Valid species in .

Abonnenc & Chassignet, 1948

Abonnenc & Chassignet, 1948 (

Type status: Holotype. Occurrence: catalogNumber: BMNHE1722138; sex: Male; Taxon: scientificName: ; Location: country: French Guiana; locality: Baduel; Event: year: 1947; month: 12; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Brazil, French Guiana

Valid species in .

Sherlock, 1962

Sherlock, 1962 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722096; sex: Male; Taxon: scientificName: ; acceptedNameUsage: ; Location: country: Colombia; stateProvince: Santander; municipality: San Vicente de Chucurí; locality: Soledad, San Vicente de Chucurí; Event: eventDate: 08-16-74; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722097; sex: Male; Taxon: scientificName: ; acceptedNameUsage: ; Location: country: Colombia; stateProvince: Santander; municipality: San Vicente de Chucurí; locality: Soledad, San Vicente de Chucurí; Event: eventDate: 16/08/44; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Brazil, Colombia, Ecuador, French Guiana, Panama, Peru, Trinidad and Tobago, Venezuela

Synonym of (Newstead, 1914). Synonymised by Young 1979.

Ogusuku, Canales & Pérez, 1997

Ogusuku, Canales & Pérez, 1997 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722012; sex: Male; Taxon: scientificName: ; Location: country: Peru; stateProvince: Huánuco; municipality: Huamalíes; locality: Monzon, Paucacu; Event: eventDate: 05-22-97; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722013; sex: Female; Taxon: scientificName: ; Location: country: Peru; stateProvince: Huánuco; municipality: Huamalíes; locality: Monzon, Paucacu; Event: eventDate: 05-22-97; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Peru

Valid species in .

Fairchild & Hertig, 1961

Fairchild & Hertig, 1961 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721997; sex: Female; Taxon: scientificName: ; acceptedNameUsage: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: Almirante; Event: eventDate: 06-19-51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721998; sex: Female; Taxon: scientificName: ; acceptedNameUsage: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: Almirante; Event: eventDate: 06-19-51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Colombia, Costa Rica, Ecuador, Panama, Peru

Synonym of (Blancas, 1959) Blancas 1959. Synonymised by Christensen and Rutledge 1973.

Fairchild & Hertig, 1957

Fairchild & Hertig, 1957 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722007; sex: Male; Taxon: scientificName: ; Location: country: Panama; locality: Cerro Campana; Event: eventDate: 04-24-52; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722008; sex: Female; Taxon: scientificName: ; Location: country: Panama; locality: Cerro Campana; Event: eventDate: 02-10-51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Colombia, Costa Rica, Ecuador, Mexico, Panama, Peru

Valid species in .

Young & Arias, 1977

Young & Arias, 1977 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721994; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Amazonas State; locality: Manaus, Estrada Torquato Tapajos, km 54 (east of Manaus); Event: eventDate: 10-21-75; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721995; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Amazonas State; locality: Manaus, Estrada Torquato Tapajos, km 54 (east of Manaus); Event: eventDate: 10-21-75; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Brazil, Venezuela

Valid species in .

Fairchild & Hertig, 1958

Fairchild & Hertig, 1958 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721986; sex: Male; Taxon: scientificName: ; Location: country: Panama; Event: eventDate: 06-20-56; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721987; sex: Male; Taxon: scientificName: ; Location: country: Panama; Event: eventDate: 06-20-56; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721988; sex: Male; Taxon: scientificName: ; Location: country: Panama; Event: eventDate: 06-20-56; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721989; sex: Male; Taxon: scientificName: ; Location: country: Panama; Event: eventDate: 06-20-56; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721990; sex: Male; Taxon: scientificName: ; Location: country: Panama; Event: eventDate: 06-20-56; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722047; sex: Male; Taxon: scientificName: ; Location: country: Panama; locality: K-19; Event: eventDate: 07-24-56; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Colombia, Costa Rica, Panama

Valid species in .

Fraiha & Ward, 1974

Fraiha & Ward, 1974 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722077; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; locality: km 46 Altamira to Itaituba road, Transamazonia highway; Event: eventDate: 08-21-71; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722078; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; locality: km 46 Altamira to Itaituba road, Transamazonia highway; Event: year: 1971; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722079; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; locality: km 46 Altamira to Itaituba road, Transamazonia highway; Event: eventDate: 08-21-71; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722080; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; locality: km 46 Altamira to Itaituba road, Transamazonia highway; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Brazil, Peru

Valid species in .

Feliciangeli, Ordoñez & Fernández, 1984

Feliciangeli, Ordoñez & Fernández, 1984 (

Type status: Holotype. Occurrence: catalogNumber: BMNHE1721984; sex: Female; Taxon: scientificName: ; Location: country: Venezuela; stateProvince: Cojedes; locality: La Vāquira; Event: eventDate: 06-20-77; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Venezuela

Valid species in .

Ogosuku, Perez, Davies & Villaseca, 1996

Ogosuku, Perez, Davies & Villaseca, 1996 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721980; sex: Female; Taxon: scientificName: ; Location: country: Peru; stateProvince: Piura; municipality: Huancabamba; locality: El Higueron; Event: eventDate: 02-21-93; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721981; sex: Male; Taxon: scientificName: ; Location: country: Peru; stateProvince: Piura; municipality: Huancabamba; locality: El Higueron; Event: eventDate: 05-23-92; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Peru

Valid species in .

Ogusuku, Canales & Pérez, 1997

Ogusuku, Canales & Pérez, 1997 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722014; sex: Female; Taxon: scientificName: ; Location: country: Peru; stateProvince: Huánuco; municipality: Huamalíes; locality: Monzon, Paucacu; Event: eventDate: 05-22-97; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722026; sex: Male; Taxon: scientificName: ; Location: country: Peru; stateProvince: Huánuco; municipality: Huamalíes; locality: Monzon, Paucacu; Event: eventDate: 05-22-97; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Peru

Valid species in .

Fairchild & Hertig, 1961

Fairchild & Hertig, 1961 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722052; sex: Female; Taxon: scientificName: ; Location: country: Honduras; stateProvince: Atlántida; municipality: Tela; locality: Lancetilla Valley, Tela; Event: eventDate: 06/01/54; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722054; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: Finca Nievecita, Almirante; Event: eventDate: 17/06/50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722055; sex: Female; Taxon: scientificName: ; Location: country: Honduras; stateProvince: Atlántida; municipality: Tela; locality: Lancetilla Valley, Tela; Event: eventDate: 23/12/53; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Brazil, Costa Rica, French Guiana, Guatemala, Honduras, Nicaragua, Panama, Venezuela

Valid species in .

Young, Pérez & Romero, 1985

Young, Pérez & Romero, 1985 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722098; sex: Male; Taxon: scientificName: ; Location: country: Peru; stateProvince: Lima; locality: Cocachacra; Event: eventDate: 01/05/63; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Peru

Valid species in .

Young & Arias, 1982

Young & Arias, 1982 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722066; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Amazonas; locality: 43km NE Manaus, at Campinas; Event: eventDate: 03-21-79; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722067; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Amazonas; locality: 25km E of Manaus, Ducke Reserve; Event: eventDate: 16-19/03/1979; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Brazil, Peru

Valid species in .

Vargas & Díaz-Nájera, 1959

Vargas & Díaz-Nájera, 1959 (

Type status: Allotype. Occurrence: catalogNumber: BMNHE1251319; sex: Female; Taxon: scientificName: ; Location: country: Mexico; stateProvince: Tabasco; municipality: Teapa; locality: Sta A-0; Event: eventDate: 12-16-53; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua

Valid species in .

(Fairchild & Hertig, 1961)

Fairchild & Hertig, 1961 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722053; sex: Female; Taxon: scientificName: ; Location: country: Panama; stateProvince: Chiriquí; municipality: Renacimiento; locality: Rio Candela, Chiriquí; Event: eventDate: 11/08/51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Costa Rica, Panama

Valid species in .

Shannon, 1929

Shannon, 1929 (

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722139; sex: Male; Taxon: scientificName: ; Location: country: Peru; stateProvince: Lima; locality: Matucana; Event: eventDate: 06-19-28; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Peru

Valid species in .

Hertig, 1948

Hertig, 1948 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722044; sex: Female; Taxon: scientificName: ; Location: country: Peru; stateProvince: Lima; municipality: Huarochiri; locality: Acequia cueva, Matucana; Event: eventDate: 08-25-45; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722045; sex: Female; Taxon: scientificName: ; Location: country: Peru; stateProvince: Lima; municipality: Huarochiri; locality: Rio Rimac, Matucana; Event: month: 8; day: 23; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Ecuador, Peru

Valid species in .

Fairchild & Hertig, 1961

Fairchild & Hertig, 1961 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722103; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Chiriqui; locality: Lagunas del Volcán; Event: eventDate: 05-19-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722104; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Chiriqui; locality: Sta. Clara, El Volcan; Event: eventDate: 08-11-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Colombia, Costa Rica, Panama, Peru, Venezuela

Valid species in .

Damasceno & Arouck, 1956

Damasceno & Arouck, 1956 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721996; sex: Male; Taxon: scientificName: ; Location: country: Brazil; locality: Rio Acará; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Brazil, French Guiana, Venezuela

Valid species in .

Osorno, Osorno-Mesa & Morales, 1972

Osorno, Osorno-Mesa & Morales, 1972 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722092; sex: Male; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Santander; municipality: Socorro; Event: eventDate: 12-09-67; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722093; sex: Female; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Santander; municipality: Barbosa; Event: eventDate: 10-30-68; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722129; sex: Female; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Santander; municipality: Barbosa; Event: eventDate: 10-30-68; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722130; sex: Female; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Santander; municipality: Güepsa; Event: eventDate: 06-22-67; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Colombia

Valid species in .

Ready & Fraiha, 1981

Ready & Fraiha, 1981 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1251320; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; locality: Altamira-Itaituba rd km 43; Event: eventDate: 08-21-71; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722063; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; locality: km 43 Altamira to Itaituba rd; Event: eventDate: 08-21-71; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722064; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; locality: km 43 Altamira to Itaituba rd; Event: eventDate: 08-21-71; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Brazil, Colombia, Ecuador, Peru

Valid species in .

Fairchild & Hertig, 1956

Fairchild & Hertig, 1956 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721991; sex: Female; Taxon: scientificName: ; Location: country: Costa Rica; stateProvince: Puntarenas; locality: Barranca, Finca Socorrito; Event: eventDate: 12-17-51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722048; sex: Male; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Cauca; municipality: Bolívar; locality: Bolívar (Cauca); Event: eventDate: 23/01/44; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Colombia, Costa Rica, Panama

Valid species in .

Summers, 1912

Summers, 1912 (

Type status: Paralectotype. Occurrence: catalogNumber: BMNHE1722082; sex: Male; Taxon: scientificName: ; Location: locality: Rio Javari, Peru/Brazil border; Event: year: 1912; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paralectotype. Occurrence: catalogNumber: BMNHE1722083; sex: Female; Taxon: scientificName: ; Location: locality: Rio Javari, Peru/Brazil border; Event: year: 1912; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paralectotype. Occurrence: catalogNumber: BMNHE1722124; sex: Female; Taxon: scientificName: ; Location: locality: Rio Javari, Peru/Brazil border; Event: year: 1912; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Lectotype. Occurrence: catalogNumber: BMNHE1722125; sex: Female; Taxon: scientificName: ; Location: locality: Rio Javari, Peru/Brazil border; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Brazil

Valid species in . Lectotype designated by Lewis 1967b.

Fairchild & Hertig, 1951

Fairchild & Hertig, 1951 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721982; sex: Female; Taxon: scientificName: ; Location: country: Panama; locality: Cerro Campana; Event: eventDate: 08-24-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1721983; sex: Male; Taxon: scientificName: ; Location: country: Panama; locality: Cerro Campana; Event: eventDate: 08-24-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722046; sex: Female; Taxon: scientificName: ; Location: country: Panama; locality: Cerro Campana; Event: eventDate: 08-24-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Colombia, Costa Rica, Panama, Peru

Valid species in .

Galvão & Coutinho, 1939

Galvão & Coutinho, 1939 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722069; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: São Paulo; municipality: Araçatuba; Event: year: 1935; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Argentina, Bolivia, Brazil, Ecuador, Paraguay, Peru

Valid species in .

Ortiz, 1965

Ortiz, 1965 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722072; sex: Male; Taxon: scientificName: ; Location: country: Venezuela; locality: Rancho Grande; Event: year: 1965; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Colombia, Peru, Venezuela

Valid species in .

Fraiha, Ward & Ready, 1981

Fraiha, Ward & Ready, 1981 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1251316; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Marabá; locality: Serra dos Carajás; Event: year: 1975; month: 2; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1251317; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Marabá; locality: Serra dos Carajás; Event: eventDate: 09-22-74; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722061; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Marabá; locality: Serra dos Carajás; Event: year: 1975; month: 2; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722062; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Marabá; locality: Serra dos Carajás; Event: eventDate: 09-23-74; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722065; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Marabá; locality: Serra dos Carajás; Event: eventDate: 09-23-74; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Brazil, Colombia, Peru

Valid species in .

Morales, Osorno-Mesa, Osorno & Hoyos, 1969

Morales, Osorno-Mesa, Osorno & Hoyos, 1969 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722094; sex: Female; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Boyacá; locality: Guayatá - Chitavita; Event: eventDate: 08-09-68; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722095; sex: Male; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Boyacá; locality: Guayatá - Chitavita; Event: eventDate: 08-09-68; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722131; sex: Female; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Boyacá; locality: Guayatá - Chitavita; Event: eventDate: 08-09-68; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722132; sex: Female; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Boyacá; locality: Guayatá - Chitavita; Event: eventDate: 08-09-68; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Colombia, Venezuela

Valid species in .

Young, 1979

Young, 1979 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722009; sex: Male; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Antioquia; municipality: Zaragoza; locality: 24km SW of Zaragoza at Rio Anori; Event: eventDate: 04-14-71; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Colombia, Ecuador, Venezuela

Valid species in .

Fairchild & Hertig, 1952

Fairchild & Hertig, 1952 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1251314; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: Almirante, Finca Nievecita; Event: eventDate: 06-21-50; eventRemarks: ; Record Level: institutionCode: NHMUK

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722060; Taxon: scientificName: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: Almirante; Event: eventDate: 22/07/50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Colombia, Costa Rica, Ecuador, Guatemala, Hounduras, Nicaragua, Panama

Valid species in .

Newstead, 1922

Newstead, 1922 (

Type status: Syntype. Occurrence: catalogNumber: BMNHE1721999; sex: Male; Taxon: scientificName: ; Location: country: Trinidad and Tabago; Event: eventDate: 01-07-22; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722000; sex: Female; Taxon: scientificName: ; Location: country: Trinidad and Tabago; Event: eventDate: 01-07-22; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722001; sex: Female; Taxon: scientificName: ; Location: country: Trinidad and Tabago; Event: eventDate: 01-07-22; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722002; sex: Male; Taxon: scientificName: ; Location: country: Trinidad and Tabago; Event: eventDate: 01-07-22; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722003; sex: Male; Taxon: scientificName: ; Location: country: Trinidad and Tabago; Event: eventDate: 01-07-22; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722004; sex: Female; Taxon: scientificName: ; Location: country: Trinidad and Tabago; Event: eventDate: 01-07-22; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722005; sex: Female; Taxon: scientificName: ; Location: country: Trinidad and Tabago; Event: eventDate: 01-07-22; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722006; sex: Female; Taxon: scientificName: ; Location: country: Trinidad and Tabago; Event: eventDate: 01-07-22; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, El Salvador, French Guiana, Guatemala, Honduras, Mexico, Nicaragua, Peru, Panama, Surinam, Trinidad and Tobago, Venezuela

Valid species in .

Fairchild & Hertig, 1952

Fairchild & Hertig, 1952 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722086; sex: Female; Taxon: scientificName: ; Location: country: Panama; stateProvince: Panamá; locality: Cerro Jefe, La Victoria; Event: eventDate: 08-29-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722087; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Colón; locality: Palenque; Event: eventDate: 09-15-49; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Belize, Colombia, Costa Rica, Ecuador, Guatemala, Mexico, Panama

Valid species in .

Mangabeira, 1942

Mangabeira, 1942 (

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722084; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Belém; locality: Aura; Event: eventDate: 10-08-40; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722085; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Belém; locality: Aura; Event: eventDate: 10-08-40; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722126; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Belém; locality: Aura; Event: eventDate: 10-08-40; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722127; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Belém; locality: Aura; Event: eventDate: 10-08-40; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722128; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Belém; locality: Aura; Event: eventDate: 10-08-40; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Syntype. Occurrence: catalogNumber: BMNHE1722140; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; municipality: Belém; locality: Aura; Event: eventDate: 10-08-40; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Brazil, Colombia, Ecuador, French Guiana, Peru, Surinam, Venezuela

Valid species in .

Ward & Fraiha, 1977

Ward & Fraiha, 1977 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722058; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; locality: Rio Jari, Monte Dourado; Event: eventDate: 06/02/76; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722059; sex: Female; Taxon: scientificName: ; Location: country: Brazil; stateProvince: Pará; locality: Rio Jari, Monte Dourado; Event: eventDate: 12/11/76; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Brazil, Colombia, French Guiana, Peru, Surinam, Venezuela

Valid species in .

Fairchild & Hertig, 1952

Fairchild & Hertig, 1952 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722027; sex: Male; Taxon: scientificName: ; acceptedNameUsage: ; Location: country: Panama; stateProvince: Panamá; locality: Serrania majé, Cerro Chucanti; Event: eventDate: 03-05-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Panama, Peru, Trinidad and Tobago, Venezuela

Synonym of (Floch & Abonnenc, 1950) Floch and Abonnenc 1950. Synonymised by Floch and Kramer 1965.

Fairchild & Hertig, 1958

Fairchild & Hertig, 1958 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722020; sex: Female; Taxon: scientificName: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: near camp No.1, Almirante; Event: eventDate: 05-17-51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722021; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: Almirante; Event: eventDate: 03-24-52; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722022; sex: Female; Taxon: scientificName: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: Almirante; Event: eventDate: 09-17-52; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722023; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: near camp No.1, Almirante; Event: eventDate: 05-17-51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722024; sex: Female; Taxon: scientificName: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: Almirante; Event: eventDate: 09-09-52; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722025; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: near camp No.1, Almirante; Event: eventDate: 05-17-51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Costa Rica, Panama

Valid species in .

Newstead, 1914

Newstead 1914 (

Sherlock, 1962

Type status: Lectotype. Occurrence: catalogNumber: BMNHE1722099; sex: Male; Taxon: scientificName: ; Location: locality: Rio Abuna, Brazil/Boliva border; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paralectotype. Occurrence: catalogNumber: BMNHE1722100; sex: Male; Taxon: scientificName: ; Location: locality: Rio Abuna, Brazil/Boliva border; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paralectotype. Occurrence: catalogNumber: BMNHE1722101; sex: Female; Taxon: scientificName: ; Location: locality: Rio Abuna, Brazil/Boliva border; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paralectotype. Occurrence: catalogNumber: BMNHE1722102; sex: Female; Taxon: scientificName: ; Location: locality: Rio Abuna, Brazil/Boliva border; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paralectotype. Occurrence: catalogNumber: BMNHE1722133; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: State of Rondonia; locality: Rio Abuna, Brazil/Boliva border; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paralectotype. Occurrence: catalogNumber: BMNHE1722135; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: State of Rondonia; locality: Rio Abuna, Brazil/Boliva border; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paralectotype. Occurrence: catalogNumber: BMNHE1722136; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: State of Rondonia; locality: Rio Abuna, Brazil/Boliva border; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paralectotype. Occurrence: catalogNumber: BMNHE1722137; sex: Male; Taxon: scientificName: ; Location: country: Brazil; stateProvince: State of Rondonia; locality: Rio Abuna, Brazil/Boliva border; Event: eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Brazil, Colombia, Ecuador, French Guiana, Panama, Peru, Trinidad and Tobago, Venezuela

Valid species in . Lectotype designated by Lewis 1967b.

Young, Brener & Wargo, 1983

Young, Brener & Wargo, 1983 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722105; sex: Female; Taxon: scientificName: ; Location: country: USA; stateProvince: California; municipality: Riverside; locality: Cahuilla Hills, Palm desert; Event: eventDate: 09-16-81; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

United States of America

Valid species in .

Fairchild & Hertig, 1952

Fairchild & Hertig, 1952 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1251315; sex: Male; Taxon: scientificName: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: Almirante, Finca Nievecita; Event: eventDate: 06-22-50; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1251318; sex: Female; Taxon: scientificName: ; Location: country: Panama; stateProvince: Bocas del Toro; locality: Almirante; Event: eventDate: 06-29-51; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Belize, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama

Valid species in .

Young & Porter, 1972

Young & Porter, 1972 (

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722056; sex: Male; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Antioquia; municipality: Zaragoza; locality: 24km S & 21 km W of Zaragoza; Event: eventDate: 23/05/70; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Type status: Paratype. Occurrence: catalogNumber: BMNHE1722057; sex: Female; Taxon: scientificName: ; Location: country: Colombia; stateProvince: Antioquia; municipality: Zaragoza; locality: 24km S & 21 km W of Zaragoza; Event: eventDate: 12/09/70; eventRemarks: ; Record Level: institutionCode: NHMUK; basisOfRecord: Preserved Specimen

Bolivia, Brazil, Colombia, Ecuador, Peru, Venezuela

Valid species in .