Checklist of British and Irish Hymenoptera - Cynipoidea.

BACKGROUND: The British and Irish checklist of Cynipoidea is revised, considerably updating the last complete checklist published in 1978. Disregarding uncertain identifications, 220 species are now known from Britain and Ireland, comprising 91 Cynipidae (including two established non-natives), 127 Figitidae and two Ibaliidae. NEW INFORMATION: One replacement name is proposed, Kleidotoma thomsoni Forshage, for the secondary homonym Kleidotoma tetratoma Thomson, 1861 (nec K. tetratoma (Hartig, 1841)).

Introduction

This paper continues the series of updated British and Irish checklists that started with Broad and Livermore (2014a), Broad and Livermore (2014b), Liston et al. (2014) and with an introduction by Broad (2014). This represents the first complete update of the British list since 1978 (Fitton et al. 1978).

The is a rather diverse superfamily of three British families (the and are entirely extralimital). The and comprise koinobiont endoparasitoids of other insects whereas the are gall-formers, or inquilines of gall-formers. A summary of cynipoid biology can be found in Gauld and Bolton (1988). Briefly, many are parasitoids of larvae although the are hyperparasitoids of primary parasitoids in aphids and some other and the are parasitoids of (); the small family are specialised parasitoids of larvae; and the are entirely phytophagous, most conspicuously as gall-formers of oaks () but with numerous species galling other (and a few other plants) and the tribe are inquilines of other cynipid galls. A few representative are illustrated in Figs 1, 2, 3.

The British checklist was updated comparatively recently (Spooner and Bowdrey 2000) and Irish checklists were published by O'Connor and Nash (1998), O'Connor et al. (2003), O'Connor (2004), O'Connor et al. (2009) and O'Connor et al. (2009). Our knowledge of the figitid fauna is still far from perfect but recent work by M. Forshage on the European and a catalogue of (Ferrer-Suay et al. 2012b plus updated synonymy in several other papers) have been timely. MF has sorted and identified the BMNH collections of British , adding many new species and distribution records. In contrast to the rather neglected and , gall wasps () have received a certain amount of attention over the years and are studied and recorded under the auspices of the British Plant Gall Society. The numbers of valid, certainly identified are listed by family and country in Table 1. The total fauna has increased by 9% since the last checklist (Quinlan 1978b). This rather small increase can be explained by the substantial increase in taxonomic work in recent years, resulting in synonymy and corrected identifications nearly keeping pace with new discoveries.

Table 1.

Numbers of confirmed British and Irish broken down by family and country, with numbers from the 1978 checklist (Quinlan 1978b) for comparison. Totals do not include uncertain identifications.

family	total valid species 1978	total valid species 2017	England	Scotland	Wales	Ireland	Isle of Man
Cynipidae	78 [inc. 1 introduced]	91 [inc. 2 introduced]	87	49	56	48	13
Figitidae	123	127	111	44	30	50	1
Ibaliidae	1	2	2	1	0	0	0
	202	220	200	94	86	98	14

Materials and methods

We reference all additions to and deletions from the 1978 British list (Quinlan 1978a) and record country-level distribution within the British Isles, i.e. England, Scotland, Wales, Ireland (as one unit) and the Isle of Man. A more complete introduction to the methods and rationale behind this checklist series can be found in Broad (2014).

Because the agamic and sexual generations of cynipids are sometimes referred to by different names, these are differentiated in the checklist. The following conventions and abbreviations are used:

[] taxon deleted from the British and Irish list and nomina dubia

BMNH Natural History Museum, London

# known introductions occurring only under artificial conditions or established in the wild

? status (including uncertain synonymy) or identification in the British Isles uncertain

misident. has been misidentified as this name

nom. dub. nomen dubium, a name of doubtful status

nom. ob. nomen oblitum, ‘forgotten name’, does not have priority over a younger name

nom. nov. nomen novum, a replacement name

nom. nud. nomen nudum, an unavailable name, with no type specimen

preocc. name preoccupied (junior homonym)

stat. rev. status revocatus, revived status (e.g. raised from synonymy)

unavailable name unavailable under provisions of the zoological code

var. variety, only available as a valid name under certain provisions of the zoological code

f. form, only available as a valid name under certain provisions of the zoological code

-a- name based on agamic (asexual) generation (used in )

-s- name based on sexual generation (used in )

Distribution data for are mainly derived by JPB from published sources, but thanks are due to the following for supplying additional data: Janet Boyd, Records Data Manager, British Plant Gall Society; Adrian Fowles, Countryside Council for Wales; Kate Hawkins, Manx Natural Heritage; David Notton, BMNH; Mark Pavett, National Museum of Wales (all pers. comm.). Distribution data for and are mainly from BMNH and cited published sources.

Word document and Excel spreadsheet versions of the checklist are available as supplementary files: Suppl. materials 1, 2.

Checklists

Latreille, 1802

Notes

Liljeblad and Ronquist (1998), Ronquist (1999) and Ronquist et al. (2015) employed a series of monophyletic tribes, in the absence of strong evidence for relationships above this level. Synonymy for includes all names that have appeared in the British literature but does not necessarily include all Palaearctic names proposed as varieties or forms. For complete synonymy please see Melika (2006a) and Melika (2006b), also Nieves-Aldrey (2001). It should be borne in mind that future molecular studies may change our understanding of some species concepts and their alternating generations.

Nieves-Aldrey, 1994

The availability of the name has been questioned and is currently being looked into. This will possibly only be solved by the ICZN.

Ashmead, 1897

Ashmead,1903

Species of removed from the British and Irish list:

[ (Kieffer, 1900, )] Added by Bagnall (1931) from the gall. Discussed by Bowdrey (1999), who concluded that this record was in error.

Barbotin, 1972

Distribution

England

added by Bowdrey (1994).

(Linnaeus, 1758)

Linnaeus, 1758

(Hartig, 1840,

(Cameron, 1875,

(Thomson, 1877,

(Thomson, 1877,

(Thomson, 1877,

England, Scotland, Wales

Quinlan & Askew, 1969

Scotland

(Kieffer, 1901)

Kieffer, 1901

England

Niblett, 1946

England

(Thomson, 1877)

Thomson, 1877

(Kieffer, 1898,

England

, added by Bagnall and Harrison (1930) on the basis of galls only (root collar of ), was synonymised by Eady and Quinlan (1963) with but erroneously so with in Fauna Europaea (Noyes et al. 2004) (Nieves-Aldrey, pers. comm.).

Förster, 1869

Förster, 1869

(Kieffer, 1898)

Kieffer, 1898

England

(Schenck, 1863)

Schenck, 1863

(Schenck, 1863,

England, Scotland

(Giraud, 1859)

Giraud, 1859

(Giraud, 1859,

(Thomson, 1877,

Wachtl, 1880

England

: galls bracts of , but not now considered to be a distinct species (Nieves-Aldrey 1994).

Förster, 1869

misspelling

(Linnaeus, 1758)

Linnaeus, 1758

(Kieffer, 1898,

England, Scotland, Wales, Isle of Man

Ashmead, 1903

Hartig, 1840

Hartig, 1843

Hartig, 1840

England, Ireland

(Perris, 1840)

Perris, 1840

(Bouché, 1834,

England, Scotland, Wales

Nieves-Aldrey, Nylander & Ronquist, 2015

Hartig, 1840

Mayr, 1873

Giraud, 1911

Giraud, 1911

England

Added by Jennings Jennings (2016).

Hartig, 1840

Hartig, 1840

Hartig, 1841

Hartig, 1841

England, Scotland, Wales, Ireland

Latreille, 1802[4]

Generic classification of oak-associated follows Melika and Abrahamson (2002). As sexual and agamic generations have often been described under different names, those based on agamic generation are denoted by the suffix '-a-', those based on the sexual generation by '-s-'.

Hartig, 1840

Förster, 1869

Förster, 1869

Radoszkowski, 1866

Ashmead, 1897

Kieffer, 1906

Rohwer & Fagan, 1917

Dettmer, 1925

Dettmer, 1925

Kinsey,1937

Kinsey, 1937

Kinsey, 1937

Kinsey, 1938

Species of removed from the British and Irish list:

[ (Giraud, 1859, ) -a-] Added by Cameron (1893) from the gall only. An agamic female in the Morley collection at Ipswich Museum is . No recent records.

[ (Olivier, 1791, ) -a-, syn. (Hartig, 1843, ) -a-] Niblett et al. (1932) (following Houard) state that this species was accidentally introduced in the 19th Century on , and never established.

[ (Boyer de Fonscolombe, 1832, ) -a-, syn. sufflator Mayr, 1882 -s-] First recorded from old galls by Bagnall and Harrison (1918). Sexual generation also included by Eady and Quinlan (1963), doubtfully British, no confirmed records.

[ (Bosc, 1792, ) -a-] Added by Eady and Quinlan (1967) on the basis of a single gall found on a road at Kew. No further records.

[ Kieffer, 1898 -a-] Added by Bagnall and Harrison (1918) from the gall only. No further records and rejected as British by Eady and Quinlan (1963).

Giraud, 1859 -s-

(Giraud, 1859,

Dalla Torre & Kieffer, 1910 -a-

England, Scotland

Sexual generation catkin galls first recorded by Traill (1873). The asexual generation, described as Dalla Torre & Kieffer, 1910, has not been recorded in Britain.

(Giraud, 1859) -a-

Giraud, 1859

England, Wales

Added by Leach (1999). First recorded from the agamic generation, the sexual generation was reported by Walker (2001b).

(Hartig, 1841) -a-

Hartig, 1841

Adler, 1881 -s-

Wachtl, 1882 -a-

England, Scotland, Wales, Ireland

(Schlechtendal, 1870) -a-

Schlechtendal, 1870

(Trotter, 1899,

Melika, Csóka & Pujade-Villar, 2000 nom. nud. -s-

England, Scotland, Ireland

was placed by Bellido et al. (2005) in synonymy with but previously regarded as a distinct species (Melika et al. 2000). Agamic generation reported by Harrison (1930) from Co. Durham and Forfarshire. The sexual generation was described as forma elianae. Form larshemi Leeuwen and Dekhuijzen-Maasland (1958), was described as the sexual generation of but is now believed to be Bellido & Pujade-Villar, 2004, fide Folliot et al. (2004), who also produced the true sexual generation of experimentally.

Wachtl, 1880 -s-

England

Added by Bowdrey (2015); an adult female was identified as this species by G. Melika with the proviso that it might be the sexual generation of a morphologically undescribed species known from DNA sequencing.

Hartig, 1840 -s-

(Hartig, 1840,

(Hartig, 1840,

(Dahlbom, 1842,

(Schenck, 1863,

Schenck, 1863 -a-

Schenck, 1863 -s-

(Förster, 1869,

(Schlechtendal, 1870,

(Schlechtendal, 1870,

England, Scotland, Wales, Ireland, Isle of Man

(Hartig, 1840) -a-

Hartig, 1840

misspelling

(Linnaeus, 1758,

Lacaze-Duthiers, 1853 -a-

(Schenck, 1863,

Adler, 1881 -s-

England, Scotland, Wales, Ireland, Isle of Man

(Giraud, 1859) -a-

Giraud, 1859

(Wachtl, 1876,

Dalla Torre & Kieffer, 1910 -a-

England

Added by Bowdrey (2009). Previously reported by Ormerod (see Cameron 1893), but reportedly as a leaf gall, so the record is suspect.

Kieffer, 1901

England

Gall recorded by Bagnall and Harrison (1919) and adult male possibly of this species by Eady and Quinlan (1963). Only the sexual generation is known, inducing bud galls of , collected in France (Dalla Torre and Kieffer 1910). Bagnall and Harrison (1918) mentioned this species from Great Britain but this record requires confirmation (Eady and Quinlan 1963). A species of uncertain status, type probably lost.

(Hartig, 1840) -a-

Hartig, 1840

Schlechtendal, 1884 -s-

England, Scotland, Ireland

Giraud, 1859 -s-

(Wachtl, 1879,

Kieffer, 1896 -a-

Belizin & Maisuradze, 1961 -s-

England

Added by Walker (2001a). Rearing experiments by Walker (2002) confirmed to represent the sexual generation of a lifecycle also involving the asexual generation galls known as , a pairing also indicated by DNA sequence data (Stone et al. 2008).

Hartig, 1840 -s-

(Hartig, 1841,

Belizin & Maisuradze, 1962 -?-

England, Scotland, Wales, Ireland

(Hartig, 1843) -a-

Hartig, 1843

auctt. misident.

(Christ, 1791,

Mayr, 1870 -s-

England, Scotland, Wales, Ireland, Isle of Man

Nineteenth Century introduction (Adler and Stratton 1894; Niblett et al. 1932).

Wiebes-Rijks, 1980 -a-

England

(Hartig, 1840) -a-

Hartig, 1840

van Leeuwen & Dekhuizen-Maasland, 1958 -s-

England, Scotland, Wales, Ireland, Isle of Man

(Hartig, 1843) -a-

Hartig, 1843

Giraud, 1859 -s-

(Kieffer, 1900,

England

Added by Mosley (1892) from Yorkshire and by (Cameron 1893) from Loch Lomond (sexual form). Not recorded subsequently until reinstated as a British species, from London specimens, by Stone and Sunnocks (1992).

(Adler, 1881) -a-

Adler, 1881

Adler, 1881 -s-

England, Scotland, Wales, Ireland

Nomenclature follows Pujade-Villar and Melika (2000).

(Radoszkowski, 1866) -a-

Radoszkowski, 1866

(Kaltenbach, 1867,

(Girauld, 1868,

(Schlechtendal, 1870,

Kieffer, 1897 -a-

Folliot, 1964 -s-

England, Scotland, Wales, Ireland

Taxonomy follows Melika et al. (2000).

Hartig, 1840 -a-

Schenck, 1863 -a-

Schenck, 1863 -a-

Schenck, 1863 -a-

Schenck, 1863 -a-

(Schlechtendal, 1870,

(Thomson, 1877,

Pigeot, 1900 -s-

f.

England, Scotland, Wales, Ireland

Synonymy of and established by Folliot (1964).

(Burgsdorff, 1783) -a-

Burgsdorff, 1783

Beyerinck, 1896 -s-

Trotter, 1899 -s-

England, Scotland, Wales, Ireland

(Linnaeus, 1761) -a-

Linnaeus, 1761

(Hartig, 1840,

(Hartig, 1841,

Adler, 1881 -s-

Tavares, 1901 -a-

England, Scotland, Wales, Ireland, Isle of Man

(Fabricius, 1798) -a-

Fabricius, 1798

Hartig, 1840 -s-

Hartig, 1840 -s-

(Hartig, 1841,

(Hartig, 1841,

Thomson, 1877 -s-

England, Scotland, Wales, Ireland

(Linnaeus, 1761) -s-

Linnaeus, 1761

(Hartig, 1840,

(Hartig, 1843,

Schenck, 1863 unjustified emendation -s-

England, Scotland, Wales, Ireland

(Hartig, 1843) -a-

Hartig, 1843

Kierych, 1965 -a-

England, Wales

Gall recorded by Bagnall and Harrison (1918) and Burkill (1933). Modern UK status uncertain. Galls of the sexual generation unconfirmed in Britain. See also under .

(Giraud, 1859) -a-

Giraud, 1859

(Schlechtendal, 1870,

England, Scotland, Wales

No sexual generation known.

(Hartig, 1843) -a-

Hartig, 1843

(Hartig, 1840,

(Bechstein & Scharfenberg, 1805,

Folliot, 1964 -a-

Folliot, 1964 -s-

England, Wales

Folliot, 1964: sexual generation, produced experimentally.

Mayr, 1870 -s-

Mayr, 1881 -s-

England

added by Jennings (2014)

(Boyer de Fonscolombe, 1832) -a-

Boyer de Fonscolombe, 1832

(Hartig, 1840,

Tschek, 1871 -s-

(Olivier, 1791,

(Dettmer, 1925,

Tavares, 1930 -a-

England, Scotland, Wales, Ireland

Hartig, 1840 -s-

Kieffer, 1900 -s-

England?

Maintained with a query; possibly the unconfirmed sexual generation of (q.v.). If specifically distinct, British records need confirmation. See Melika (2006a).

Mayr, 1881

(Giraud, 1859) -a-

Giraud, 1859

England

added by Crawley (1997)

Westwood, 1840

Hartig, 1840

Hartig, 1840 preocc.

Walsh, 1864

Guérin-Méneville, 1865

Förster, 1869

Rondani, 1871 preocc.

(Olivier, 1791) -s-

Olivier, 1791

(Bosc, 1791,

(D'Anthoine, 1794,

(D'Anthoine, 1794,

(Fabricius, 1798,

(Hartig, 1840,

(Stadelman, 1892,

England, Scotland, Wales, Ireland, Isle of Man

Förster, 1869

Dettmer, 1930

Species of removed from the British and Irish list:

[ (Giraud, 1859, ) -a-] Recorded by Rolfe (1881) and Askew (1959) but in error fide Eady and Quinlan (1963).

(Dettmer, 1930) -s-

Dettmer, 1930

England, Wales, Ireland

(Giraud, 1859) -a-

Giraud, 1859

Dettmer, 1933 -a-

Dettmer, 1933 -a-

Förster, 1869 -s- synonymy by

England, Wales

Förster, 1869 established as the sexual generation by Pujade-Villar et al. (2007), who produce evidence that may be a separate species.

Linnaeus, 1758

misapplied

Förster, 1869

Species of excluded from the British and Irish list:

[quercus (Fourcroy, 1785, ) -a-; syn. pubescentis (Mayr, 1881, ) -s-] Both generations reported by Bagnall and Harrison (1918), Bagnall and Harrison (1919) from several localities in Northumberland and Tyne & Wear, perhaps in error due to confusion with the similar quercusfolii. There are no subsequent records and the species requires confirmation as British.

Hartig, 1840 -a-

Folliot, 1964 -s-

England, Scotland, Wales, Ireland

Hartig, 1840 -a-

Niblett, 1948 -s-

England, Scotland, Wales, Ireland

Hartig, 1840 -a-

(Schlechtendal, 1870,

England, Scotland, Wales, Ireland

Hartig, 1840 -a-

(Adler, 1881,

Kinsey, 1930 -s-

England, Scotland, Wales, Ireland

Linnaeus, 1758 -a-

Gleditsch, 1774 -a-

(Olivier, 1791,

D'Anthoine, 1794 -a-

(D'Anthoine, 1794,

(Giraud, 1868,

(Tschek, 1869,

(Schlechtendal, 1870,

England, Scotland, Wales, Ireland

Galls of the sexual generation were recorded as by Bagnall and Harrison (1918); there appear to be no further records.

Giraud, 1859

Species of removed from the British and Irish list:

[ Giraud, 1859 -a-] Galls recorded once by Fitch (1874); no further UK records.

Yasumatsu, 1951

England

Added by Malumphy (2015), EPPO (2015). This highly invasive, East Asian species has spread across Europe and has recently been reported from Kent and Hertfordshire. As the UK has Protected Zone Status for this global pest, apparently successful attempts were made to eradicate at the two outbreak sites and it should not be formally added to the British list at this stage.

Hartig, 1840

Hartig, 1840

Förster, 1869

Ashmead, 1887

Kinsey, 1923

Kinsey, 1923

Monzen, 1954

Belizin & Maisuradze, 1961

Species of removed from the British and Irish list:

[ (Bignell, 1892, )] Described by Bignell (1892), no subsequent records.

(Schenck, 1863) -s-

Schenck, 1863

Schenck, 1863 -a-

Schlechtendal, 1870

Tavares, 1928 -s-

England, Scotland, Wales, Ireland, Isle of Man

Form ‘’ described by Entwistle and Hails (1997).

(Curtis, 1838) -a-

Curtis, 1838

(Boyer de Fonscolombe, 1832,

(Hartig, 1840,

Beyerinck, 1882 -s-

England, Scotland, Wales, Ireland

Transferred from by Pujade-Villar et al. (1998).

(Geoffroy, 1785) -a-

Geoffroy, 1785

(Olivier, 1790,

Hartig, 1840 -a-

Hartig -a-

(Curtis, 1843,

Schenck, 1863 -a-

(Schlechtendal, 1870,

Hieronymus, 1890 -s-

Dettmer, 1925 -s-

England, Scotland, Wales, Ireland

Hartig, 1840 -s-

(Hartig, 1840,

Hartig, 1841 -s-

Hartig, 1841 -a-

Hartig, 1841 -a-

(Giraud, 1859,

Mayr, 1870 -a-

(Schlechtendal, 1870,

Mayr, 1870 -a-

England, Scotland, Ireland

(Linnaeus, 1758) -s-

Linnaeus, 1758

(Linnaeus, 1758,

(Fourcroy, 1785,

(Fourcroy, 1785,

(Olivier, 1791,

(Fabricius, 1793,

(D'Anthoine, 1794,

(D'Anthoine, 1794,

Hartig, 1840 -a-

(Hartig, 1840,

(Hartig, 1841,

(Blanchard, 1849,

(Duméril, 1860,

Schenck, 1863 -a-

Schenck, 1863 -a-

(Küstenmacher, 1894,

England, Scotland, Wales, Ireland, Isle of Man

(Hartig, 1841) -s-

Hartig, 1841

Hartig, 1841 -a-

(Schenck, 1863,

England, Scotland, Wales, Ireland

Mayr, 1881

Kieffer, 1903

Kiefer, 1903

(Mayr, 1882) -a-

Mayr, 1882

England, Wales

Added by Robbins (2007) based on old agamic gall only; subsequently found at Imperial Wharf and Royal Botanic Gardens, Kew, both London (D.G. Notton, pers. comm.). Galls introduced and .

(Mayr, 1882) -a-

Mayr, 1882

(Kieffer, 1902,

Wales

Added by Robbins (2007), based on old agamic gall only.

(Fabricius, 1798) -s-

Fabricius, 1798

(Lichtenstein, 1877,

(Lichtenstein, 1877,

Mayr, 1882 -s-

Mayr, 1882 -s-

England

Added by Hancy and Hancy (2004). Added from sexual generation gall, adults reared. Recent records from Devon, Cornwall and Hampshire (Isle of Wight). Catkin galls reported by Biggs (2011) from Isle of Wight as forma fusifex, which has no taxonomic status.

Kinsey, 1923

(Kollar, 1857) -a-

Kollar, 1857

(Giraud, 1859,

(Giraud, 1859,

England

added by Redfern (2006)

Hartig, 1840

Ashmead, 1897

Kovatev, 1965

Kovalev, 1965

Species of removed from the British and Irish list:

[synaspis (Hartig, 1841, ) -a-; syn. megapteropsis Wriese, 1900 -s-] Recorded by Bagnall and Harrison (1918) from the agamic gall; no further records.

(Panzer, 1801) -s-

Hartig, 1840 -s-

(Hartig, 1840,

England, Scotland, Wales, Ireland

Nieves-Aldrey, Nylander & Ronquist, 2015

Hartig, 1840

Ashmead, 1897

Species of removed from the British and Irish list:

[mayri Reinhard, 1876] The status of mayri as a British species appears to rest solely on three published references to the species: Cameron (1893), subsequently quoted in Connold (1909) and repeated by Swanton (1912). Nowhere does Cameron claim to have evidence of this species’ occurrence in Britain, merely comparing its gall to other British species of and . As there appear to be no genuine UK records it should therefore be excluded.

(Bouché, 1834)

Bouché, 1834

Cameron, 1889

(Marshall, 1867,

England, Wales, Ireland

Förster, 1869

(Ratzeburg, 1831)

Ratzeburg, 1831

England, Scotland, Wales

(Hartig, 1840)

Hartig, 1840

(Schenck, 1863,

Dettmer, 1925

England, Scotland, Ireland

Dettmer, 1924

England, Scotland, Wales, Ireland

Close to , taxonomic status needs revision (note in Fauna Europaea).

Förster, 1869

(Thomson, 1877)

Thomson, 1877

Schlechtendal, 1880

England, Scotland, Wales, Ireland

(Retzius, 1783)

Retzius, 1783

(Curtis, 1838,

(Hartig, 1840,

Schenck, 1863

(Thomson, 1877,

(Thomson, 1877,

England, Wales, Ireland, Isle of Man

Nieves-Aldrey, Nylander & Ronquist, 2015

Förster, 1860

Ashmead, 1897

(Hedicke, 1939)

Hedicke, 1939

England, Wales

(Förster, 1860)

Förster, 1860

(Thomson, 1877,

Diakontschuk, 1980

Diakontschuk, 1980

England, Scotland

(Kieffer, 1887)

Kieffer, 1887

Stefani, 1903

England, Wales, Ireland, Isle of Man

(Ashmead, 1897)

Ashmead, 1897

England, Scotland

Galls recorded from Co. Durham (Bagnall 1917), Derbyshire (Fordham 1917), Northumberland (Bagnall 1917) and Scotland (Bagnall 1932). No recent records.

Mayr, 1881

Synonymised with (Eady and Quinlan 1963); re-established by Nieves-Aldrey (1994) but Melika (2006a) again treated as a synonym of .

Species of removed from the British and Irish list:

[ (De Stefani, 1900, )] Galls recorded from Norfolk and Surrey by Bagnall and Burkill (1935) but no recent records. Possibly confused in the literature with which also galls (Nieves-Aldrey 2001).

(Perris, 1873)

England

Galls recorded in Norfolk and Derbyshire by Bagnall and Harrison (1934).

(Tavares, 1904)

Tavares, 1904

England

added by Jennings (2005)

Latreille, 1802

Hartig, 1840

Geoffroy, 1762

Hartig, 1840

Förster, 1869

Walsh, 1864

Kieffer, 1902

Sakugami, 1949

(Hartig, 1840)

Hartig, 1840

(Förster, 1869,

England

Galls not distinguishable from those of the smooth form of nervosa; records from Scotland, Wales and the Isle of Man are based on galls only.

(Schlechtendal, 1877)

Schlechtendal, 1877

(Beijerinck, 1883,

England

(Curtis, 1838)

Curtis, 1838

(Hartig, 1840,

(Giraud, 1859,

(Kieffer, 1904,

(Loiselle, 1912,

(Niblett, 1943,

England, Wales

(Linnaeus, 1758)

Linnaeus, 1758

Fourcroy, 1785

England, Scotland, Wales, Ireland, Isle of Man

(Giraud, 1859)

Giraud, 1859

(Inchbald, 1861,

England, Scotland, Wales, Ireland, Isle of Man

Ashmead, 1896

Taxonomy follows Pujade-Villar et al. (2003).

Dalla Torre & Kieffer, 1910

(Hartig, 1840)

Hartig, 1840

(Hartig, 1840,

England, Scotland, Ireland

Hartig, 1840

Förster, 1869

Species of removed from the British and Irish list:

[hayneanus (Ratzeberg, 1833, )] Added by Morley (1931); specimen in his collection at Ipswich Museum redetermined as (Eady 1952).

Hartig, 1841

England, Scotland

Eady, 1952

England, Wales, Ireland

Giraud, 1911

England

Added by Jennings (2017); identified by G. Melika.

(Curtis, 1838)

Curtis, 1838

Mayr, 1872

Tavares, 1920

Dettmer, 1924

England, Wales, Ireland

Hartig, 1840

misident.

England, Scotland, Wales, Ireland

This species has hitherto been known as in Britain but it was shown in Pujade-Villar (2005) that this name refers to another species (of ) and the widespread species should be called . Melika (2006a) proposed that (Boyer de Fonscolombe, 1832, ) should be retained as the valid name but did not make an application to ICZN to overturn the type designation of .

Hartig, 1840

Hartig, 1841

Hartig, 1843 preocc.

England, Scotland, Wales, Ireland

Hartig, 1841

Dalla Torre, 1893

England, Wales, Ireland

Mayr, 1872

England, Wales

Hartig, 1840

Hartig, 1840

Hartig, 1840

Hartig, 1840

Hartig, 1841

Hartig, 1841

Hartig, 1841

Hartig, 1841

Hartig, 1841

Mayr, 1872

Mayr, 1873

Giraud, 1911

Dettmer, 1924

Dettmer, 1924

England, Wales, Ireland

Mayr, 1872

England

Mayr, 1872

England, Scotland, Wales, Ireland

Hartig, 1840

England, Ireland

(Dalman, 1823)

Dalman, 1823

Hartig, 1840

Hartig, 1841

(Hartig, 1841,

Dalla Torre 1893 unjustified emendation

Giraud, 1911

Giraud, 1911

Dettmer, 1924 preocc.

England

Hartig, 1840

Hartig, 1841

Hartig, 1841

Mayr, 1872

(Olivier, 1791)

Olivier, 1791

(D'Anthoine, 1794,

(Boyer de Fonscolombe, 1832,

Hartig, 1841

Hartig, 1843

Kollar, 1843

Dettmer, 1924 preocc.

England, Scotland, Wales, Ireland

Mayr, 1872

Giraud, 1911

Giraud, 1911

Vassileva-Samnalieva, 1986

England

Added by Chinery & Williams in Melika (2006a).

Hartig, 1840

Some Irish records from O'Connor et al. (2003).

Thomson, 1862

Förster, 1869

Kovalev, 1979 synonymy by

Kovalev, 1979 synonymy by

Synonymy follows Fergusson (1986) except where otherwise noted.

Haliday in Walker, 1835

Fergusson, 1985

England, Ireland

added by Fergusson (1985)

(Dalman, 1823)

Dalman, 1823

(Westwood, 1833,

(Westwood, 1833,

Reinhard, 1860

Reinhard, 1860

Cameron, 1883

Cameron, 1883

Cameron, 1887

England, Ireland, Isle of Man

Fergusson, 1985

England

added by Fergusson (1985)

Fergusson, 1985

England, Scotland

added by Fergusson (1985)

Dalman, 1823

Hartig, 1840

Förster, 1869 preocc.

Dalle Torre & Kieffer, 1910

(Dalman, 1818)

Walker, 1835

Walker, 1835

(Zetterstedt, 1838,

(Hartig, 1840,

misspelling

England, Ireland

Walker, 1835

Walker, 1835

(Hartig, 1841,

Dahlbom, 1842

(Förster, 1869,

England, Ireland

Hartig, 1843

Hedicke, 1914

Western Palaearctic species were revised by Mata-Casanova et al. (2015).

(Giraud, 1860)

Giraud, 1860

(Giraud, 1860,

(Cameron, 1883,

England, Scotland, Ireland

Kieffer, 1901

misident.

England, Scotland, Ireland

Dalla Torre & Kieffer, 1910

Thomson, 1862

Dahlbom, 1842

Haliday in Curtis, 1829 preocc.

Giraud, 1860

(Villers, 1789)

Villers, 1789

(Rossi, 1790,

(Boyer de Fonscolombe, 1832,

(Boyer de Fonscolombe, 1832,

Kieffer, 1907 (

Ireland

After the taxonomic changes in the revision by Ros-Farré and Pujade-Villar (2013) it is not entirely certain which nominal species is present in the British material. The authors cite as British but with a reference to Fergusson (1986), thus not based on reexamination of specimens.

Dahlbom, 1842

Nomenclature follows Ros-Farré and Pujade-Villar (2009).

Dahlbom, 1842

Dahlbom, 1842

(Cameron, 1879,

Giraud, 1860

Dahlbom, 1856 unjustified emendation

Ashmead, 1887

(Cameron, 1888,

(Kieffer, 1901,

Dettmer, 1924

(Kieffer, 1901,

(Kieffer, 1901,

England, Ireland

Haliday in Walker, 1835

Hartig, 1840 preocc.

Hartig, 1843

Förster, 1869

Transferred from to by Buffington et al. (2007).

Synonymy follows Fergusson (1986).

Giraud, 1860

misident.

(Dahlbom, 1842,

(Reinhard, 1860,

England, Ireland

(Kieffer, 1903)

Kieffer, 1903

England, Ireland

(Hartig, 1840)

Hartig, 1840

Cameron, 1883

England, Ireland

Giraud, 1860

Förster, 1869

(Cameron, 1879,

England, Scotland, Wales, Ireland

Giraud, 1860

Kieffer, 1901

Kieffer, 1901

(Kieffer, 1901)

Kieffer, 1901

Wales

Dalla Torre & Kieffer, 1910

Thomson, 1862 unavailable

Hellén, 1931

Kierych, 1979

Kovalev, 1994

Except where noted, nomenclature follows Menke and Evenhuis (1991) and Ferrer-Suay et al. (2012d). Further taxonomic changes have been made in a series of papers by Ferrer-Suay and colleagues (Ferrer-Suay et al. 2012a, Ferrer-Suay et al. 2012b, Ferrer-Suay et al. 2012c, Ferrer-Suay et al. 2013a, Ferrer-Suay et al. 2013b, Ferrer-Suay et al. 2013c, Ferrer-Suay et al. 2014a, Ferrer-Suay et al. 2015, Ferrer-Suay et al. 2013d, Ferrer-Suay et al. 2014b). Tribal subdivisions have been abandoned following Paretas-Martínez et al. (2007), who found that was paraphyletic with respect to . Some Welsh occurrence records from Baker (2013).

Förster, 1869

Westwood, 1833 preocc.

Hartig, 1840 preocc.

Förster, 1869

Förster, 1869

Kierych, 1988

Kovalev, 1994

Quinlan (1978a) listed 36 species for Britain and Ireland, which Fergusson (1986) reduced to 11, mainly through synonymy. However, Fergusson did not work with much reared material and Evenhuis (1985) and Müller et al. (1999) have adopted much narrower species limits than Fergusson, an approach borne out by FVV’s work on the biology and taxonomy of the genus (see Van Veen et al. 2003).

species of excluded from the British and Irish list:

[ (Hartig, 1841, ) incertae sedis]

[ (Giraud, 1860, ) incertae sedis; misspelling]

These species were listed by Quinlan (1978a) but not recognised as British or Irish by Fergusson (1986) and not found by FVV.

species of uncertain status:

[ (Kieffer, 1900, ) nom. dub.] Listed by Fergusson (1986) as a synonym of , the type material of has not been located and it has not been possible to interpret the name (Ferrer-Suay et al. 2012b).

Fergusson, 1986

England

added by Fergusson (1986)

Hellén, 1963

England

Det. Van Veen, added here. The species identified as by Müller et al. (1999) is actually , although the true has also been found in Silwood Park.

(Kieffer, 1902)

Kieffer, 1902

misident.

Evenhuis, 1976

Scotland, Wales

Raised from synonymy with by Ferrer-Suay et al. (2012a).

(Cameron, 1886)

Cameron, 1886

(Cameron, 1886,

(Cameron, 1889,

Scotland

Raised from synonymy with by Ferrer-Suay et al. (2013).

(Hartig, 1840)

Hartig, 1840

England, Scotland, Wales, Ireland

(Thomson, 1862)

Thomson, 1862

misident.

England

(Hartig, 1841)

Hartig, 1841

misident.

(Cameron, 1886,

(Cameron, 1883,

(Cameron, 1883,

(Cameron, 1889,

Kieffer, 1902 (

(Kieffer, 1902,

Kieffer, 1904

(Hedicke, 1928,

(Hartig, 1841,

(Thomson, 1862,

England, Wales

Synonymised with by Fergusson (1986) but treated as a valid species by Ferrer-Suay et al. (2012b) following the advice of FVV. is listed as a synonym of in Fauna Europaea (Noyes et al. 2004), following alternate synonymisations under (Quinlan 1974) and (Evenhuis 1982). In Müller et al. (1999) this was recorded as ‘ f1’.

(Hartig, 1841)

Hartig, 1841

England

Synonymised under by Fergusson (1986) but this has not been supported by other workers (Menke and Evenhuis 1991, Van Veen et al. 2003).

(Thomson, 1862)

Thomson, 1862

Kieffer, 1902 (

(Kieffer, 1904,

Belizin, 1966 synonymy by

England, Wales

(Zetterstedt, 1838)

Zetterstedt, 1838

(Hartig, 1841,

(Cameron, 1886,

(Ashmead, 1887,

(Kieffer, 1902,

(Froggatt, 1904,

(De Santis, 1937,

England

Synonymised under by Fergusson (1986) but raised from synonymy by Menke and Evenhuis (1991), a result supported by Van Veen et al. (2003).

(Cameron, 1889)

Cameron, 1889

Scotland

Raised from synonymy with by Ferrer-Suay et al. (2013).

(Hartig, 1840)

Hartig, 1840

(Kieffer, 1904,

England

Considered a nomen dubium in Ferrer-Suay et al. (2014) but Evenhuis (1982) studied the type and subsequently identified British specimens as belonging to this taxon, so we see no problem with considering it a valid British species. It was listed as in Quinlan (1978a) but not mentioned by Fergusson (1986) and then recorded again by Müller et al. (1999).

(Hartig, 1840)

Hartig, 1840

misident.

(Hartig, 1841,

(Thomson, 1862,

England, Scotland

Removed from synonymy with by Pujade-Villar et al. (2011).

(Thomson, 1862)

Thomson, 1862

England

Synonymised under by Fergusson (1986) but considered here to be a valid species (Ferrer-Suay et al. 2012b).

(Hartig, 1841)

Hartig, 1841

England, Wales

added by Van Veen et al. (2003)

(Hartig, 1841)

Hartig, 1841

misident.

misident.

misident.

misident.

misident.

misspelling

(Cameron, 1889,

Kieffer, 1902

Kieffer, 1902 (

England, Scotland, Wales

is recorded as a misidentification of by Fergusson (1986) but erroneously listed as a valid species, occurring in Britain, in Fauna Europaea (Noyes et al. 2004), but with the comment that the species might be synonymous with .

(Kieffer, 1900)

Kieffer, 1900

Scotland

Synonymised under by Fergusson (1986) but considered here to be a valid species (Ferrer-Suay et al. 2012b, following pers. comm. by FVV).

(Cameron, 1883)

Cameron, 1883

Scotland

Raised from synonymy with brevis by Ferrer-Suay et al. (2012a).

(Thomson, 1862)

Thomson, 1862

Wales

added by Baker (2013)

(Hartig, 1840)

Hartig, 1840

Kieffer, 1904 (

England

det. Van Veen, added here.

(Curtis, 1838)

Curtis, 1838

(Zetterstedt, 1838,

(Giraud, 1860,

(Thomson, 1862,

Andrews, 1978

(Curtis, 1838)

Curtis, 1838

England, Wales, Ireland

(Cameron, 1883)

Cameron, 1883

Scotland, Wales

Raised from synonymy with by Ferrer-Suay et al. (2013).

(Hartig, 1840)

Hartig, 1840

(Hartig, 1841,

England

J. Pujade-Villar (pers. comm.) has seen English material.

(Cameron, 1879)

Cameron, 1879

(Kieffer, 1902,

Kieffer, 1922

England, Scotland, Ireland

(Kieffer, 1902)

Kieffer, 1902

England

added by Sanders and Van Veen (2010)

(Thomson, 1862)

Thomson, 1862

(Hartig, 1840,

(Förster, 1869,

(Kieffer, 1904,

England

added by Müller et al. (1999)

Fergusson, 1986

England, Ireland

added by Fergusson (1986)

(Giraud, 1860)

Giraud, 1860

England

Excluded from the British and Irish list by Fergusson (1986) but reinstated by Van Veen et al. (2003).

(Westwood, 1833)

Westwood, 1833

(Curtis, 1838,

(Zetterstedt, 1838,

(Hartig, 1840,

(Fitch, 1861,

(Thomson, 1877,

(Buckton, 1879,

(Buckton, 1879,

(Cameron, 1883,

(Kieffer, 1902,

(Kieffer, 1902,

(Kieffer, 1902,

(Kieffer, 1909,

(Kieffer, 1904,

(Girault, 1932

England, Scotland, Wales

Fergusson, 1986

(Hartig, 1841)

Hartig, 1841

(Thomson, 1862,

England, Wales

Förster, 1869

Haliday in Marshall, 1870

Thomson, 1877

misspelling

Förster, 1869

(Haliday, 1870,

(Thomson, 1877,

(Belizin, 1966,

England, Scotland, Wales

Förster, 1869

Förster, 1869

Thomson, 1877

Kieffer, 1902

Bréthes, 1913

Fergusson (1986) established (including only ) as a synonym of , it was then raised from synonymy by Ronquist (1999) only to be synonymised again with by Pujade-Villar and Paretas-Martínez (2006).

(Cameron, 1889)

Cameron, 1889

England, Ireland

(Hartig, 1840)

Hartig, 1840

Wales

Listed by Quinlan (1978a) as a species of , not mentioned by Fergusson (1986), then recorded as British by Baker (2013) based on specimens identified by M. Ferrer-Suay.

(Förster, 1869)

Förster, 1869

England, Wales

(Thomson, 1877, ) was resurrected as a valid species by Pujade-Villar and Paretas-Martínez (2006). The authors did not examine material from a wide range but state that ‘is only represented by type material’, implying that the real is the widespread one.

(Cameron, 1883)

Cameron, 1883

Cameron, 1888

England, Wales

(Hartig, 1841)

Hartig, 1841

(Thomson, 1862,

(Cameron, 1889,

(Ashmead, 1898,

(Kieffer, 1902,

(Kieffer, 1902,

(Kieffer, 1902,

(Kieffer, 1902,

(Kieffer, 1904,

(Kieffer, 1904,

(Kieffer, 1909,

(Kieffer, 1909,

(Girault, 1931,

(Hellén, 1931,

(Ionescu, 1963

England, Wales

Förster, 1869

(Thomson, 1877,

Kieffer, 1901

England

Thomson, 1862

has been a subfamily of rather than a family of its own since Ronquist (1999). Tribal classification follows Forshage and Nordlander (2008). Synomymy and distribution data taken from Quinlan (1978b) and Nordlander (1978), Nordlander (1980), Nordlander (1981), Forshage and Nordlander (2008), supplemented by Fauna Europaea (Noyes et al. 2004) and MF’s identifications of material in BMNH. Some Irish records from O'Connor (2004).

Belizin, 1961

Förster, 1869

Kieffer, 1904

England, Wales, Ireland

Förster, 1869

Förster, 1869

Förster, 1869

(Giraud, 1860)

Giraud, 1860

England

(Giraud, 1860)

Giraud, 1860

(Thomson, 1862,

Förster, 1869

England, Ireland

Förster, 1869

Kieffer, 1901

(Giraud, 1860,

Scotland

Supposed English specimens in BMNH are all (det. MF).

Thomson, 1862

Westwood, 1833

Agassiz, 1846

Kieffer, 1901

Förster, 1869

Westwood, 1833

(Hartig, 1840,

(Kieffer, 1901,

England, Scotland, Ireland

(Hartig, 1840, Cothonaspis)

Hartig, 1840

Dahlbom, 1842

Belizin, 1973

England, Scotland

Förster, 1869

Synonymy from Nordlander (1980).

(Hartig, 1841)

Hartig, 1841

BMNH, det. Forshage, added here.

(Kieffer, 1901)

Kieffer, 1901

nec (Förster, 1869,

(Cameron, 1883,

(Kieffer, 1904,

(Kieffer, 1904,

(Kieffer, 1907,

(Hellén, 1960,

England

Treated as by Quinlan (1978b).

(Thomson, 1862)

Thomson, 1862

(Rondani, 1875,

(Kieffer, 1904,

(Kieffer, 1904,

(Kieffer, 1916,

(Weld, 1944,

England, Ireland

Treated as by Quinlan (1978b).

(Hartig, 1841)

Hartig, 1841

England

BMNH, det. Forshage, added here.

Förster, 1869

Förster, 1869 synonymy by

Förster, 1869

Förster, 1869

Förster, 1869

Förster, 1869

Förster, 1869

Förster, 1869

Förster, 1869

Ashmead, 1886

Tavares, 1924

Ashmead, 1903

Synonymy mostly from Nordlander (1981). Several new species of are present in Britain and several new synonymies will result from MF’s unpublished revision. Some changes have been made in anticipation of this publication as they affect names described from British specimens.

species of excluded from the British and Irish list:

[ Cameron, 1883] This name will be placed in synonymy.

(Thomson, 1862)

Thomson, 1862

England

BMNH, det. Forshage, added here.

(Thomson, 1861)

Thomson, 1861

Kerrich & Quinlan, 1960

England

(Hartig, 1840)

Hartig, 1840

Cameron, 1883

England, Scotland, Ireland

(Hartig, 1841)

Hartig, 1841

England, Ireland

(Hartig, 1841)

Hartig, 1841

England, Scotland, Wales, Ireland

(Thomson, 1877)

Thomson, 1877

England

BMNH, det. Forshage, added here. Generic combination not yet formally published.

(Thomson, 1862)

Thomson, 1862

England

BMNH, det. Forshage, added here.

(Cameron, 1883)

Cameron, 1883

(Cameron, 1889,

(Kieffer, 1902,

England, Scotland

(Cameron, 1888)

Cameron, 1888

England, Scotland, Wales, Ireland

(Hartig, 1840)

Hartig, 1840

(Dahlbom, 1846,

(Kieffer, 1904,

England, Ireland

(Zetterstedt, 1838)

Zetterstedt, 1838

(Cameron, 1883,

England, Scotland

(Giraud, 1860)

Giraud, 1860

England

BMNH, det. Forshage, added here.

(Westwood, 1835)

Westwood, 1835

(Hartig, 1841,

(Giraud, 1860,

(Thomson, 1862,

Förster, 1869

nec (Latreille, 1805,

(Förster, 1869,

Cameron, 1889

(Cameron, 1889,

(Kieffer, 1901,

(Kieffer, 1902,

nec (Thomson, 1862,

(Kieffer, 1905,

England, Scotland, Wales, Ireland

(Förster, 1855)

Förster, 1855

(Cameron, 1879,

England, Ireland

(Cameron, 1889)

Cameron, 1889

England, Scotland

Recently removed from synonymy with in Forshage et al. (2013).

(Hedicke, 1914)

Hedicke, 1914

England

BMNH, det. Forshage, added here.

(Giraud, 1860)

Giraud, 1860

England

BMNH, det. Forshage, added here.

(Hartig, 1841)

Hartig, 1841

(Kieffer, 1901,

(Thomson, 1862,

England

Belizin, 1961

Förster, 1869

Belizin, 1961

species of excluded from the British and Irish list:

[antennale Kieffer, 1904; England, Ireland] This name will be placed in synonymy

Förster, 1869

England

BMNH, det. Forshage, added here.

Riley, 1879

Ashmead, 1896

(Kieffer, 1904)

Kieffer, 1904

England, Wales

BMNH, det. Forshage, added here.

Förster, 1869

Förster, 1869

Kieffer, 1904

England

BMNH, det. Forshage, added here.

(Hellén, 1960)

Hellén, 1960

(Belizin, 1968,

England, Scotland

BMNH, det. Forshage, added here.

Thomson, 1861

Förster, 1869

Förster, 1869

Förster, 1869

Kieffer, 1904

Rohwer & Fagan, 1917

(Hartig, 1840)

Hartig, 1840

(Hartig, 1841,

(Dahlbom, 1842,

Kieffer, 1901

(Kieffer, 1902,

(Kieffer, 1904,

(Kieffer, 1904,

Ionescu, 1963

England, Wales, Ireland

Förster, 1869

Förster, 1869

(Hartig, 1841)

Hartig, 1841

(Giraud, 1860,

(Mayer, 1923,

England

Belizin, 1968

Quinlan, 1975

(Thomson, 1877)

Thomson, 1877

(Kieffer, 1901,

(Hedicke, 1913,

England, Ireland

Hellén, 1960

Hartig, 1840

Kieffer, 1901

Hartig, 1841

(Dalla Torre & Kieffer, 1910,

England

Nordlander, 1976

England

Hartig, 1841

England, Scotland

Förster, 1869

(Thomson, 1877)

Thomson, 1877

England

Westwood, 1833

Förster, 1869

Förster, 1869

Förster, 1869

Förster, 1869

Förster, 1869

Förster, 1869

Förster, 1869

Förster, 1869

Cameron, 1890

Kieffer, 1901

Kieffer, 1901

Kieffer, 1901

Rohwer & Fagan, 1917

Weld, 1951

Subgeneric classification was finally abandoned and all subgeneric names considered mere synonyms in Forshage and Nordlander (2008).

Species of removed from the British and Irish list:

[myrmecophila Kieffer, 1908] Recorded from Britain in Fauna Europaea (Noyes et al. 2004) but in error.

Cameron, 1889

Scotland

Cameron, 1888

England, Scotland, Ireland

(Zetterstedt, 1838)

Zetterstedt, 1838

Cameron, 1888

England, Scotland

Usually regarded as a species of but published as a new combination in by Jonsell et al. (1999) in an ecological paper and listed as such in Fauna Europaea (Noyes et al. 2004). The Zetterstedt, refers to various small spp.

Thomson, 1877

(Kieffer, 1904,

England, Wales, Ireland

Cameron, 1889

England, Scotland

Cameron, 1889

England

Cameron, 1889

England

Thomson, 1861

Thomson, 1861

England

Thomson, 1862

England, Scotland

Cameron, 1889

England, Scotland

Cameron, 1889

England, Scotland, Ireland

Cameron, 1889

Belizin, 1964

England, Ireland

(Hartig, 1840)

Hartig, 1840

Cameron, 1888

Cameron, 1888

(Kieffer, 1904,

(Kieffer, 1904,

England, Scotland, Ireland

Thomson, 1861

Cameron, 1886

England, Scotland, Ireland

Cameron, 1886

Scotland, Ireland

Westwood, 1833

England, Wales

(Dahlbom, 1842)

Dahlbom, 1842

England, Ireland

Cameron, 1886

England, Scotland

Cameron, 1880

England, Scotland

(Walker, 1834)

Walker, 1834

(Förster, 1869,

England, Wales, Ireland

Forshage, sp. nov., replacement name

Thomson, 1861 preocc. Name preoccupied by

England, Scotland, Ireland

(Giraud, 1860)

Giraud, 1860

(Förster, 1869,

Thomson, 1877

England

Cameron, 1889

England, Scotland

Kovalev, 1989

Förster, 1869

Förster, 1869

Kieffer, 1901

Belizin, 1966

Synonymy taken from Nordlander (1978) .

Species of removed from the British and Irish list:

[ Nordlander, 1978] was listed by Quinlan (1978b) but this was most likely a mistake. Nordlander (1978) mentions that some Swedish paratypes were deposited in BMNH. There are no British or Irish specimens identified as in the collection.

(Hartig, 1840)

Hartig, 1840

(Hartig, 1840,

(Hartig, 1841,

(Giraud, 1860,

(Thomson, 1862,

(Förster, 1869,

(Kieffer, 1901,

Kurdjumov, 1912

England, Scotland, Wales, Ireland

(Hartig, 1840)

Hartig, 1840

England, Wales, Ireland

BMNH, det. Forshage, added here.

Benoit, 1956

BMNH, det. Forshage, added here. Only a single male specimen has been found amongst British material; it is not identifiable to species level with our current state of knowledge, and is listed here as a record of the genus from Britain.

Hartig, 1840

has been transferred to by Buffington et al. (2007).

Synonymy follows Fergusson (1986) except where otherwise noted.

Hartig, 1840

misspelling

Removed from synonymy with by Ronquist (1999).

(Hartig, 1840)

Hartig, 1840

Hartig, 1840

(Dahlbom, 1842,

(Giraud, 1860,

England

Latreille, 1802

Hartig, 1840

Förster, 1869

Hedicke, 1913

Species of excluded from the British and Irish list:

[ Dahlbom, 1842]

[ Kieffer, 1901]

[ Dahlbom, 1842]

Included in Quinlan (1978a) but not by Fergusson (1986).

Bouché, 1834

England, Ireland

Giraud, 1860

misident.

Hellén, 1937

England, Ireland

Fergusson, 1986

England, Ireland

added by Fergusson (1986)

(Rossi, 1794)

Rossi, 1794

(Rossi, 1794,

(Herrich-Schäffer in Panzer, 1801,

(Hartig, 1840,

(Hedicke, 1913,

England

Thomson, 1862

Thomson, 1862

England, Scotland, Ireland

(Dahlbom, 1842)

Dahlbom, 1842

England, Ireland

Hartig, 1840

(Zetterstedt, 1838)

Zetterstedt, 1838

Hartig, 1840

(Hedicke, 1913,

England, Wales, Ireland

Kieffer, 1901

(Giraud, 1860)

Giraud, 1860

Wales, Ireland

added by Fergusson (1986)

Förster, 1869

Förster, 1869

Ashmead, 1887

(Dahlbom, 1842)

Dahlbom, 1842

(Hartig, 1843,

England, Ireland

Thomson, 1862

Latreille, 1802

Panzer, 1806

(Hockenwarth, 1785)

Hockenwarth, 1785

(Fabricius, 1793,

Norton, 1862

Jacobson, 1899

Yoshimoto, 1970

England

Cresson, 1879

Borries, 1891

Dalla Torre & Kieffer, 1910

England, Scotland

Checklist of the British and Irish

Data type: formatted text

Brief description: Word document version of the checklist

File: oo_124446.docx

Checklist of British and Irish

Data type: spreadsheet

Brief description: Excel spreadsheet version of the checklist

File: oo_124447.xlsx