Checklist of British and Irish Hymenoptera - aculeates (Apoidea, Chrysidoidea and Vespoidea).

BACKGROUND: The checklist of British and Irish aculeate Hymenoptera (Apoidea, Chrysidoidea and Vespoidea) is revised. Species distribution is summarised for all species at the level of country (England, Scotland, Wales, Ireland and Isle of Man). NEW INFORMATION: The 601 native species represent an increase of 25 on the 1978 checklist, comprising mostly new discoveries. This increase is nearly balanced by the 23 species now presumed to be extinct in Britain and Ireland.

Introduction

The checklist of British and Irish aculeates is essentially that of Else et al. (2004) but with several additions and updated taxonomy. This continues the series of chapters, starting with Broad and Livermore (2014a), Broad and Livermore (2014b) and Liston et al. (2014) comprising the first complete updating of the 1978 checklist (Fitton et al. 1978).

Unlike most groups of , the aculeate ants, bees and wasps are well-served by identification literature and are actively collected and recorded by a relatively large number of amateur as well as professional entomologists. We thus have a better idea of the actual size of the fauna than we do for sawflies and parasitoid wasps. There is thus a much better idea of change in the aculeate fauna and numerous extinctions and colonisations have been documented.

The aculeate encompass three superfamilies and 18 families in Britain and Ireland (as discussed in Broad 2014, this figure is likely to change), with a wide range of ecologies and body forms. Figs 1, 2, 3, 4, 5, 6 depict some of this variety (and the website of the Bees, Wasps and Ants Recording Society has many more images). With multiple evolutions of eusociality, importance as pollinators, predators and parasitoids and general conspicuousness, the aculeates have always attracted attention and very good summaries of their biology can be found in a diverse array of texts for a variety of different audiences, from Lubbock (1882) to Gauld and Bolton (1988).

Materials and methods

The rationale and background to the updated checklist are laid out in Broad (2014). The aim is to account for all the names in the 1978 checklist, document all additions to and deletions from that list and to list the distributions of all species at the country level, i.e. England, Scotland, Wales, Ireland (as one unit) and the Isle of Man. We do not usually report distribution data for introduced species that only reproduce indoors or in greenhouses.

Sources of classification are listed under each superfamily. The checklist of bees is essentially similar to that of Else's (in prep.) handbook to the British fauna, except that we employ six families rather than including all bees in and, as discussed in the introduction to this checklist series (Broad 2014), we do not consider the fauna of the Channel Islands. We adopt a multi-family classification of the bees to maintain unity with larger-scale checklists (particularly that of M. Kuhlmann's Checklist of the Western Palaearctic Bees) and because this is the system employed in the most authoritative treatise on bee classification (Michener 2007), as well as most recent papers dealing with bee phylogeny.

Country-level distribution data mostly derive from the distribution atlases produced by the Bees, Wasps and Ants Recording Society (Edwards 1997, Edwards 1998, Edwards and Telfer 2001, Edwards and Telfer 2002, Edwards and Broad 2005, Edwards and Broad 2006, Edwards and Roy 2009, Collins and Roy 2012, with maps updated on their website), the catalogue of Irish (O'Connor et al. 2009), and some additional literature references. Note that the , and are particularly poorly recorded.

The following abbreviations and conventions are employed:

[] taxon deleted from the British and Irish list

NHM Natural History Museum, London

# known or suspected introductions with at least temporarily self-sustaining populations

? status (including uncertain synonymy) or identification in the British Isles uncertain

misident. has been misidentified as this name

nom. dub. nomen dubium, a name of unknown or doubtful application

nom. ob. nomen oblitum, ‘forgotten name’, does not have priority over a younger name

nom. nov. nomen novum, a replacement name

nom. nud. nomen nudum, an unavailable name, with no type specimen

preocc. name preoccupied (junior homonym)

stat. rev. status revocatus, revived status (e.g., raised from synonymy)

unavailable not meeting the requirements of the International Code of Zoological Nomenclature

var. variety, only available as a valid name under the provisions of article 45.6 of the ICZN

Text and spreadsheet versions of the checklists are available as supplementary files: Suppl. materials 1, 2.

Checklists

Latreille, 1802

Notes

The higher-level classification of the follows Pulawski (2013) Engel (2005) and M. Kuhlmann's online checklist of Western Palaearctic bees. The ‘traditional’ family is paraphyletic with respect to the bee families (here referred to as the , an informal taxon, following Engel 2005) and in Britain and Ireland comprises the families and . Species-level classification of and follows Pulawski (2013). Figs 1, 2: habitus of selected British .

Latreille, 1802

Lepeletier, 1845

Latreille, 1796

Panzer, 1806

(Schrank, 1781)

Schrank, 1781

(Panzer, 1798,

(Panzer, 1804,

(Jurine, 1807,

Curtis, 1829

Robert, 1833

Smith, 1875

Distribution

England

Spinola, 1843

(Dahlbom, 1832)

Dahlbom, 1832

misident.

(Zetterstedt, 1838,

(Smith, 1845,

England, Scotland, Wales

Latreille, 1802

Dalla Torre, 1897

Wesmael, 1852

(Fabricius, 1798)

Fabricius, 1798

(Curtis, 1836,

England

Latreille, 1802

Ashmead, 1899

(Shuckard, 1837)

Shuckard, 1837

misident.

Tsuneki, 1971

Kazenas, 1971

England, Wales

(Linnaeus, 1761)

Linnaeus, 1761

(Linnaeus, 1761,

(Harris, 1776,

(Rossi, 1790,

(Fabricius, 1793,

(Fabricius, 1804,

(Donovan, 1808,

(Eversmann, 1849,

(Bondroit, 1933,

England, Scotland, Wales, Ireland

Latreille, 1805

Panzer, 1805

Stephens, 1829

Lepeletier, 1832

Lepeletier, 1832

Zavadil, 1948

(Lepeletier, 1832)

Lepeletier, 1832

England, Wales

(Fabricius, 1804)

Fabricius, 1804

(Lepeletier, 1832,

(Mocsáry, 1878,

England, Wales

Shuckard, 1837

Jurine, 1807

Dahlbom, 1843

Fox, 1893

(Panzer, 1801)

Panzer, 1801

(Tsuneki, 1963,

Kazenas, 1989

England, Scotland, Wales, Ireland, Isle of Man

Lepeletier, 1832

Asmead, 1899

Asmead, 1899

(Rossi, 1794)

Rossi, 1794

England

Latreille, 1804

Latreille, 1796

Costa, 1859

Jurine, 1807

Lepeletier, 1845

Chevrier, 1867

England, Wales

(Fabricius, 1798)

Fabricius, 1798

(Fabricius, 1804,

Lepeletier, 1845

Wesmael, 1852

England

(Forster, 1771)

Forster, 1771

(Linnaeus, 1767,

(Fabricius, 1775,

(Fabricius, 1793,

(Turton, 1802,

Lepeletier, 1845

Gussakovskij, 1932

England, Scotland, Wales, Ireland

(Rossi, 1790)

Rossi, 1790

England, Wales

Latreille, 1802

Latreille, 1802

Fabricius, 1775

Lepeletier & Brullé, 1835

Dahlbom, 1845

Costa, 1871

Ashmead, 1899

Ashmead, 1899

Pate, 1944

Pate, 1944

Pate, 1944

Pate, 1947

Pate, 1944

Pate, 1944

Pate, 1944

Marshakov, 1977

(Linnaeus, 1758)

Linnaeus, 1758

(Schreber, 1784,

(Christ, 1791,

(Christ, 1791,

(Christ, 1791,

Panzer, 1797

Mocsáry, 1901

Kokujev, 1927

England, Scotland, Wales, Isle of Man

(Schreber, 1784)

Schreber, 1784

Panzer, 1797

Panzer, 1797

Fabricius, 1798

England, Scotland, Wales, Ireland, Isle of Man

(von Scheven, 1781)

von Scheven, 1781

(Schreber, 1784,

Panzer, 1801

(Lepeletier & Brullé, 1835,

England

Lepeletier & Brullé, 1835

Perkins, 1913

(Dahlbom, 1844)

Dahlbom, 1844

England

added by Archer (2007) based on a pers. comm. from D. Baldock

(Vander Linden, 1829)

Vander Linden, 1829

Dahlbom, 1842

Šnoflak, 1948

Zavadil, 1948

England, Wales

Perkins, 1913

Tsuneki, 1974

(Panzer, 1798)

Panzer, 1798

(Panzer, 1799,

(Schrank, 1802,

(Lepeletier & Brullé, 1835,

Lepeletier & Brullé, 1835

(Costa, 1871,

(Yasumatsu, 1942,

(Tsuneki, 1974,

England, Wales

Lepeletier & Brullé, 1835

Thomson, 1874

(Lepeletier & Brullé, 1835)

Lepeletier & Brullé, 1835

(Lepeletier & Brullé, 1835,

(Gimmerthal, 1836,

(Dahlbom, 1842,

(Dahlbom, 1845,

(Banks, 1921,

(Sandhouse, 1938,

England, Scotland, Wales

(Shuckard, 1837)

Shuckard, 1837

(Dahlbom, 1838,

Tsuneki, 1960

England, Scotland, Wales, Ireland

(Shuckard, 1837)

Shuckard, 1837

(Dahlbom, 1838,

(Herrich-Schäffer, 1841,

(Matsumura, 1912,

Tsuneki, 1976

England, Wales, Ireland

(Linnaeus, 1758)

Linnaeus, 1758

(Dahlbom, 1838,

(Herrich-Schäffer, 1841,

(Wesmael, 1852,

(Provancher, 1882,

(Provancher, 1888,

(Fox, 1895,

(Dalla Torre, 1897,

(Ashmead, 1901,

(Bradley, 1906,

(Mickel, 1916,

(Mickel, 1916,

England, Scotland

(Rossi, 1790)

Rossi, 1790

misident.

(Haliday, 1833,

Lepeletier & Brullé, 1835

Lepeletier & Brullé, 1835

(Smith, 1856,

(Costa, 1884,

(Radoszkowski, 1887,

(Richards, 1935,

England, Scotland, Wales, Ireland, Isle of Man

(Lepeletier & Brullé, 1835)

Lepeletier & Brullé, 1835

(Shuckard, 1837,

Herrich-Schäffer, 1841

(Thomson, 1870,

Kohl, 1880

(Provancher, 1888,

(Verhoeff, 1891,

Tsuneki, 1967

Tsuneki, 1976

Tsuneki, 1979

England, Wales

(Kohl, 1892)

Kohl, 1892

(Gussakovskij, 1932,

(Iwata, 1938,

Tsuneki, 1977

England

(Shuckard, 1837)

Shuckard, 1837

misident.

(Shuckard, 1837,

(Schenck, 1857,

(Nielson, 1901,

England, Scotland, Wales

Lepeletier & Brullé, 1835

Ashmead, 1899

Holmberg, 1903

(Morawitz, 1866)

Morawitz, 1866

(Thomson, 1870,

England

added by Packer (1981)

(Vander Linden, 1829)

Vander Linden,

Lepeletier & Brullé, 1835

Lepeletier & Brullé, 1835

Lepeletier & Brullé, 1835

Lepeletier & Brullé, 1835

Lepeletier & Brullé, 1835

Lepeletier & Brullé, 1835

(Shuckard, 1837,

(Shuckard, 1837,

(Shuckard, 1837,

(Shuckard, 1837,

(Shuckard, 1837,

(Eversmann, 1849,

(Smith, 1851,

(Fox, 1895,

(Rohwer, 1912,

(Richards, 1928,

England, Scotland, Wales, Isle of Man

The British population is considered to belong to the subspecies Lepeletier & Brullé (synonyms: ).

(Vander Linden, 1829)

Vander Linden,

Lepeletier & Brullé, 1835

England

Lepeletier & Brullé, 1835

(Zetterstedt, 1838,

(Wesmael, 1852,

(Smith, 1856,

(Schulz, 1906,

England, Scotland, Wales

(Linnaeus, 1767)

Linnaeus, 1767

(Schreber, 1784,

(Fabricius, 1787,

Lepeletier & Brullé, 1835

(Gimmerthal, 1836,

(Eversmann, 1849,

(Smith, 1856,

England, Scotland, Isle of Man

(Shuckard, 1837)

Shuckard, 1837

misident.

De Stefani Perez, 1884 preocc,

England, Scotland, Wales, Ireland, Isle of Man

Lepeletier & Brullé, 1835 nomen protectum

misspelling

Lepeletier & Brullé, 1835

Lepeletier & Brullé, 1835

(Shuckard, 1837,

Lepeletier, 1845

(Smith, 1856,

(Smith, 1856,

England, Scotland, Wales, Ireland, Isle of Man

(Vander Linden, 1829)

Vander Linden,

(Lepeletier & Brullé, 1835,

(Lepeletier & Brullé, 1835,

England, Scotland, Wales, Ireland

Morawitz, 1866

Lepeletier & Brullé, 1835

(Panzer, 1798,

(Dahlbom, 1845,

(Schulz, 1906,

England, Wales

(Fabricius, 1781)

Fabricius, 1781

(Rossi, 1790,

(Olivier, 1792,

(Olivier, 1792,

(Panzer, 1797,

(Illiger, 1807,

(Lepeletier & Brullé, 1835,

(von Sebold, 1844,

England, Scotland, Wales, Ireland

Thomson, 1874

(Fabricius, 1793)

Fabricius, 1793

(Fabricius, 1793,

(Latreille, 1805,

Vander Linden, 1829,

Lepeletier & Brullé, 1835

(Dahlbom, 1838,

(Dahlbom, 1838,

England, Scotland, Wales, Ireland

Dahlbom, 1845

Morawitz, 1864

(Thomson, 1870)

Thomson, 1870

misident.

England, Scotland, Wales, Ireland, Isle of Man

(Panzer, 1804)

Panzer, 1804

(Spinola, 1806,

(Lepeletier & Brullé, 1835,

(Lepeletier & Brullé, 1835,

(Shuckard, 1837,

(Smith, 1856,

(Smith, 1856,

(Packard, 1866,

(Packard, 1866,

(Packard, 1866,

(Viereck, 1908,

England, Scotland, Wales, Ireland, Isle of Man

(Zetterstedt, 1838)

Zetterstedt, 1838

(Herrich-Schäffer, 1841,

(Cresson, 1865,

(Cresson, 1865,

(Packard, 1866,

(Thomson, 1870,

(Verhöff, 1892,

(Matsumura, 1911,

(Matsumura, 1912,

England, Wales

(Fabricius, 1775)

Fabricius, 1775

misident.

(Fabricius, 1787,

(Retzius, 1783,

(Olivier, 1792,

(Preyssler, 1793,

(Panzer, 1797,

(Panzer, 1798,

(Schrank, 1802,

(Spinola, 1806,

(Lepeletier & Brullé, 1835,

(Blanchard, 1940,

(Perkins, 1899,

Pate, 1946

England, Wales

Dahlbom, 1845

(Zetterstedt, 1838)

Zetterstedt, 1838

(Zetterstedt, 1838,

(Herrich-Schäffer, 1841,

(Packard, 1866,

(Kohl, 1878,

(Mickel, 1916,

England

(Lepeletier & Brullé, 1835)

Lepeletier & Brullé, 1835

(Lepeletier & Brullé, 1835,

(Lepeletier & Brullé, 1835,

(Dahlbom, 1838,

(Herrich-Schäffer, 1841,

(Smith, 1856,

(Cresson, 1865,

(Packard, 1866,

(Packard, 1866,

(Rohwer, 1908,

(Strand, 1916,

England, Wales

Ashmead, 1899

Ashmead, 1899

(Fabricius, 1804)

Fabricius, 1804

misident.

(Say, 1824,

(Herrich-Schäffer, 1841,

(Smith, 1869,

(Costa, 1871,

(Walker, 1871,

(Taschenberg, 1875,

(De Stefani Perez, 1884,

(Viereck, 1910,

(Cockerell, 1910,

(Rohwer, 1917,

England, Scotland, Wales, Ireland

(Dufour & Perris, 184)

Dufour & Perris, 1840

(Herrich-Schäffer, 1841,

(Wesmael, 1852,

(Costa, 1871,

England, Wales

Ashmead, 1899

(Olivier, 1792)

Olivier, 1792

misident.

(Olivier, 1792,

(Olivier, 1792,

(Olivier, 1792,

(Panzer, 1799,

(Lepeletier & Brullé, 1835,

(Lepeletier & Brullé, 1835,

(Lepeletier & Brullé, 1835,

(Shuckard, 1837,

(Dahlbom, 1838,

(Dahlbom, 1845,

(Smith, 1856,

(Provancher, 1882,

(Morawitz, 1892,

England, Scotland, Wales, Ireland

(Panzer, 1805)

Panzer, 1805

(Lepeletier & Brullé, 1835,

(Lepeletier & Brullé, 1835,

(Lepeletier & Brullé, 1835,

(Dahlbom, 1845,

(Schenck, 1857,

(Smith, 1858,

(Morawitz, 1866,

(Costa, 1871,

England, Wales

Dahlbom, 1844

Pate, 1944

Pate, 1944

Pate, 1944

Leclercq, 1956

Leclercq, 1977

(Vander Linden, 1829)

Vander Linden, 1829

(Lepeletier & Brullé, 1835,

(Gribodo, 1884,

England, Wales

Billberg, 1820

Richards, 1935

Lepeletier & Brullé, 1835 preocc.

(Schreber, 1759)

Schreber, 1759

(Christ, 1791,

(Panzer, 1797,

(Fabricius, 1804,

(Thunberg, 1815,

(Herrich-Schäffer, 1841,

(Dufour, 1841,

England

Lepeletier & Brullé, 1835

Wesmael, 1852

Morawitz, 1866

(Fabricius, 1793)

Fabricius, 1793

(Dahlbom, 1838,

England, Scotland, Wales

(Vander Linden, 1829)

Vander Linden, 1829

Lepeletier & Brullé, 1835

(Kohl, 1905,

Tsuneki, 1967

Tsuneki, 1972

England

(Rossi, 1794)

Rossi, 1794

Lepeletier & Brullé, 1835

(Šnoflak, 1948,

England

Represented by the subspecies armatus (Vander Linden, 1829, ).

Stephens, 1829

Stephens, 1829

Risso, 1826 nom. ob.

Lepeletier & Brullé, 1835

(Linnaeus, 1758)

Linnaeus, 1758

(Panzer, 1799,

England, Scotland, Wales, Ireland, Isle of Man

Lepeletier & Brullé, 1835

Herrich-Schäffer, 1840

Ashmead, 1899

(Scopoli, 1763)

Scopoli, 1763

(Fabricius, 1798,

(Fabricius, 1798,

Rohwer, 1908

England, Scotland, Wales, Ireland

Wesmael, 1852

Kiesenwetter, 1849 preocc.

(Morawitz, 1866,

(Morawitz, 1888,

England

Latreille, 1810

Kohl, 1883

Arnold, 1922

Arnold, 1923

(Spinola, 1805)

Spinola, 1805

misident.

(de Saussure, 1867,

England, Wales, Isle of Man

Added by Edwards (1998); earlier records of mostly refer to .

(Schenck, 1857)

Schenck, 1857

(Thomson, 1870,

England

Only recorded as a vagrant in Deal, Kent; resident on the Channel Isles, which lie outside of the geographical coverage of this checklist.

(Panzer, 1805)

Panzer, 1805

misident.

(Spinola, 1808,

(Panzer, 1809,

(Panzer, 1809,

(Cresson, 1865,

(Provancher, 1882,

Fox, 1894

Fox, 1894

Fox, 1894

Bingham, 1897

Nurse, 1903

Rohwer, 1911

Mickel, 1916

Mickel, 1916

Mickel, 1916

England, Scotland, Wales, Ireland, Isle of Man

(Panzer, 1809)

Panzer, 1809

misident.

(Drapiez, 1819,

England

Fox, 1894

Jurine, 1807

Ashmead, 1897

Cockerell, 1898

Ashmead, 1898

Lepeletier, 1845

Smith, 1858

England

Jurine, 1807

(Panzer, 1809,

Verhöff, 1890

de Andrade, 1960

England

added by Knowles and Else (2006)

Dahlbom, 1844

misident.

Balthazar, 1957

England

Latreille, 1809

Brèthes, 1913

Williams, 1928

Valkeila, 1974

England

added by Felton (1987)

Gayubo & Felton, 2000

misident.

England

Added by Felton (1987), initially as Latreille, 1809 but described later as a new species (Gayubo and Felton 2000).

Leach, 1815

Latreille, 1796

Dahlbom, 1845

Costa, 1882

Kohl, 1924

Pate, 1937

Pate, 1937

Noskiewicz & Chudoba, 1950

Curtis, 1833

misident.

(Donovan, 1806,

Shuckard, 1837

Shuckard, 1837

England, Wales

Dahlbom, 1845

Gerstäcker, 1867

England, Scotland, Wales, Isle of Man

(Linnaeus, 1758)

Linnaeus, 1758

(Fabricius, 1793,

(Fabricius, 1798,

(Donovan, 1806,

Olivier, 1812

Say, 1824

Smith, 1856

Cresson, 1865

Gerstäcker, 1867

Provancher, 1883

Giner Marí, 1943

England, Scotland, Wales, Ireland, Isle of Man

Lepeletier, 1845

Latreille, 1796

Panzer, 1806

Jurine, 1807

Richards, 1934

Arnold, 1959

Smith, 1851

England, Scotland, Wales, Ireland

Lepeletier & Serville, 1828

Zetterstedt, 1840

Antropov, 1985

England, Wales, Ireland

(Linnaeus, 1758)

Linnaeus, 1758

(Scopoli, 1763,

Fox, 1891

Tsuneki, 1956

de Beaumont, 1957

Giner Marí, 1959

England, Wales

de Beaumont, 1945

England, Scotland, Wales

added by Pulawski (1984)

de Beaumont, 1945

Tsuneki, 1956

England

added by Felton (1988)

Fox, 1895

Panzer, 1806

Jurine, 1807 preocc.

(Fabricius, 1793)

Fabricius, 1793

(Fabricius, 1793,

England

Latreille, 1802

Fabricius, 1790

(Linnaeus, 1758)

Linnaeus, 1758

(Linnaeus, 1758,

(Harris, 1776,

(Schrank, 1781,

(Retzius, 1783,

(Geoffroy, 1785,

(Geoffroy, 1785,

(Fabricius, 1787,

(Villers, 1789,

(Gmelin, 1790,

(Rossi, 1790,

(Rossi, 1790,

(Christ, 1791,

(Olivier, 1792,

(Olivier, 1792,

(Olivier, 1792,

(Fabricius, 1793,

(Panzer, 1794,

(Schrank, 1796,

Jurine, 1807

Handlirsch, 1888

England, Scotland, Wales, Ireland, Isle of Man

(Thunberg, 1791)

Thunberg, 1791

(Fabricius, 1787,

(Villers, 1789,

(Olivier, 1792,

Fabricius, 1793 preocc.

(Panzer, 1797,

(Panzer, 1797,

Fabricius, 1804

England, Wales

Dahlbom, 1835

Dahlbom, 1835

Curtis, 1834

Rohwer, 1915

Tsuneki, 1972

Tsuneki, 1974

Spooner, 1938

misident.

England

Shuckard, 1837

England, Wales

(Fabricius, 1793)

Fabricius, 1793

Krombein, 1939

England, Wales, Isle of Man

(Vander Linden, 1829)

Vander Linden, 1829

England, Scotland, Wales

Shuckard, 1837

Shuckard, 1837

Verhöff, 1890

Verhöff, 1890

Faester, 1947

Gussakovskij, 1952

England

Shuckard, 1837

England, Wales

Kohl, 1893

England

added by Richards (1980)

(Curtis, 1834)

Curtis, 1834

misident.

misident.

Morawitz, 1864

England

(Vander Linden, 1829)

Vander Linden, 1829

Verhöff, 1890

Yasumatsu, 1934

England, Wales

Dahlbom, 1842

England, Scotland, Wales, Ireland

Dahlbom, 1844

misident.

Morawitz, 1864

Krombein, 1938

England, Wales

Dahlbom, 1844

England

added by Guichard (2002)

Latreille, 1796

Panzer, 1806

Westwood, 1837

Shuckard, 1837

Westwood, 1840

Kohl, 1883

Tsuneki, 1972

Taxonomy follows Dollfuss (1995).

(Kohl, 1888)

Kohl, 1888

misident.

Valkeila, 1972

Valkeila, 1972

Merisuo, 1972

England

Richards (1980) refers to this species as (Wagner, 1932, ). There has been much confusion over the identities of these taxa (see Knowles in Collins and Roy 2012) but Pulawski (2013) treats and as separate species.

Say, 1824

(Morawitz, 1864,

(Puton, 1871,

Fox, 1829

England, Scotland, Wales, Ireland

(Shuckard, 1837)

Shuckard, 1837

misspelling

misident.

(Chevrier, 1870,

(Müller, 1911,

(Wagner, 1918,

(Wagner, 1918,

(Wagner, 1918,

(Wagner, 1918,

Wagner, 1931

Wagner, 1932

Gussakovskij, 1952

Gussakovskij, 1952

Merisuo, 1972

Merisuo, 1972

Valkeila, 1972

Valkeila, 1972

Valkeila, 1972

Tsuneki, 1977

England, Scotland, Wales, Ireland, Isle of Man

(Fabricius, 1793)

Fabricius, 1793

Say, 1824

Gimmerthal, 1836

Shuckard, 1837

Cresson, 1865 preocc.

Dalla Torre, 1897

Dalla Torre, 1897

Cameron, 1908

Rohwer, 1917

Gussakovkij, 1932

England, Scotland, Wales, Ireland, Isle of Man

Vander Linden, 1829

(Smith, 1851,

Thomson, 1870

Thomson, 1870

Tsuneki, 1951

England, Wales

(Dahlbom, 1844)

Dahlbom, 1844

misspelling

(Panzer, 1798,

(Gimmerthal, 1836,

(Morawitz, 1864,

Perkins, 1929

(Bondroit, 1931,

Gussakovskij, 1952

Valkeila, 1972

Valkeila, 1972

Valkeila, 1972

England, Scotland

Shuckard, 1838

Shuckard, 1837 preocc.

Rayment, 1930 preocc.

Rayment, 1935

Tsuneki, 1971

Blüthgen, 1953

Valkeila, 1957

England

Blüthgen, 1953

misident.

England

Has been called (Dahlbom, 1844, ) in several publications, following the synonymy by Dollfuss (1986), but was reinstated as a valid species by Vikberg (2000).

(Dahlbom, 1844)

Dahlbom, 1844

Dolfuss, 1983

Tsuneki, 1971

England

Blüthgen, 1953

England, Ireland

(Vander Linden, 1829)

Vander Linden, 1829

(Radoszkowski, 1877,

Blüthgen, 1953

England, Scotland, Wales

Panzer, 1805

Zetterstedt, 1838

Rohwer, 1911

Krombein, 1973

Panzer, 1804

Jurine, 1807

England

added by Allen (1987)

Morawitz, 1864

Tsuneki, 1954

Tsuneki, 1954

England, Wales

Costa, 1858

Shuckard, 1837

Wesmael, 1852 preocc.

Gussakovskij, 1937 preocc.

(Jurine, 1807)

Jurine, 1807

misident.

(Panzer, 1805,

England

Bondroit, 1934

(Gussakovskij, 1937,

England, Wales

(Fabricius, 1804)

Fabricius, 1804

misident.

England, Scotland, Wales

(Fabricius, 1787)

Fabricius, 1787

Wesmael, 1852

Packard, 1867

Smith, 1908

(Gussakovskij, 1937,

Finnamore, 1980

England, Wales

Malloch, 1933

(Morawitz, 1891)

Morawitz, 1891

(Tournier, 1899,

(Bondroit, 1932,

(Gussakovskij, 1932,

(Yasumatsu, 1937,

England

(Wesmael, 1852)

Wesmael, 1852

England, Scotland, Wales

(Bondroit, 1934)

Bondroit, 1934

misident.

(Gussakovskij, 1934,

(Gussakovskij, 1937,

(Spooner, 1948,

England, Wales, Ireland

(Richards, 1948)

Richards, 1948

England

(Vander Linden, 1829)

Vander Linden, 1829

(Dahlbom, 1842,

(Saunders, 1904,

(van Lith, 1976,

England

added by Else and Felton (1994)

Latreille, 1796

Rafinesque, 1815

Stephens, 1829

Wissmann, 1849

Wesmael, 1852

Cameron, 1899

(Olivier, 1792)

Olivier, 1792

(Fabricius, 1794,

(Fabricius, 1804,

(Jurine, 1807,

(Jurine, 1807,

England

Kohl, 1897

misident.

Viereck, 1901

Schulz, 1911

Gussakovskij, 1934

Yasumatsu, 1938

is a misidentification by American authors in the older literature

(Dahlbom, 1843)

Dahlbom, 1843

(Schenck, 1857,

(Schenck, 1857,

England, Wales

(Panzer, 1798)

Panzer, 1798

(Fabricius, 1804,

(Costa, 1861,

(Costa, 1871,

(Tournier, 1899,

(Tournier, 1899,

(Tournier, 1899,

(Tournier, 1899,

Harttig, 1931

Merisuo, 1937

England, Wales, Ireland

(Tournier, 1889)

Tournier, 1889

(Tournier, 1899,

(Tournier, 1899,

England

Latreille, 1802

Lepeletier, 1845

Latreille, 1802

Spinola, 1839

Dahlbom, 1844

Dahlbom, 1845

Shestakov, 1923

Brèthes, 1913

Minkiewicz, 1934

Gussakovskij, 1935

Pate, 1937

(Linnaeus, 1758)

Linnaeus, 1758

(Forster, 1771,

(Harris, 1776,

(Harris, 1776,

(Fabricius, 1781,

(Gmelin, 1790,

(Fabricius, 1794,

Schletterer, 1887

England, Wales

(Panzer, 1799)

Panzer, 1799

Spinola, 1806

England

(Rossi, 1792)

Rossi, 1792

misident.

Dahlbom, 1844 preocc.

Lepeletier, 1845

England

(Fabricius, 1793)

Fabricius, 1793

misident.

(Schrank, 1802,

(Schrank, 1802,

(Fabricius, 1804,

(Fabricius, 1804,

Latreille, 1809

Costa, 1867

England

(Linnaeus, 1771)

Linnaeus, 1771

(Fabricius, 1790,

(Christ, 1791,

(Panzer, 1797,

(Panzer, 1799,

(Schrank, 1802,

(Thunberg, 1815,

(Thunberg, 1815,

Nurse, 1903

Schulz, 1904

Eck, 1979

England

(Panzer, 1799)

Panzer, 1799

(Panzer, 1799,

(Spinola, 1805,

(Thunberg, 1815,

Lepeletier, 1845

Shestakov, 1923

England

Latreille, 1802

Fabricius, 1790

Panzer, 1806

Jurine, 1807

Westwood, 1835

Dahlbom, 1844

Ashmead, 1899

Ashmead, 1899

Banks, 1913

(Fabricius, 1775)

Fabricius, 1775

(Linnaeus, 1767,

(Geoffroy, 1785,

(Christ, 1791,

(Olivier, 1792,

(Rossi, 1792,

Panzer, 1797

Panzer, 1799

Latreille, 1799

Dahlbom, 1845

England, Wales

Latreille, 1802

André, 1886

Kirby, 1798

misident.

Jurine, 1807

Latreille, 1845

Gussakovskij, 1928

Leclercq, 1961

Curtis, 1836

misident.

(Dahlbom, 1843,

Šnoflak, 1943

Wilcke, 1945

England, Wales

(Linnaeus, 1758)

Linnaeus, 1758

(Poda, 1761,

(Geoffroy, 1785,

(Christ, 1791,

Kirby, 1798

Sowerby, 1805

(Jurine, 1807,

Dahlbom, 1845

Cameron, 1889

Gussakovskij, 1932

England, Scotland, Wales, Ireland

Fernald, 1927

Dahlbom, 1842 preocc.

Spinola, 1853 suppressed

(Kirby, 1798)

Kirby, 1798

misident.

(Mercet, 1906,

England, Wales

(Scopoli, 1763)

Scopoli, 1763

misident.

(Fabricius, 1787,

(Gmelin, 1790,

(Kirby, 1798,

England, Wales

Latreille, 1804

Engel (2005) proposed that the informal name , without formal rank, be used for the bees (here denoted as an 'infraorder' due to restrictions in the available fields). Note that 'apiformes' and the corresponding 'spheciformes') is an equivalent term proposed by Brothers (1975)

Latreille, 1802

Latreille, 1802

Fabricius, 1775

Taxonomy mostly follows Gusenleitner and Schwarz (2002).

Fabricius, 1775

Illiger, 1801

Gistel, 1850

Smith, 1847

England, Wales, Ireland

(Kirby, 1802)

Kirby, 1802

England, Scotland, Wales, Ireland, Isle of Man

Smith, 1847

England, Scotland, Wales, Ireland, Isle of Man

(Müller, 1766)

Müller, 1766

(Gmelin, 1790,

England, Scotland, Wales, Ireland

(Linnaeus, 1758)

Linnaeus, 1758

(Kirby, 1802,

England, Scotland, Wales

Zetterstedt, 1838

England, Scotland, Wales, Ireland, Isle of Man

(Scopoli, 1763)

Scopoli, 1763

(Kirby, 1802,

Smith, 1855 preocc.

England, Wales, Ireland

Perkins, 1914

Perkins, 1895, preocc.

England, Scotland, Wales

(Kirby, 1802)

Kirby, 1802

(Rossius, 1792,

(Kirby, 1802,

England, Wales

Hedicke, 1933

(Fabricius, 1775)

Fabricius, 1775

(Kirby, 1802,

England, Wales

Pérez, 1890

Imhoff, 1832

Smith, 1847

England, Wales, Isle of Man

Hedicke, 1933

(Christ, 1791)

Christ, 1791

Smith, 1849

England, Wales

Hedicke, 1933

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

England, Scotland, Wales, Ireland, Isle of Man

(Kirby, 1802)

Kirby, 1802

England, Scotland, Wales, Ireland, Isle of Man

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

England, Scotland, Wales

Smith, 1851

England

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

England

Probably extinct in Britain.

Hedicke, 1933

Lanham, 1949

LaBerge, 1964

Fabricius, 1775

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

Smith, 1847 preocc.

Smith, 1849

Smith, 1849 preocc.

England, Scotland, Wales, Ireland, Isle of Man

Nylander, 1848

England, Scotland

Pérez, 1890

(Kirby, 1802)

Kirby, 1802

Smith, 1847

England, Wales, Isle of Man

Pérez, 1890

Stephens, 1846

Smith, 1847

Smith, 1847

England, Wales

Smith, 1847

Smith, 1847

England

Panzer, 1801

misident.

(Kirby, 1802,

Illiger, 1806

misspelling

England, Wales, Ireland

is the summer brood of rosae

Perkins, 1916

misident.

Perkins, 1921

Perkins, 1921

England, Scotland, Wales, Ireland, Isle of Man

Although usually referred to as in Britain (e.g. Else et al. 2004), Schwarz et al. (1996) and Gusenleitner and Schwarz (2002) have treated as a synonym of Peréz, 1916; however, Else (in prep.), following P. Westrich’s (in lit.) interpretation of the type of , regards this is a separate species, with a junior synonym of .

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

Peréz, 1916

England, Wales

Hedicke, 1933

Smith, 1844

England

(Kirby, 1802)

Kirby, 1802

(Christ, 1791,

(Kirby, 1802,

England, Scotland, Wales, Ireland, Isle of Man

Warncke, 1968

Fabricius, 1776

(Schrank, 1781,

(Kirby, 1802,

Smith, 1849

England, Scotland, Wales, Ireland

Pérez, 1890

Hedicke, 1933

Lanham, 1949 preocc.

Lanham, 1950

(Linnaeus, 1758)

Linnaeus, 1758

England, Scotland, Wales, Ireland, Isle of Man

(Kirby, 1802)

Kirby, 1802

Smith, 1848

England, Scotland, Wales, Ireland, Isle of Man

The population on the Isles of Scilly and Channel Islands has been described as the subspecies sarnia Richards, 1979.

(Müller, 1776)

Müller, 1776

Olivier, 1789

Smith, 1847

England, Wales, Ireland

(Fabricius, 1775)

Fabricius, 1775

(Kirby, 1802,

England, Wales

Panzer, 1799

England

Probably extinct in Britain.

Ashmead, 1899

Hedicke, 1933

Perkins, 1914

Perkins, 1914

England

Perkins, 1915

England

Eversmann, 1852

Probably extinct in Britain.

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

Smith, 1855 preocc.

England, Wales, Ireland

Perkins, 1914

Perkins, 1914

England

(Kirby, 1802)

Kirby, 1802

England

Probably extinct in Britain.

Nylander, 1848

Probably extinct in Britain.

Friese, 1887

misident.

England, Wales

Pérez, 1903

misident.

Perkins, 1914

England, Scotland, Wales, Ireland, Isle of Man

Nylander, 1848

England, Scotland, Wales, Ireland

Pérez, 1890

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

England, Scotland, Wales

Schenck, 1853

Imhoff, 1868

England

Hirashima & Tadauchi, 1975

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

England, Scotland, Wales, Ireland, Isle of Man

Hedicke, 1933

Hedicke, 1933

Hedicke, 1933

Hirashima & LaBerge, 1965

Hirashima, LaBerge & Ikudomem 1884

(Kirby, 1802)

Kirby, 1802

Smith, 1847

Smith, 1847

Smith, 1847

Smith, 1847

England, Wales

Thomson, 1872

England

Although treated as a synonym of by Gusenleitner and Schwarz (2002) here nigrospina is treated as a valid species, following Baker (1994).

Fabricius, 1781

misident.

Smith, 1853

Smith, 1872

England

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

(Kirby, 1802,

England

Hedicke, 1933

Fabricius, 1781

misident.

England, Wales, Ireland

Warncke, 1968

Fabricius, 1793

Smith, 1847

England

Smith, 1847

England

Probably extinct in Britain.

Schmid-Egger, 2005

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

England, Wales

Robertson, 1902

LaBerge, 1964

(Kirby, 1802)

Kirby, 1802

Smith, 1847

England, Scotland, Wales, Ireland, Isle of Man

Pérez, 1890

Viereck, 1924

Timberlake, 1949

Schmiedeknecht, 1884

Stöckhert, 1930

England, Wales

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

England, Wales

Schenck, 1861

England

Probably extinct in Britain.

Hedicke, 1933

Alfken, 1899

England

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

Thomson, 1870

England, Wales

Smith, 1849

Christ, 1791 nom. dub.

England, Scotland, Wales

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

England, Scotland, Wales, Ireland, Isle of Man

Osytshnjuk, 1984

Nylander, 1848

misident.

England, Scotland, Wales, Ireland, Isle of Man

Robertson, 1902

(Fabricius, 1781)

Fabricius, 1781

misident.

England, Scotland, Wales, Ireland, Isle of Man

Hedicke, 1933

Panzer, 1799

(Kirby, 1802,

(Kirby, 1802,

Smith, 1849

England, Wales, Ireland

Imhoff, 1832

Nylander, 1852 preocc.

Schenck, 1853

England

Leach, 1815

Leach, 1815

Panzer, 1806

(Kirby, 1802)

Kirby, 1802

misident.

England, Wales

(Scopoli, 1763)

Scopoli, 1763

(Kirby, 1802,

England, Wales

Latreille, 1802

Latreille, 1802

Dahlbom, 1835

Latreille, 1803

Latreille, 1803

(Pallas, 1772)

Pallas, 1772

misident.

(Fabricius, 1775,

England, Wales

Patton, 1879

(Panzer, 1798)

Panzer, 1798

England, Scotland, Wales

Brooks, 1988

(Panzer, 1798)

Panzer, 1798

(Kirby, 1802,

England, Wales

Klug, 1807

Latreille, 1809

(Panzer, 1798)

Panzer, 1798

England, Ireland

Brooks, 1988

(Linnaeus, 1758)

Linnaeus, 1758

(Kirby, 1802,

(Kirby, 1802,

England

Latreille, 1802

Linnaeus, 1758

Linnaeus, 1758

Linnaeus, 1761

England, Scotland, Wales, Ireland, Isle of Man

There have been various subspecies names proposed for honey bees, with the supposedly native north west European population being . See Engel (1999) for a full taxonomy of .

Latreille, 1802

Latreille, 1802

Various subspecific names have been used for British but these are generally poorly justified and we have treated these as synonymous here. Subgeneric classification follows Williams et al. (2008).

Latreille, 1802

Vogt, 1911

Full synonymy for this subgenus is given by Williams et al. (2012), who shed some light on the confusing classification of and relative to

(Fabricius, 1775)

Fabricius, 1775

Scotland, Ireland

added by Bertsch et al. (2005)

(Linnaeus, 1761)

Linnaeus, 1761

England, Scotland, Wales, Ireland, Isle of Man

Vogt, 1911

Scotland, Ireland

(Linnaeus, 1758)

Linnaeus, 1758

(Harris, 1776,

England, Scotland, Wales, Ireland, Isle of Man

Vogt, 1911

(Kirby, 1802)

Kirby, 1802

England

Extinct in Britain since the 1940s.

Dalla Torre, 1880

(Fabricius, 1777)

Fabricius, 1777

England, Scotland, Wales

Dalla Torre, 1880

Vogt, 1911

(Linnaeus, 1761)

Linnaeus, 1761

Smith, 1846

Sladen, 1912

Stelfox, 1938

England, Scotland, Wales, Ireland, Isle of Man

Populations in Ireland have been referred to the subspecies .

(Fabricius, 1775)

Fabricius, 1775

(Harris, 1776,

(Kirby, 1802,

(Kirby, 1802,

England

English populations have been referred to the subspecies .

Dalla Torre, 1880

Vogt, 1911

(Linnaeus, 1758)

Linnaeus, 1758

England, Scotland, Wales, Ireland, Isle of Man

Lepeletier, 1832

Popov, 1931

Frison, 1927

Frison, 1927

Popov, 1931

(Kirby, 1802)

Kirby, 1802

England, Scotland, Wales, Ireland

(Seidl, 1837)

Seidl, 1837

(Pérez, 1884,

England, Scotland, Wales, Ireland, Isle of Man

(Panzer, 1801)

Panzer, 1801

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

(Richards, 1936,

England, Scotland, Wales, Ireland, Isle of Man

Populations in Western Scotland have been referred to the subspecies .

(Fabricius, 1793)

Fabricius, 1793

(Kirby, 1802,

England, Wales, Ireland

(Lepeletier, 1832)

Lepeletier, 1832

misident.

England, Scotland, Wales, Ireland

(Geoffroy, 1785)

Geoffroy, 1785

England, Scotland, Wales, Ireland

Dalla Torre, 1880

Vogt, 1911

(Linnaeus, 1758)

Linnaeus, 1758

England, Scotland, Wales

added by Goulson and Williams (2001)

(Kirby, 1802)

Kirby, 1802

misident.

(Kirby, 1802,

Vogt, 1911

Wild, 1931

Richards, 1933 preocc.

Kruseman, 1953

Rasmont, 1983

England, Scotland, Wales, Ireland, Isle of Man

The following populations have been given subspecies names: on the Hebrides; on the Isle of Man; on the Shetlands ( is a replacement name for the preccupied ).

Smith, 1849

misident.

Pittioni, 1942

England, Scotland, Wales, Ireland, Isle of Man

(Linnaeus, 1761)

Linnaeus, 1761

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

England, Scotland, Wales, Ireland, Isle of Man

Vogt, 1911

Morawitz, 1869

England, Scotland, Wales, Ireland, Isle of Man

(Linnaeus, 1758)

Linnaeus, 1758

Smith, 1844

(Kirby, 1802,

England, Wales

Extinct in Britain since the 1980s but an attempt is being made to reintroduce the species to southern England. British populations have been referred to the subspecies whereas the introduced individuals from Sweden have been ascribed to the nominate subspecies.

Dalla Torre, 1880

Krüger, 1917

Illiger, 1806

Panzer, 1806

Seidl, 1837

Smith, 1876 preocc.

Yarrow, 1978

England, Scotland, Wales, Isle of Man

(Linnaeus, 1758)

Linnaeus, 1758

misident.

misident.

Evans, 1901 preocc.

Vogt, 1909

Vogt, 1911

Stelfox, 1933

Richards, 1935

Richards, 1935

Yarrow, 1978

Baker, 1996

England, Scotland, Wales, Ireland, Isle of Man

The following populations have been given subspecies names: on the Irish mainland and southern England; on the Aran islands; on Orkney; on the Isles of Scilly; in mainland Scotland and Northern England; on the Shetland islands.

(Scopoli, 1763)

Scopoli, 1763

(Harris, 1776,

(Fabricius, 1787,

(Gmelin, 1790,

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

Stevens, 1846

White, 1851

Vogt, 1909

England, Scotland, Wales, Ireland, Isle of Man

The following populations have been given subspecies names: in southern England and Wales; in Ireland; in Scotland and Northern England.

(Panzer, 1805)

Panzer, 1805

England

There is confusion surrounding the dates of capture of the supposedly British specimens of , collected near Deal in Kent (summarised by Alford 1975).

(Müller, 1776)

Müller, 1776

(Kirby, 1802,

(Kirby, 1802,

England, Scotland, Wales, Ireland

(Linnaeus, 1761)

Linnaeus, 1761

Pérez, 1879

Vogt, 1909

England, Scotland, Wales, Ireland

Latreille, 1802

Scopoli, 1770

Scopoli, 1770

(Linnaeus, 1758)

Linnaeus, 1758

(Fabricius, 1775,

England, Wales

Pérez, 1879

misident. [?]

England

Probably extinct in Britain.

Westwood, 1840

Latreille, 1802

(Forster, 1771)

Forster, 1771

(Fabricius, 1775,

(Panzer, 1799,

England, Wales

(Scopoli, 1770)

Scopoli, 1770

England

Probably extinct in Britain.

Latreille, 1802

Roberston, 1903

Latreille, 1802

(Panzer, 1799)

Panzer, 1799

Thomson, 1870

England, Wales

(Linnaeus, 1758)

Linnaeus, 1758

misident.

Thomson, 1870

England, Wales, Scotland, Isle of Man

Latreille, 1802

Scopoli, 1763

Herrich-Schäffer, 1839

Smith, 1846

England, Ireland

Herrich-Schäffer, 1839

Stephens, 1846

England, Wales

Smith, 1844

misident.

England

Herrich-Schäffer, 1839

England

Lepeletier, 1841

England

(Linnaeus, 1767)

Linnaeus, 1767

(Kirby, 1802,

(Kirby, 1802,

England, Scotland, Wales, Ireland, Isle of Man

(Linnaeus, 1767)

Linnaeus, 1767

misident.

(Kirby, 1802,

Smith, 1876

England, Scotland, Wales

Panzer, 1798

England, Wales, Ireland

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

England, Scotland, Wales, Ireland, Isle of Man

(Kirby, 1802)

Kirby, 1802

misident. [?]

England, Wales

Panzer, 1798

England, Wales

Fabricius, 1793

Panzer, 1798

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

(Kirby, 1802,

England, Wales

(Kirby, 1802)

Kirby, 1802

misident.

(Kirby, 1802,

England, Scotland, Wales, Ireland, Isle of Man

Schenck, 1861

England

Pérez, 1884

Perkins, 1917

England, Wales

Brullé, 1832

misident.

misident.

misident.

misident.

Friese, 1921

England, Wales

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

England, Wales

(Kirby, 1802)

Kirby, 1802

Zetterstedt, 1838

Smith, 1844

England, Scotland, Wales, Ireland, Isle of Man

(Kirby, 1802)

Kirby, 1802

misident.

England, Scotland, Wales, Ireland, Isle of Man

Nylander, 1848

Smith, 1851

England, Scotland, Wales, Ireland, Isle of Man

Lepeletier, 1841

misident.

England, Scotland, Wales, Ireland

Panzer, 1799

Alfken, 1901

England, Scotland, Wales, Isle of Man

(Linnaeus, 1758)

Linnaeus, 1758

Thomson, 1872

England, Scotland, Wales, Ireland, Isle of Man

Fabricius, 1793

misident.

(Kirby, 1802,

(Kirby, 1802,

England, Scotland, Wales, Ireland, Isle of Man

Panzer, 1799

(Kirby, 1802,

(Kirby, 1802,

England

(Kirby, 1802)

Kirby, 1802

misident.

Curtis, 1832

England, Wales

Jurine, 1807

England, Wales

Fabricius, 1793

(Kirby, 1802,

(Kirby, 1802,

Smith, 1844

England, Scotland, Wales, Ireland

Latreille, 1802

Latreille, 1802

Latreille, 1802

Hirashima, Moure & Daly, 1971

(Kirby, 1802)

Kirby, 1802

England

Latreille, 1802

Latreille, 1809

Latreille, 1809

(Linnaeus, 1758)

Linnaeus, 1758

England

Lepeletier, 1841

Lepeletier, 1841

Latreille, 1802

(Linnaeus, 1761)

Linnaeus, 1761

England, Wales

The British population has been described as a separate subspecies, O’Toole, 1974, and Else, Field & O’Toole (in Edwards 1997) suggest that the British population may be specifically distinct.

Smith, 1846

England, Scotland, Wales, Ireland

Eversmann, 1852

Morawitz, 1876

England, Scotland, Ireland

(Geoffroy, 1785)

Geoffroy, 1785

England, Scotland, Wales

Verhoeff, 1944

England

Schmidt & Westrich, 1993

England

added by Cross (2002)

Smith, 1846

England, Wales

Schenck, 1853

Thomson, 1872

England, Wales, Ireland, Isle of Man

(Linnaeus, 1758)

Linnaeus, 1758

England, Scotland, Wales, Ireland, Isle of Man

Viereck, 1916

Fabricius, 1793

Popov, 1939

Curtis, 1831

Smith, 1842

England

Fabricius, 1793

Nylander, 1852

(Smith, 1872,

England, Scotland, Wales, Ireland

Alfken, 1912

Smith, 1842

England

Probably extinct in Britain

Popov, 1939

(Kirby, 1802)

Kirby, 1802

misident.

Forster, 1871

(Saunders, 1894,

England

Notton and Dathe (2008) designated as lectotype a female specimen that corresponds to the species here called , with the species generally called taking the name .

(Kirby, 1802)

Kirby, 1802

misident.

England, Wales

Popov, 1939

Nylander, 1852

(Smith, 1853,

England

Fabricius, 1804

Perkins, R.C.L., 1899

Nylander, 1852

(Smith, 1869,

England, Scotland, Wales, Ireland, Isle of Man

Nylander, 1852

England, Scotland, Wales, Ireland

The subspecies Blüthgen, 1926, is recognised from Britain.

Förster, 1871

misident.

Thomson, 1872

England, Wales

Förster, 1871

Förster, 1871

(Perkins, 1900,

England

(Panzer, 1798)

Panzer, 1798

England, Wales

Popov, 1939

Smith, 1842

England, Scotland, Wales, Ireland, Isle of Man

Thomson, 1869

Thomson, 1869

Latreille, 1804

Latreille, 1804

Blüthgen, 1931

misident.

misident.

England

Smith, 1848

England

Probably extinct in Britain.

(Christ, 1791)

Christ, 1791

Smith, 1870

Perkins, 1922

England, Scotland, Wales, Ireland, Isle of Man

Robertson, 1918

Smith, 1853

Alfken, 1907

Blüthgen, 1926

misident.

England

(Rossi, 1792)

Rossi, 1792

Smith, 1849

Probably extinct in Britain.

(Linnaeus, 1758)

Linnaeus, 1758

England, Scotland, Wales, Ireland

Curtis, 1833

Nomenclature for some species ('' s. l.) updated from Pesenko (2007) but, as Pesenko’s study was limited to the Palaearctic region, we do not follow his elevation of to a valid genus, nor have we adopted his numerous subgenera.

Robertson, 1902

(Pérez, 1903)

Pérez, 1903

England, Scotland, Wales, Ireland, Isle of Man

The Irish population has been described as the subspecies hibernicum Ebmer, 1970, with other populations referred to the subspecies scoticum Ebmer, 1970.

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

(Brullé, 1832,

(Blüthgen, 1918,

England, Scotland, Wales, Ireland, Isle of Man

(Fabricius, 1793)

Fabricius, 1793

England, Scotland, Wales, Isle of Man

(Kirby, 1802)

Kirby, 1802

England, Scotland, Wales, Isle of Man

Cockerell, 1897

(Perkins, 1895)

Perkins, 1895

England

(Schenck, 1869)

Schenck, 1869

England

(Kirby, 1802)

Kirby, 1802

(Saunders, 1910,

England, Wales, Ireland

(Kirby, 1802)

Kirby, 1802

England, Scotland, Wales, Ireland, Isle of Man

(Schenck, 1853)

Schenck, 1853

misident.

England, Wales

(Brullé, 1832)

Brullé, 1832

(Saunders, 1879,

England

(Schenck, 1853)

Schenck, 1853

(Saunders, 1879,

England, Scotland, Wales, Ireland, Isle of Man

(Morawitz, 1872)

Morawitz, 1872

England

(Zetterstedt, 1838)

Zetterstedt, 1838

(Smith, 1870,

England, Scotland, Wales, Ireland

(Alfken, 1914)

Alfken, 1914

England

(Schenck, 1870)

Schenck, 1870

(Warncke, 1986,

England

added by Hawkins (2011)

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

England, Scotland, Wales, Ireland, Isle of Man

Curtis, 1833

(Kirby, 1802)

Kirby, 1802

(Kirby, 1802,

England, Wales

(Schenck, 1853)

Schenck, 1853

(Perkins, 1913,

England, Wales

(Smith, 1848)

Smith, 1848

England, Wales

(Kirby, 1802)

Kirby, 1802

England

(Kirby, 1802)

Kirby, 1802

misident.

England

(Kirby, 1802)

Kirby, 1802

Curtis, 1833

England, Wales

Warncke, 1975

(Schrank, 1781)

Schrank, 1781

(Smith, 1853,

England, Scotland, Wales, Ireland

(Smith, 1848)

Smith, 1848

England, Wales

Ashmead, 1899

(Fabricius, 1781)

Fabricius, 1781

England, Scotland, Wales, Ireland, Isle of Man

(Scopoli, 1763)

Scopoli, 1763

(Fabricius, 1793,

(Kirby, 1802,

(Kirby, 1802,

England, Scotland, Wales, Ireland, Isle of Man

(Pérez, 1903)

Pérez, 1903

misident.

(Nylander, 1848,

(Alfken, 1905,

England, Scotland, Wales, Ireland

(Kirby, 1802)

Kirby, 1802

misident.

England, Scotland, Wales

(Kirby, 1802)

Kirby, 1802

England

Probably extinct in Britain.

(Schenck, 1869)

Schenck, 1869

misident.

England

(Kirby, 1802)

Kirby, 1802

(Smith, 1848,

England

(Schenck, 1853)

Schenck, 1853

(Schenck, 1853,

England, Wales

Latreille, 1804

Thomson, 1870

von Hagens, 1874

England, Scotland, Wales

(Linnaeus, 1767)

Linnaeus, 1767

(Kirby, 1802,

Wesmael, 1835

England, Wales, Isle of Man

von Hagens, 1882

England, Wales

(Kirby, 1802)

Kirby, 1802

von Hagens, 1882

von Hagens, 1882

England, Scotland, Wales, Ireland, Isle of Man

(Linnaeus, 1758)

Linnaeus, 1758

(Kirby, 1802,

(Kirby, 1802,

England, Scotland, Wales

von Hagens, 1882

England, Scotland, Wales, Ireland

von Hagens, 1882

England, Wales

von Hagens, 1882

von Hagens, 1882

England

(Kirby, 1802)

Kirby, 1802

Smith, 1845

England, Scotland, Wales, Ireland, Isle of Man

von Hagens, 1874

England

Smith, 1845

Thomson, 1870

England, Scotland, Wales, Ireland

Thomson, 1870

England, Wales

Thomson, 1870

England, Wales

von Hagens, 1875

misident.

misident.

England, Wales

Wesmael, 1835

England, Wales

von Hagens, 1875

England, Wales

Schenck, 1866

Lepeletier, 1841

(Nylander, 1852)

Nylander, 1852

England

Probably extinct in Britain.

Lepeletier, 1841

Schenck, 1861

England

Probably extinct in Britain.

Spinola, 1808

Spinola, 1808

England

Probably extinct in Britain.

Latreille, 1802

Latreille, 1802

Ashmead, 1899

Fabricius, 1804

(Linnaeus, 1758)

Linnaeus, 1758

England, Scotland, Wales

Represented by the subspecies nigrithorax Dalla Torre, 1877

Panzer, 1806

Nylander, 1848

England

added by Else and Spooner (1987)

(Klug, 1807)

Klug, 1807

Smith, 1843

England, Wales

(Kirby, 1802)

Kirby, 1802

England, Wales

(Kirby, 1802)

Kirby, 1802

(Panzer, 1798,

England, Scotland, Wales

Latreille, 1802

Latreille, 1809

Tkalcu, 1974

Lepeletier, 1841

Probably extinct in Britain.

Mitchell, 1973

(Kirby, 1802)

Kirby, 1802

England, Wales, Ireland

Nylander, 1848

England, Wales

Latreille, 1809

(Illiger, 1806)

Illiger, 1806

Curtis, 1831

England, Wales, Ireland

Lepeletier, 1841

Smith, 1855

England, Scotland, Wales, Ireland

(Linnaeus, 1758)

Linnaeus, 1758

England, Wales

Lepeletier & Serville, 1825

Smith, 1843

England, Wales

Latreille, 1802

Thomson, 1872

Curtis, 1828

Pérez, 1879 synonymy by

misident.

England, Wales

Latreille, 1802

(Linnaeus, 1758)

Linnaeus, 1758

England, Scotland, Wales, Ireland, Isle of Man

Thomson, 1872

England

Probably extinct in Britain.

(Kirby, 1802)

Kirby, 1802

England, Wales, Ireland

Smith, 1844

England, Scotland, Wales, Ireland, Isle of Man

Includes the subspecies hiberniae Perkins, 1925.

Friese, 1899

(Lepeletier, 1841)

Lepeletier, 1841

(Smith, 1844,

(Smith, 1844,

England

Probably extinct in Britain.

Robertson, 1903

(Kirby, 1802)

Kirby, 1802

England, Scotland, Wales

(Kirby, 1802)

Kirby, 1802

England, Wales, Ireland, Isle of Man

(Kirby, 1802)

Kirby, 1802

England, Scotland, Wales, Ireland, Isle of Man

Includes the nominate subspecies and hibernica Perkins, 1925

Newman, 1834

Latreille, 1809

(Kirby, 1802)

Kirby, 1802

England

(Linnaeus, 1758)

Linnaeus, 1758

(Linnaeus, 1767,

England, Wales

Spinola, 1808

Peréz, 1890

England

added by Else (in prep.)

(Linnaeus, 1758)

Linnaeus, 1758

England

Klug, 1807

Cresson, 1864

(Thomson, 1872)

Thomson, 1872

misident.

England, Wales

(Kirby, 1802)

Kirby, 1802

(Dufour & Perris, 1840,

England

Probably extinct in Britain.

Lepeletier & Serville, 1825

(Kirby, 1802)

Kirby, 1802

England, Wales

Panzer, 1806

Thomson, 1872

Panzer, 1799

England, Scotland, Wales, Ireland

(Linnaeus, 1758)

Linnaeus, 1758

(Linnaeus, 1761,

England, Scotland, Wales

(Fabricius, 1804)

Fabricius, 1804

(Panzer, 1798,

England

(Kirby, 1802)

Kirby, 1802

misident.

England, Wales

Schmiedeknecht, 1884

(Zetterstedt, 1838)

Zetterstedt, 1838

misident.

Scotland

Curtis, 1828

England, Scotland, Wales

Smith, 1846

England, Wales

Gerstäcker, 1869

Scotland

added by Else in Else and Spooner (1987)

(Kirby, 1802)

Kirby, 1802

Curtis, 1828

England, Wales

Tkalc, 1974

(Schrank, 1781)

Schrank, 1781

England, Wales

Panzer, 1806

(Linnaeus, 1758)

Linnaeus, 1758

(Linnaeus, 1758,

Smith, 1844

England, Scotland, Wales, Ireland

Schenck, 1860

Börner, 1919

According to Danforth et al. (2006), may be better treated as a separate family, but this requires further corroboration.

Latreille, 1802

(Fabricius, 1793)

Fabricius, 1793

(Harris, 1776,

(Panzer, 1797,

(Kirby, 1802,

England, Wales

Robertson, 1904

Panzer, 1809

Warncke, 1973

misident.

England

Schenck, 1860

Kirby, 1802

Leach, 1815

Radoszkowski, 1891

Morawitz, 1876

England

(Fabricius, 1775)

Fabricius, 1775

Kirby, 1802

England, Scotland, Wales, Isle of Man

Kirby, 1802

(Panzer, 1799)

Panzer, 1799

England, Wales

Kirby, 1802

(Nylander, 1852,

England, Wales

Some distribution records taken from Archer and Burn (2012). Fig. 3: habitus of selected British .

Haliday, 1833

Taxonomy mostly follows Gordh and Móczár (1990). Distribution data from Perkins (1976).

Haliday, 1833

Latreille, 1802

Förster, 1856

Thomson, 1862 synonymy by

Thomson, 1862

Kieffer, 1905

(Thomson, 1862)

Thomson, 1862

England

added by Burn (1997)

(Förster, 1860)

Förster, 1860

Kieffer, 1905

England, Scotland, Wales, Isle of Man

Richards, 1939

England

(Jurine, 1807)

Jurine, 1807

Haliday, 1834

Curtis, 1838

Förster, 1851

(Thomson, 1862,

(Marshall, 1874,

Kieffer, 1905

Hellén, 1920

Arle, 1929

England, Scotland, Wales

Förster, 1856

Cameron, 1883

Kieffer, 1905

Kieffer, 1914

(Förster, 1851)

Förster, 1851

(Förster, 1851,

Thomson, 1862

Westwood, 1874

Kieffer, 1905 preocc.

Kieffer, 1905

England

Kieffer, 1914

Westwood, 1833

Förster, 1850

Kirchner, 1867

Westwood, 1833

(Förster, 1850,

Kieffer, 1906

Kieffer, 1906

England, Ireland

is listed as a separate species (described from English specimens, now missing) by Gordh and Móczár (1990) but we follow Richards (1939) who believed this not to be a separate species from .

(Ashmead, 1887)

Ashmead, 1887

(Ashmead, 1887,

Girad, 1898

Duchaussoy, 1920

Hoffer, 1936

Richards, 1939

England

(Ashmead, 1893)

Ashmead, 1893

Kieffer, 1907

Brethes, 1913

Fouts, 1920

Gahan, 1931

Westwood, 1832

Motshultsky, 1863

Saussure, 1892

Kieffer, 1904 synonymy by

Kieffer, 1913

Kieffer, 1913

Kieffer, 1913

Thomson, 1862

Westwood, 1874

Westwood, 1874

Keiffer, 1906

Kieffer, 1906

England

Westwood, 1832

England

Cameron, 1888]

Supposedly described from the New Forest, the type is lost and the species has not been satisfactorily interpreted since.

Kieffer, 1905

Kieffer, 1913

Brethes, 1913

(Fabricius, 1798)

Fabricius, 1798

(Ashmead, 1901,

(Brethes, 1913)

Brethes, 1913

(Turner & Waterston, 1921,

Ashmead, 1893

see Notton et al. (2014) for synonymy and distribution

(Förster, 1860)

Förster, 1860

(Kieffer, 1906,

(Kieffer, 1906,

England

(Say, 1836)

Say, 1836

England

Added by Notton et al. (2014); probably an accidental introduction.

Kieffer, 1905

Hedqvist, 1975

(Ashmead, 1893)

Ashmead, 1893

England

Richards, 1939

(Kieffer, 1914)

Kieffer, 1914

Gahan, 1931

Kieffer, 1914

Klug, 1808

Kieffer, 1905

Kieffer, 1911

Kieffer, 1914

(Fabricius, 1805)

Fabricius, 1805

(Menozzi, 1925,

England

Kieffer, 1904

Kieffer, 1905

Cameron, 1909

Kieffer, 1910

Kieffer, 1913

Kieffer, 1913

Kieffer, 1914

Benoit, 1957

Oglobin, 1963

(Haliday, 1838)

Haliday, 1838

(Westwood, 1874,

Kieffer, 1906

Kieffer, 1906

Chitty, 1906

Kieffer, 1906

England

Latreille, 1802

Morice, 1900

Latreille, 1802

Latreille, 1802

(Linnaeus, 1761)

Linnaeus, 1761

(Panzer, 1798,

Lepeletier, 1805

Mocsáry, 1889

England, Wales

Synonymy follows Rosa et al. (2015).

Móczár, 1962

(Fabricius, 1793)

Fabricius, 1793

England

Latreille, 1802

Some distribution data from Morgan (1984).

Latreille, 1802

Linnaeus, 1761

The number of species recognised in the group has varied from author to author. Soon et al. (2014) analysed species limits and found there to be 15 described European species plus cryptic species, a result more in line with Morgan’s (Morgan 1984) recognition of multiple species than, for example, Kunz’s (Kunz 1994) much more conservative treatment.

Schenck, 1856

Tournier, 1879

England, Scotland, Wales

Valkeila, 1971

England

added by Soon et al. (2014)

Linnaeus, 1761

England

Förster, 1853

Chevrier, 1862

England

added by Morgan (1984)

(Linnaeus, 1758)

Linnaeus, 1758

England, Scotland, Wales, Ireland, Isle of Man

Wesmael, 1839

Hellén, 1919 preocc.

Linsenmaier, 1959

misident.

England

Schenck, 1856

England, Scotland, Wales, Ireland, Isle of Man

Abeille de Perrin, 1879

England, Scotland

Linsenmaier, 1951

England, Wales, Ireland

Linsenmaier, 1951

England

Shuckard, 1836

Wesmael, 1839

England, Scotland, Wales, Isle of Man

Linsenmaier, 1968

Linsenmaier, 1959 preocc.

England

Dahlbom, 1854

England

added by Soon et al. (2014)

Linsenmaier, 1959

misident.

England, Scotland, Wales, Ireland, Isle of Man

Linnaeus, 1761

Linnaeus, 1767

England, Wales

Dahlbom, 1845

(Gerstäcker, 1869)

Gerstäcker, 1869

(Thomson, 1870,

Scotland

(Harris, 1776)

Harris, 1776

(Abeille de Perrin, 1878,

England, Wales

Linsenmaier, 1951

(Shuckard, 1836)

Shuckard, 1836

England, Wales

Lichtenstein, 1876

(Linnaeus, 1758)

Linnaeus, 1758

England, Scotland, Wales

Dahlbom, 1854

Spinola, 1806

Förster, 1853

(Fabricius, 1804)

Fabricius, 1804

(Panzer, 1798,

misident.

England

Abeille de Perrin, 1878

(Latreille, 1801)

Latreille, 1801

England, Scotland, Wales, Ireland, Isle of Man

(Dahlbom, 1854)

Dahlbom, 1854

England

(Dahlbom, 1845)

Dahlbom, 1845

(Dahlbom, 1854,

England, Wales

(Rossi, 1790)

Rossi, 1790

England

Latreille, 1802

Linsenmaier, 1959

misident.

England

(Scopoli, 1763)

Scopoli, 1763

(Fabricius, 1775,

(Fabricius, 1793,

England

added by Baldock and Hawkins (2013); tentative identification but subsequently confirmed (e.g. Earwaker and Sutton 2015).

Dahlbom, 1854

misident.

England

Panzer, 1801

(Fabricius, 1787)

Fabricius, 1787

England, Wales

(Mocsáry, 1887)

Mocsáry, 1887

England, Scotland, Wales

Treated as a subspecies of aeneus in Fauna Europaea.

Abeille de Perrin, 1879

(Dahlbom, 1831)

Dahlbom, 1831

England

Ashmead, 1902

(Linnaeus, 1758)

Linnaeus, 1758

England, Scotland, Wales, Ireland

(Scopoli, 1763)

Scopoli, 1763

England, Wales

Haliday, 1833

Some distributional data and synonymy from Olmi (1984), supplemented by Perkins (1976) and Olmi (1989).

Perkins, 1912

Jurine, 1807

Haliday, 1838

Perkins, 1905

Kieffer, 1905

Kieffer, 1905

Richards, 1939

in part, Perkins, 1976

England, Scotland, Wales, Ireland

(Dalman, 1823)

Dalman, 1823

(Walker, 1837,

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1906

Kieffer, 1906

(Chitty, 1908,

Kieffer, 1914

England, Scotland, Wales, Ireland

(Dalman, 1818)

Dalman, 1818

(Dalman, 1818,

(Thomson, 1860,

Kieffer, 1905

Kieffer, 1905

(Berland, 1928,

Richards, 1939

Moczar, 1983

England, Scotland, Wales, Ireland

(Haupt, 1941)

Haupt, 1941

Hellén, 1935 nom. nud.

Jansson, 1950

Hellén, 1953

England, Wales

added by Burn (1995)

(Thomson, 1860)

Thomson, 1860

Burn, 1990

England

(Dalman, 1818)

Dalman, 1818

Kieffer, 1905

Haupt, 1941

England, Scotland, Wales, Ireland

(Haliday, 1828)

Haliday, 1828

(Thomson, 1860,

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1906

(Richards, 1935,

England, Scotland, Wales, Ireland, Isle of Man

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

(Chitty, 1908,

England, Scotland, Wales, Ireland

(Haliday, 1837)

Haliday, 1837

misident.

(Haliday, 1837,

(Thomson, 1860,

Kieffer, 1905

Kieffer, 1905

(Chitty, 1908,

England, Scotland, Ireland

Latreille, 1809

(Dalman, 1818,

(Haliday, 1828,

(Walker, 1837,

(Walker, 1837,

(Haliday, 1837,

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

(Chitty, 1908,

(Berland, 1928,

Richards, 1939

Perkins, 1976

England, Scotland, Wales, Ireland

(Dalman, 1818)

Dalman, 1818

(Dalman, 1823,

(Ljungh, 1824,

(Haliday, 1828,

(Walker, 1837,

(Walker, 1837,

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

(Chitty, 1908,

(Chitty, 1908,

(Maneval, 1935,

(Haupt, 1941,

Moczar, 1983

England, Scotland, Wales, Ireland

Included as a synonym of brachycerum by Richards (1978).

Kieffer, 1905

England

added by Olmi (1989)

(Haliday, 1837)

Haliday, 1837

Kieffer, 1905

Kieffer, 1905

(Haupt, 1941,

England

Kieffer, 1905

Kieffer, 1905

(Chitty, 1908,

(Berland, 1928,

(Maneval, 1935,

(Richards, 1936,

(Ponomarenko, 1970,

England, Scotland, Ireland

Kieffer, 1905

Kieffer, 1913

Kieffer, 1913

(Dalman, 1818)

Dalman, 1818

(Dalman, 1818,

(Dalman, 1818,

(Dalman, 1818,

(Dalman, 1823,

(Haliday, 1828,

(Walker, 1837,

(Walker, 1837,

(Walker, 1837,

(Thomson, 1860,

(Thomson, 1860,

(Förster, 1861,

(Kieffer, 1905,

(Kieffer, 1905,

(Kieffer, 1905,

(Kieffer, 1905,

(Kieffer, 1905,

(Kieffer, 1905,

(Kieffer, 1905,

(Kieffer, 1905,

(Kieffer, 1905,

(Kieffer, 1905,

(Kieffer, 1905,

(Kieffer, 1906,

(Kieffer, 1906,

(Chitty, 1908,

(Chitty, 1908,

(Chitty, 1908,

(Kieffer, 1914,

(Kieffer, 1914,

(Oglobin, 1924,

(Oglobin, 1924,

(Haupt, 1941,

(Ponomarenko, 1970,

(Richards, 1971,

(Moczar, 1983,

(Moczar, 1983,

England, Scotland, Wales, Ireland, Isle of Man

Perkins, 1912

Dalman, 1823

Benoit, 1951

Benoit, 1951

Data on five species reared in Wales from Jervis (1977).

(Dalman, 1823)

Dalman, 1823

Richards, 1939

England, Scotland, Wales, Ireland

Richards, 1939

Richards, 1939

England, Wales

(Dalman, 1818)

Dalman, 1818

(Ratzeburg, 1848,

England, Scotland, Wales, Ireland

Kieffer, 1905

England, Scotland, Wales, Ireland

Olmi, 1984

England

added by Burn (1995)

Richards, 1939

England, Scotland, Wales, Ireland

Richards, 1939

Förster, 1856

Ruthe, 1859

England

Kieffer, 1906

Latreille, 1804

Westwood, 1835

Perkins, 1905

Perkins, 1905

Kieffer, 1906

Kieffer, 1906

Perkins, 1907

(Linnaeus, 1767)

Linnaeus, 1767

Latreille, 1805

(Westwood, 1835,

(Kieffer, 1914,

England

Kieffer, 1904

(Richards, 1939,

England

Kieffer, 1906

Ljungh, 1810

Haliday, 1829-30

Haliday, 1833

Perkins, 1905

Perkins, 1905

Perkins, 1905

Perkins, 1905

Perkins, 1905

Kieffer, 1906

Perkins, 1907

Perkins, 1907

Perkins, 1907

Kieffer, 1907

Kieffer, 1913

Kieffer, 1914

Hellén, 1930

Oglobin, 1932

Haupt, 1938

Richards, 1939

Richards, 1939

Richards, 1939

Richards, 1939

Benoit, 1953

Benoit, 1953

Ponomarenko, 1966

Olmi & Currado, 1976

Perkins, 1976

Kieffer, 1905

(Richards, 1939,

England, Wales

(Haliday, 1828)

Haliday, 1828

(Haliday, 1833,

(Westwood, 1833,

Kieffer, 1905

Kieffer, 1906

Haupt, 1916

(Heikinheimo, 1957,

England, Scotland, Wales, Ireland

(Thunberg, 1827)

Thunberg, 1827

Westwood, 1833

Thomson, 1860

(Förster, 1861,

Szépligeti, 1901,

Kieffer, 1905

Kieffer, 1907

Sahlberg, 1910

Strand, 1919

Richards, 1939

Ponomarenko, 1965

Ponomarenko, 1970

England, Scotland, Wales, Ireland

Kieffer, 1905

Sahlberg, 1910

(Ceballos, 1927,

England, Wales

Kieffer, 1905

Thomson, 1860 preocc.

Kieffer, 1905

Sahlberg, 1910

Picard, 1932

(Haupt, 1938,

Hellén, 1953

Ponomarenko, 1965

England, Wales, Ireland

(Richards, 1939)

Richards, 1939

England, Wales

Raatikainen, 1961

Kieffer, 1906

Hellén, 1935

England

added by Burn (1997)

Klug, 1810

Kieffer, 1904

Kieffer, 1904

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1905

Kieffer, 1906

Strand, 1919

England, Wales

Dalman, 1818

Westwood, 1833

Sahlberg, 1910

(Ponomarenko, 1966,

England

Kieffer, 1905

(Moczar, 1965,

(Ponomarenko, 1965,

England, Scotland, Wales, Ireland

Perkins, 1905

Richards, 1939

(Westwood, 1833)

Westwood, 1833

England

Förster, 1856

Westwood, 1833

Westwood, 1833

Nees, 1834

Agassiz, 1846

De Romand, 1846

Förster, 1856

Ashmead, 1893

Brues, 1933

(Westwood, 1833)

Westwood, 1833

(Westwood, 1833,

(Nees, 1834,

(Kieffer, 1906,

(Kieffer, 1906,

(Kieffer, 1906,

Kieffer, 1906

England, Scotland, Wales

Some distribution data for , , and from Archer (2014). Figs 4, 5, 6: habitus of selected British .

Latreille, 1809

Nomenclature, synonymy and higher classification based on Bolton (1995), Bolton (2003) and AntWeb, with additional references given.

Forel, 1878

[LINEPITHEMA Mayr, 1866 # humile (Mayr, 1868, )] Although this highly invasive species has the potential to establish itself in hothouses, given the correct climate, we are unaware of any established colonies being found in Britain. Any introductions have been of isolated workers.

Förster, 1850

Dufour, 1857

Forel, 1910

Santschi, 1925

Wheeler, 1928

Kurian, 1955

(Latreille, 1798)

Latreille, 1798

(Losana, 1834,

(Nylander, 1849,

Förster, 1850

Emery, 1925

Emery, 1925

Emery, 1925

Karavaiev, 1927

England

Seifert, 2012

misident.

misident.

England

Seifert (2012) clarified the identity of the north European .

(Fabricius, 1793)

Fabricius, 1793

(Jerdon, 1851,

(Smith, 1857,

(Smith, 1860,

Santschi, 1928

Santschi, 1928

Latreille, 1802

Linnaeus, 1758

Schuckard, 1840

Wheeler, 1913

Forel, 1913

Forel, 1913

Müller, 1923

Lomnicki, 1925

Yarrow, 1955

Ruzsky, 1920 preocc.

Scotland, Ireland

Latreille, 1798

Forel, 1874

Ruzsky, 1896

Forel, 1904

Wheeler, 1913

Ruzsky, 1914

Ruzsky, 1915

Santschi, 1928

Kuznetsov-Ugamsky, 1929

Dlussky, 1967

England, Wales

Nylander, 1846

Forel, 1874

Forel, 1874

Emery, 1909

Bondroit, 1918

Scholz, 1924

Creighton, 1935

Betrem, 1954

Dlussky, 1964

England, Scotland

Linnaeus, 1758

Scopoli, 1763

Geoffroy, 1785

Razoumowski, 1789

Christ, 1791

Nylander, 1846

Wheeler, 1913

Kuznetsov-Ugamsky, 1926

Stitz, 1930

England, Scotland, Wales, Ireland, Isle of Man

Bondroit, 1917

Vashkevich, 1924

England, Scotland, Wales, Ireland, Isle of Man

Zetterstedt, 1838

Nylander, 1846

Wheeler, 1913

Bondroit, 1920

Ruzsky, 1926

Sadil, 1953

England, Scotland, Wales, Ireland

Nylander, 1846

misident.

White, 1884 preocc.

Nasonov, 1889

Ruzsky, 1915

Emery, 1925

Emery, 1925

Stärcke, 1935

England, Wales

Not a homonym under article 23.9.5 of the Code (Seifert 2004).

Retzius, 1783

Bondroit, 1912

Bondroit, 1917

Bondroit, 1918

Ruzsky, 1926

Stärcke, 1942

Gösswald, 1951

England

Probably extinct in Britain (Edwards and Broad 2005).

Linnaeus, 1761

Christ, 1791

Panzer, 1798

Nylander, 1849

Schenck, 1852

Smith, 1858

Forel, 1874

Nasonov, 1889

Bondroit, 1917

England, Wales

Fabricius, 1793

Leach, 1825

Förster, 1850

Forel, 1874

Smith, 1878

Smith, 1878

Lomnicki, 1925

England

Latreille, 1798

Nylander, 1846

Emery, 1895

Ruzsky, 1903

Ruzsky, 1905

Forel, 1909

Santschi, 1925

Santschi, 1925

Kuznetsov-Ugamsky, 1926

Kuznetsov-Ugamsky, 1926

Kuznetzsov-Ugamsky, 1926

Kuznetsov-Ugamsky, 1928

Santschi, 1928

Karavaiev, 1929

England, Scotland, Wales

Fabricius, 1804

Ruzsky, 1912

Ruzsky, 1912

Wilson, 1955

Faber, 1967

(Foerster, 1850)

Foerster, 1850

Emery, 1893

Röszler, 1942

England, Scotland, Wales, Ireland

(Latreille, 1798)

Latreille, 1798

(Latreille, 1798,

(Förster, 1850,

Forel, 1874

(Donisthorpe, 1926,

England, Wales

(Olivier, 1792)

Olivier, 1792

Forel, 1874

Forel, 1874

Forel, 1874

Zimmermann, 1935

Stärcke, 1944

England

added by Pontin (2008); further records are given by Smith and Williams (2008)

(Fabricius, 1781)

Fabricius, 1781

(Fabricius, 1804,

Emery, 1893

Ruzsky, 1902

Ruzsky, 1905

Wheeler, 1917

Wheeler, 1917

(Bondroit, 1918,

Santschi, 1925

Menozzi, 1925

Karavaiev, 1926

Cook, 1953

England, Scotland, Wales, Ireland, Isle of Man

(Latreille, 1798)

Latreille, 1798

England, Scotland, Wales, Ireland, Isle of Man

(Bondroit, 1920)

Bondroit, 1920

England, Wales

(Nylander, 1846)

Nylander, 1846

England, Scotland, Wales, Ireland

Van Loon, Boomsma & Andrasfalvy, 1990

England

added by Fox (2009)

(Linnaeus, 1758)

Linnaeus, 1758

(Christ, 1791,

(Schenck, 1852,

Forel, 1874

Ruzsky, 1905

(Kuznetzsov-Ugamsky, 1927,

(Kuznetsov-Ugamsky, 1928,

Röszler, 1943

England, Scotland, Wales, Ireland, Isle of Man

Seifert, 1991

England, Scotland, Wales, Ireland

added by Seifert (1992)

Seifert, 1992

England, Wales

added by Seifert (1992)

(Bondroit, 1918)

Bondroit, 1918

England, Wales, Isle of Man

(Nylander, 1846)

Nylander, 1846

(Walsh, 1863,

Ruzsky, 1902

Donisthorpe, 1914

(Bondroit, 1918,

Wheeler, 1928

Stärcke, 1937

Santschi, 1941

(Röszler, 1943,

Buren, 1944

England, Scotland, Wales, Ireland, Isle of Man

Motschulsky, 1863

(Latreille, 1802)

Latreille, 1802

(Jerdon, 1851,

(Nylander, 1856,

Motschoulsky, 1863

Mayr, 1861

Faber, 1969

Faber, 1969

Forel, 1895

Santschi, 1911

Santschi, 1920

Emery, 1921

Two nests persisted on the Isle of Wight from 2007-2008 (BB, pers. obs., specimens in NHM, BB coll. and B. Seifert coll.) but appear to have died out since. A native of the Azores and Madeira.

Lepeletier, 1835

The tribal classification follows Ward et al. (2015).

Smith, 1858

Westwood, 1841

(Fabricius, 1793)

Fabricius, 1793

(Forskål, 1775,

(Losana, 1834,

(Heer, 1852,

(Smith, 1855,

(Smith, 1857,

(Smith, 1858,

Smith, 1858

(Smith, 1859,

Mayr, 1862

(Gerstäcker, 1862,

Forel, 1891

Forel, 1891

Forel, 1905

Emery, 1915

A tramp species, in the past regarded as a proper introduction in Britain but no established colonies have been reported in the period 1970–present. On a world-wide scale its importance as a tramp species seems to be diminishing. The origin of is certainly Afrotropical as it is a member of a large and successful species group that is otherwise confined to that region.

Forel, 1893

Emery, 1869

Crawley, 1920 was described from a worker casually introduced in Britain; included here only because the type-locality is Britain.

Lund, 1831

(Olivier, 1792, ) is an occasional accidental import with cork from southern Europe. It has never established in Britain and, with the decrease in the cork trade, imports of this species will probably become even rarer.

Mayr, 1855

Wheeler, 1904

(Nylander, 1846)

Nylander, 1846

(Förster, 1850,

(Wasmann, 1906,

England, Scotland

Mayr, 1855

Kutter, 1945

Ruzsky, 1904

Many species previously placed in have been transferred to by Bolton (2003).

(Fabricius, 1793)

Fabricius, 1793

(Zetterstedt, 1838,

Ruzsky, 1905

Ruzsky, 1905

Ruzsky, 1920

(Kuznetsov-Ugamsky, 1928,

England, Scotland, Wales, Ireland, Isle of Man

Curtis, 1829

Mann, 1921

(Latreille, 1802)

Latreille, 1802

Curtis, 1829

(Nylander, 1849,

(Förster, 1850,

Forel, 1914

Bondroit, 1918

Karavaiev, 1930

Karavaiev, 1930

Santschi, 1939

England, Wales

Mayr, 1853

Schenck, 1853

(Schenck, 1852)

Schenck, 1852

(Schenck, 1852,

Donisthorpe, 1936

England

Mayr, 1861

Roger, 1863

Emery, 1895

Ruzsky, 1902

Wheeler, 1905

Mann, 1920

Mann, 1920

Menozzi, 1923

Menozzi, 1925

Smith, 1950

Hamann & Klemm, 1967

(Curtis, 1854)

Curtis, 1854

misident.;

(Bondroit, 1918,

England, Wales

(Schenck, 1852)

Schenck, 1852

(Nylander, 1856,

(Bondroit, 1918,

England

(Foerster, 1850)

Foerster, 1850

(Schenck, 1852,

(Forel, 1874,

(Ruzsky, 1902,

England, Wales

(Latreille, 1798)

Latreille, 1798

(Emery, 1869,

(Forel, 1874,

(Ruzsky, 1905,

(Karavaiev, 1937,

(Karavaiev, 1937,

(Arnol’di, 1977,

England

Recorded from England as an accidental introduction by Fox (2011).

Mayr, 1855

Roger, 1857

Forel, 1874

Forel, 1887

Forel, 1890

Mayr, 1901

Emery, 1912

Wheeler, 1920

Kratochvil, 1941

Kutter, 1950

(Schenck, 1852)

Schenck, 1852

(Creighton, 1934,

England

(Nylander, 1846)

Nylander, 1846

misident.

(Guérin-Méneville, 1852,

(Mayr, 1853,

(Smith, 1860,

(Smith, 1862,

(Linnaeus, 1758)

Linnaeus, 1758

(Leach, 1825,

(Nylander, 1846,

(Foerster, 1850,

Viehmeyer, 1914

Donisthorpe, 1915

Santschi, 1927

Santschi, 1927

Ruzsky, 1936

(Enzmann, 1946,

Consani & Zngheri, 1952

Wang & Xiao, 1988

England, Scotland, Wales, Ireland, Isle of Man

s.l. is now recognised as comprising several species but the species present in Britain is apparently the true , at least on the basis of specimens from southern England (Schlick-Steiner et al. 2006).

(Smith, 1851)

Smith, 1851

(Smith, 1859,

Emery, 1877

Forel, 1902

Santschi, 1910

Forel, 1913

Santschi, 1914

(Borgmeier, 1928,

Santschi, 1928

Lepeletier, 1835

Latreille, 1804

Emery, 1907

Menozzi, 1925

Arnol’di, 1930

Cole, 1957

Arnol’di, 1968

Elmes, 1978

England, Wales

(Arnol’di, 1930)

Arnol’di, 1930

misident.

(Samsinak, 1957,

van Boven, 1970

(Kutter, 1973,

England, Wales

Nylander, 1846

Curtis, 1854

Bondroit, 1911

Stärcke, 1927

Karavaiev, 1929

Karavaiev, 1929

Karavaiev, 1929

Santschi, 1931

Arnol’di, 1934

Karavaiev, 1934

Stitz, 1939

England, Scotland, Wales

Finzi, 1926

Scotland

added by Seifert (2000)

(Linnaeus, 1758)

Linnaeus, 1758

Nylander, 1846

Curtis, 1854

Forel, 1901

Finzi, 1926

Weber, 1947

Seifert, 1993

England

Nylander, 1846

Say, 1836

Nylander, 1849

Forel, 1874

Wheeler, 1928

Santschi, 1937

Weber, 1947

Brian & Brian, 1949

Brian & Brian, 1949

Sadil, 1952

England, Scotland, Wales, Ireland, Isle of Man

Meinert, 1861

Forel, 1874

England, Scotland, Wales, Ireland, Isle of Man

Nylander, 1846

Forel, 1915

Bondroit, 1920

Karavaiev, 1926

Sadil, 1952

England, Scotland, Wales, Ireland, Isle of Man

Viereck, 1903

Finzi, 1926

Arnol'di, 1934

Ruzsky, 1946

Boer & Noordijk, 2005

England, Wales, Ireland

Bondroit, 1918

Emery, 1924

Finzi, 1926

Karavaiev, 1926

(Karavaiev, 1926,

Santschi, 1931

Stärcke, 1942

Stärcke, 1942

Sadil, 1952

Sadil, 1952

Arnol’di, 1976

Radchenko & Elmes, 2010

England

Nylander, 1846

Curtis, 1854

Ruzsky, 1895

Wasmann, 1910

Donisthorpe, 1915

Forel, 1915

Ruzsky, 1920

Kuznetsov-Ugamsky, 1928

Kupyanskaya, 1990

England, Scotland, Wales

Bondroit, 1920

England, Wales

added by Elmes et al. (2003)

Forel, 1893

Mayr, 1855

(Jerdon, 1851)

Jerdon, 1851

(Linnaeus, 1758)

Linnaeus, 1758

(Fabricius, 1793,

(Shuckard, 1838,

(Jerdon, 1851,

(Smith, 1857,

(Smith, 1858,

(Smith, 1858,

Westwood, 1840

Mayr, 1855

Forel, 1915

Forel, 1918

Santschi, 1923

Creighton, 1930

Santschi, 1930

Creighton, 1930

Kusnezov, 1953

Kusnezov, 1953

Kusnezov, 1957

Kusnezov, 1957

(Latreille, 1798)

Latreille, 1798

(Nylander, 1849,

Ruzsky, 1905

Ruzsky, 1905

Emery, 1909

Santschi, 1934

Santschi, 1934

Santschi, 1934

Bernard, 1950

Bernard, 1950

Bernard, 1950

Bernard, 1950

Bernard, 1950

Bernard, 1950

Bernard, 1950

Bernard, 1950

Bernard, 1950

Bernard, 1950

Bernard, 1950

Ettershank, 1966

(Bernard, 1978,

(Bernard, 1978,

(Bernard, 1978,

(Bernard, 1978,

England

Ashmead, 1905

Westwood, 1839

Mayr, 1861

Santschi, 1921

(Förster, 1850)

Förster, 1850

(Förster, 1850,

Karavaiev, 1926

Arnol’di, 1928

Begdon, 1932

Casevitz-Weulerrse, 1990

England, Wales, Ireland

added by DuBois (1993)

Westwood, 1839

England

Records from Ireland may relate to (Fox in Collins and Roy 2012).

Lepeletier, 1835

Santschi, 1938

(Forel, 1912, ) was described from a worker casually introduced to Britain.

(Roger, 1859)

Roger, 1859

(Roger, 1859,

(Charsley, 1877,

(Forel, 1892,

(Santschi, 1911,

(Santschi, 1912,

(Forel, 1914,

(Santschi, 1914,

(Santschi, 1914,

(Forel, 1916,

(Santschi, 1923,

(Wheeler, 1927,

(Wheeler, 1931,

(Arnol’di, 1932,

(Weber, 1942,

(Bernard, 1953,

(Bernard, 1953,

(Stefani, 1970,

England, Scotland, Wales, Ireland

(Forel, 1893)

Forel, 1893

(Emery, 1899,

(Forel, 1899,

(Santschi, 1911,

(Forel, 1911,

England

Latreille, 1804

Wheeler, 1933

Wheeler, 1933

Bernard, 1950

(Latreille, 1802)

Latreille, 1802

(Latreille, 1802,

Emery, 1898

Santschi, 1921

England, Wales

Emery, 1895

Emery, 1916

England

added by Attewell et al. (2010)

Latreille, 1802

Nomenclature follows Lelej (2002), who includes full synonymy. Only names that have been used in the British literature are included here.

Latreille, 1802

Linnaeus, 1758

Linnaeus, 1758

England, Scotland

Thomson, 1870

(Fabricius, 1787)

Fabricius, 1787

England

Fox, 1894

Latreille, 1796

Panzer, 1801

(Fabricius, 1793,

England, Wales, Ireland, Isle of Man

The nominate subspecies occurs in England and Wales, atra erythrocephala Yarrow, 1954 in Ireland and the Isle of Man.

Latreille, 1805

Banks, 1910

Classification follows Wahis (1986), Wahis (2006). Some distribution data from Day (1988).

Lepeletier, 1845

Lepeletier, 1845

Spinola, 1841

Costa, 1859

Kohl, 1844

(Scopoli, 1763)

Scopoli, 1763

(Fabricius, 1804,

(Schenck, 1857,

Lepeletier, 1845

Lepeletier, 1845

Haupt, 1962

Haupt, 1962

England, Wales

Pate, 1946

Lepeletier, 1845

(Spinola, 1808)

Spinola, 1808

(Dahlbom, 1829,

(Dahlbom, 1829,

(Herrich-Schäffer, 1830,

(Zetterstedt, 1838,

(Dahlbom, 1843,

(Lepeletier, 1845,

(Lepeletier, 1845,

(Lepeletier, 1845,

(Costa, 1881,

(Verhoeff, 1892,

England, Wales

Panzer, 1806

Brullé, 1833

Wolf, 1965

Bilberg, 1820

Fabricius, 1804

Dahlbom, 1845

(Rossi, 1792)

Rossi, 1792

(Spinola, 1808,

Vander Linden, 1827,

(Dufour, 1841,

(Lepeletier, 1845,

(Lepeletier, 1845,

(Marquet, 1879,

(Dalla Torre, 1897,

Haupt, 1927

England, Wales

Fox, 1897

Sustera, 1912

(Geoffroy, 1785)

Geoffroy, 1785

(Fabricius, 1798,

(Dahlbom, 1843,

England

(Magretti, 1886)

Magretti, 1886

(Haupt, 1927,

England, Scotland, Wales

(Linnaeus, 1758)

Linnaeus, 1758

(Kohl, 1888,

(Ferton, 1897,

(Haupt, 1927,

England, Scotland, Wales, Ireland

Schiødte, 1837

Schiødte, 1837

Junco, 1946

(Shuckard, 1837)

Shuckard, 1837

Schiødte, 1837

Junco, 1946

England, Scotland, Wales

Wahis, 2006

Haupt, 1927 preocc.

Wolf, 2004 preocc.

England, Ireland

Haupt, 1927

England

(Fabricius, 1775)

Fabricius, 1775

(Scopoli, 1763,

(Harris, 1780,

Dahlbom, 1845

Haupt, 1927

Wolf, 1959

England, Scotland, Wales, Ireland, Isle of Man

Haupt, 1927

England, Wales, Ireland

(Fabricius, 1793)

Fabricius, 1793

(Dahlbom, 1829,

(Costa, 1887,

Radoszkowski, 1888

Verhoeff, 1892

(Saunders, 1896,

Sustera, 1938

Wolf, 1967

England, Wales

Dahlbom, 1845

(Zetterstedt, 1879,

Gussakovskij, 1930

Haupt, 1937

Sustera, 1938

Blüthgen, 1944

Wahis, 1979 nom. auct.

England, Scotland, Wales

(Lepeletier, 1845)

Lepeletier, 1845

Dubois, 1920

England

(Schiødte, 1837)

Schiødte, 1837

England, Wales

Haupt, 1927

England, Scotland, Wales

Junco, 1946

Haupt, 1949

Dahlbom, 1843

Junco, 1946

England

(Harris, 1780)

Harris, 1780

misident.

(Panzer, 1804,

(Dahlbom, 1829,

(Lepeletier, 1845,

(Smith, 1851,

Wolf, 1960

England, Scotland, Wales, Ireland

Haupt, 1927

Haupt, 1927

Wolf, 1975

England, Wales

Latreille, 1805

Ashmead, 1902

Ashmead, 1902

Banks, 1917

(Spinola, 1808)

Spinola, 1808

(Dahlbom, 1829,

(Dahlbom, 1832,

(Lepeletier, 1845,

England, Wales

(Vander Linden, 1827)

Vander Linden, 1827

(Lepeletier, 1845,

(Kohl, 1879,

(Tournier, 1889,

(Tournier, 1889,

(Moczar, 1944,

England

added by Baldock (2006)

Dufour, 1834

Dufour, 1834

(Aurivillius, 1907)

Aurivillius, 1907

(Perkins, 1917,

Haupt, 1937

Moczar, 1944

England, Wales

(Dahlbom, 1845)

Dahlbom, 1845

(Wesmael, 1851,

(Smith, 1877,

(Morawitz, 1891,

(Morawitz, 1891,

(Tournier, 1889,

England, Scotland, Wales, Ireland

(Scopoli, 1763)

Scopoli, 1763

(Fabricius, 1775,

(Tischbein, 1850,

(Schenck, 1857,

(Tournier, 1889,

(Tournier, 1889,

Banks, 1939

Dreisbach, 1950

England, Scotland, Wales, Ireland, Isle of Man

Howard, 1901

Ashmead, 1902

(Vander Linden, 1827)

Vander Linden, 1827

(Herrich-Schäffer, 1830,

(Shuckard, 1835,

(Dahlbom, 1843,

Haupt, 1962

(Haupt, 1962,

Wolf & Diniz, 1970

England, Wales

(Linnaeus, 1758)

Linnaeus, 1758

(Linnaeus, 1761,

(Dahlbom, 1843,

(Smith, 1879,

(Tournier, 1890,

(Tournier, 1890,

(Tournier, 1890,

(Tournier, 1890,

(Tournier, 1890,

(Dalla Torre, 1897,

(Mantero, 1905,

England, Wales, Ireland

Spinola, 1808

Spinola, 1808

Vander Linden, 1827

(Kohl, 1838,

England, Wales

Kincaid, 1900

Wilcke, 1942

Dreisbach, 1949

Wolf, 1965

Wolf, 1965

Wolf, 1965

Wolf, 1965

(Wesmael, 1851)

Wesmael, 1851

(Harris, 1870,

(Thomson, 1870,

(Tournier, 1890,

(Tournier, 1890,

(Tournier, 1890,

(Tournier, 1890,

(Tournier, 1890,

(Tournier, 1890,

(Wolf, 1966,

Wolf & Moczar, 1972

England, Scotland, Wales, Ireland, Isle of Man

(Dahlbom, 1843)

Dahlbom, 1843

(Brullé, 1840,

(Pérez, 1895,

(Dalla Torre, 1897,

(Saunders, 1904,

(Haupt, 1927,

(Haupt, 1928,

(Haupt, 1930,

(Junco, 1960,

(Wolf, 1965,

(Wolf, 1965,

(Wolf, 1966,

(Wolf, 1966,

England, Wales

(Dahlbom, 1843)

Dahlbom, 1843

misident.

(Tournier, 1889,

(Haupt, 1927,

(Blüthgen, 1961,

(Wolf, 1965,

England, Wales

(Thomson, 1870)

Thomson, 1870

England, Wales

Banks, 1939

(Dahlbom, 1842)

Dahlbom, 1842

(Dahlbom, 1843,

(Dahlbom, 1843,

(Lepeletier, 1845,

(Dalla Torre, 1897,

(Priesner, 1960,

Wolf, 1970

England, Wales

(Schiødte, 1837)

Schiødte, 1837

Wolf, 1970

England, Scotland, Wales

Kincaid, 1900

Ashmead, 1902

(Haupt, 1927)

Haupt, 1927

(Haupt, 1937,

Wolf, 1975

England

Schiødte, 1837

Ashmead, 1902

(Tournier, 1889)

Tournier, 1889

Haupt, 1930

Blüthgen, 1944

England

added by Baldock (2006)

(Linnaeus, 1758)

Linnaeus, 1758

(Rossius, 1794,

(Lepeletier, 1845,

(Wesmael, 1851,

(Costa, 1887,

(Tournier, 1889,

(Tournier, 1889,

(Tournier, 1889,

(Dalla Torre, 1897,

(Dalla Torre, 1897,

Wolf, 1961

England, Wales, Ireland

Lepeletier, 1845

Ashmead, 1902

Arnold, 1937

Pate, 1946

Haupt, 1930

Haupt, 1930

Moczar, 1946

Dreisbach, 1950

Wolf, 1970

Wolf, 1970

(Shuckard, 1835)

Shuckard, 1835

(Thomson, 1870,

Wolf, 1964

England, Scotland, Wales, Ireland, Isle of Man

(Vander Linden, 1827)

Vander Linden, 1827

Lepeletier, 1845

Costa, 1882

(Dalla Torre, 1897,

Wolf, 1970

Wolf, 1970

England

(Linnaeus, 1758)

Linnaeus, 1758

(Kohl, 1886,

(Thomson, 1870,

Wolf, 1970

England

Dahlbom, 1843

Dahlbom, 1842

Costa, 1886

(Fabricius, 1793)

Fabricius, 1793

(Latreille, 1809,

(Lepeletier, 1845,

(Eversmann, 1849,

(Marquet, 1879,

Morawitz, 1888

(Dalla Torre, 1897,

England

Fabricius, 1798

Priesner, 1955

(Fabricius, 1775)

Fabricius, 1775

(Fabricius, 1787,

Fabricius, 1798

Smith, 1879 preocc.

Tournier, 1889

Dalla Torre, 1897

Wolf & Diniz, 1970

Wolf, 1972

England, Scotland, Wales, Ireland, Isle of Man

Radoszkowski, 1888

Latreille, 1796

Schulz, 1906

Morice & Durrant, 1915

(Fabricius, 1775)

Fabricius, 1775

(Müller, 1776,

(Fourcroy, 1785,

(Panzer, 1799,

(Costa, 1882,

Costa, 1893

England, Scotland, Wales, Ireland, Isle of Man

(Fabricius, 1798)

Fabricius, 1798

Costa, 1887

England

Latreille, 1810

Pic, 1920

(Linnaeus, 1758)

Linnaeus, 1758

England, Wales

Latreille, 1796

(Fabricius, 1781)

Fabricius, 1781

England, Wales

Leach, 1915

Latreille, 1804

misspelling

Taxonomy follows Agnoli (2005).

Latreille, 1792

(Latreille, 1792,

Latreille, 1805

Latreille, 1806

(Latreille, 1809,

(Costa, 1858,

England, Wales

Fabricius, 1775

Fabricus, 1775

England, Wales

Vander Linden, 1827

England, Scotland, Wales, Ireland, Isle of Man

Laicharting, 1781

Taxonomy mostly follows van der Vecht and Fischer (1972) and Fauna Europaea (Mitroiu et al. 2004).

Wesmael, 1836

(Panzer, 1798)

Panzer, 1798

(Curtis, 1826,

England, Scotland, Wales

Thomson, 1874

(Panzer, 1799,

England

Although this species has been referred to as in British faunistic works (e.g. Richards 1980, Archer 2003, Archer 2014), the type of Panzer is lost and its identification as this species, whilst probable, is not certain (Blüthgen 1959, van der Vecht and Fischer 1972), therefore is the name most usually used for this species (e.g. in Fauna Europaea: Mitroiu et al. 2004, accessed 2016).

(Panzer, 1798)

Panzer, 1798

(Fabricius, 1793,

England, Scotland, Wales, Ireland

(Curtis, 1826)

Curtis, 1826

(Thomson, 1870,

(Thomson, 1870,

(Christ, 1791,

England, Scotland, Wales, Ireland, Isle of Man

(Wesmael, 1836)

Wesmael, 1836

(Herrich-Schäffer, 1839,

(Herrich-Schäffer, 1839,

England, Scotland, Wales, Ireland, Isle of Man

The subspecies hibernicus Blüthgen, 1937 occurs in Ireland and the Outer Hebrides.

(Linnaeus, 1758)

Linnaeus, 1758

(Christ, 1791,

(Herrich-Schäffer, 1839,

England, Scotland, Wales, Ireland, Isle of Man

(Linnaeus, 1758)

Linnaeus, 1758

England, Scotland, Wales, Ireland, Isle of Man

(Curtis, 1826)

Curtis, 1826

misident.

(Zetterstedt, 1838,

England, Scotland, Wales, Ireland, Isle of Man

(Müller, 1776)

Müller, 1776

(Zetterstedt, 1838,

(Herrich-Schäffer, 1839,

England, Scotland, Wales, Ireland, Isle of Man

Latreille, 1802

(Linnaeus, 1758)

Linnaeus, 1758

(Christ, 1791,

England, Wales

Dalla Torre, 1904

Blüthgen, 1938

(Fabricius, 1793)

Fabricius, 1793

(Thomson, 1870,

England

Blüthgen, 1938

(Shuckard, 1837)

Shuckard, 1837

England, Wales

Thomson, 1874

(Herrich-Schäffer, 1839)

Herrich-Schäffer, 1839

(Lepeletier, 1841,

England

Latreille, 1802

Latreille, 1802

(Gmelin, 1790)

Gmelin, 1790

England, Wales

(Linnaeus, 1758)

Linnaeus, 1758

(Fabricius, 1793,

(Zetterstedt, 1838,

England, Scotland, Wales, Ireland, Isle of Man

Blüthgen, 1938

(Gmelin, 1790)

Gmelin, 1790

England

Morawitz, 1867

England

Saussure, 1856

(Saussure, 1855)

Saussure, 1855

misident.

(Smith, 1857,

England

Wesmael, 1836

(Linnaeus, 1761)

Linnaeus, 1761

(Fabricius, 1793,

(Baldeni, 1894,

Richards, 1935

England, Scotland, Wales, Ireland, Isle of Man

(Curtis, 1826)

Curtis, 1826

misident.

England

(Panzer, 1798)

Panzer, 1798

England, Wales

(Brullé, 1832)

Brullé, 1832

England, Wales

Taxonomy follows Carpenter (1996).

Latreille, 1802

(Christ, 1791)

Christ, 1791

misident.

Herrich-Schäffer, 1840 nom. nud.

Herrich-Schäffer, 1841

Guérin, 1844

Erichson, 1849

Rudow, 1889

Dusmet, 1903

Buysson, 1912

Weyrauch, 1938

Weyrauch, 1939 preocc.

Giordani Soika, 1970

Gusenleitner, 1976

England

(Linnaeus, 1767)

Linnaeus, 1767

Radoszkowski, 1872

Kohl, 1898

(Weyrauch, 1938,

Ireland

Added by McClenaghan (1979) (as ), who collected a single male in Co. Down; it is unknown whether a nest had been established but this southern European species is an unlikely colonist.

Nomenclature follows Carpenter and Kojima (1997) and Kimsey and Carpenter (2012).

Rohwer, 1916

Bischoff, 1931

Blüthgen, 1943

Blüthgen, 1943

(Retzius, 1783)

Retzius, 1783

(Lepeletier, 1836,

(Herrich-Schäffer, 1841,

(Schenck, 1853,

(Schenck, 1853,

(Schenck, 1853,

(Pérez, 1910,

Paul, 1943

Ishikawa, 1969

Lee, 1986 preocc.

England, Scotland, Wales, Isle of Man

added by Falk (1982)

(Fabricius, 1781)

Fabricius, 1781

(Leach, 1814,

(Kirby, 1837,

(Zetterstedt, 1838,

(Sladen, 1918,

(Friese, 1919,

Blüthgen, 1937

England, Scotland, Wales, Ireland, Isle of Man

(Fabricius, 1793)

Fabricius, 1793

(von Schrank, 1802,

(Schenck, 1853,

Yamane, 1975

Eck, 1983

Eck, 1983

England, Wales

added by Allen and Archer (1989)

(Scopoli, 1763)

Scopoli, 1763

(Harris, 1776,

(Fabricius, 1793,

(Latreille, 1802,

(Fowler, 1833,

(Smith, 1843,

(Costa, 1858,

(De Buysson, 1905,

Lee, 1986

England, Scotland, Wales, Ireland, Isle of Man

Linnaeus, 1758

Dalla Torre, 1904

van der Vecht, 1959

Linnaeus, 1758

Harris, 1776

Retzius, 1783

Geoffroy, 1785

Christ, 1791

Smith, 1852

Radoszkowski, 1863

Gribodo, 1892 preocc.

du Buysson, 1902

Cameron, 1903

du Buysson, 1905

Pérez, 1910

Pérez, 1910

Birula, 1925

Birula, 1925

Birula, 1925 preocc.

Bequaert, 1931

Bequaert, 1931

England, Wales

British populations are considered to belong to the subspecies .

Thomson, 1869

Schmiedeknecht, 1881

Blüthgen, 1938

Blüthgen, 1943

Archer, 1982

(Panzer, 1799)

Panzer, 1799

(Smith, 1843,

(Smith, 1849,

(Schilling, 1850,

England, Scotland, Wales, Ireland

(Fabricius, 1793)

Fabricius, 1793

(Scopoli, 1763,

(Olivier, 1792,

England, Scotland, Wales, Ireland

(Linnaeus, 1758)

Linnaeus, 1758

(Radoszkowski, 1861,

(André, 1884,

Archer, 1981

Lee, 1986

Lee, 1986

England, Scotland, Wales, Ireland, Isle of Man

(Linnaeus, 1758)

Linnaeus, 1758

(Panzer, 1799,

England, Scotland, Wales, Ireland, Isle of Man

Checklist of the British and Irish aculeates

Data type: formatted text

Brief description: Word document version of the checklist

File: oo_83170.docx

Checklist of the British and Irish aculeates

Data type: spreadsheet

Brief description: Excel spreadsheet version of the checklist

File: oo_83127.xlsx