One, four or 100 genera? A new classification of the cone snails.

We present a new classification for the genus Conus sensu lato (family Conidae), based on molecular phylogenetic analyses of 329 species. This classification departs from both the traditional classification in only one genus and from a recently proposed shell- and radula-based classification scheme that separates members of this group into five families and 115 genera. Roughly 140 genus-group names are available for Recent cone snails. We propose to place all cone snails within a single family (Conidae) containing four genera-Conus, Conasprella, Profundiconus and Californiconus (with Conus alone encompassing about 85% of known species)-based on the clear separation of cone snails into four distinct and well-supported groups/lineages in molecular phylogenetic analyses. Within Conus and Conasprella, we recognize 57 and 11 subgenera, respectively, that represent well-supported subgroupings within these genera, which we interpret as evidence of intrageneric distinctiveness. We allocate the 803 Recent species of Conidae listed as valid in the World Register of Marine Species into these four genera and 71 subgenera, with an estimate of the confidence for placement of species in these taxonomic categories based on whether molecular or radula and/or shell data were used in these determinations. Our proposed classification effectively departs from previous schemes by (1) limiting the number of accepted genera, (2) retaining the majority of species within the genus Conus and (3) assigning members of these genera to species groups/subgenera to enable the effective communication of these groups, all of which we hope will encourage acceptance of this scheme.

INTRODUCTION

Cone snails are members of a hyperdiverse group of marine gastropods that are well known and appreciated for their history of rapid diversification, their tremendously diverse and complex venoms, and the elegance of their shells. They include such emblematic species as the once mythical ‘Glory-of-the-Sea’ Cone (Conus gloriamaris) and the deadly Geography Cone (Conus geographus). From 30 valid species known to Linnaeus, the current accepted number of species stands at 803 (WoRMS, 2014). These live in the tropical and warm temperate seas of the world, from the intertidal to depths of almost 1,000 m. Like most species in the superfamily Conoidea, cones are characterized by the possession of a venom gland and a highly modified radular tooth that they use as a harpoon to inject venom into their prey. Toxins included in these venoms are being studied for the possibilities they offer to understand the mechanisms underlying the evolution of species interactions, ecological diversification and the functioning of the nervous system, and also for potential therapeutic applications.

After more than two centuries of naming and overnaming, the species-level systematics of cones is undergoing a reappraisal based, among others, on molecular characters, and this is leading to a more stable taxonomy. Conversely, the supraspecific classification of the cone snails has in the last 20 years become more unstable than ever. A plethora of nominal (sub)genera reflect subtle differences in shell form and radular morphology, but these are obscuring phylogenetic relationships between terminal taxa. While toxinologists are longing for a predictive classification that can be used as a roadmap for bioprospecting (Olivera, 2006; Puillandre & Holford, 2010), the two genus-level classifications of cones proposed in the last 25 years (da Motta, 1991; Tucker & Tenorio, 2009, 2013) have remained ignored outside the world of cone-shell collectors.

Whereas the general and characteristic shape of the shell has empirically brought together all cone snails in what for two centuries has been called the family Conidae, the relationships between the species within Conidae, and in particular their eventual classification in genera and subgenera, is still debated. The first modern divisions of the family Conidae recognized one family, 14 groups “which may even represent subfamilies”, and 29 genera (Cotton, 1945: 231), two subfamilies and eight genera (Walls, 1978) and eight genera and 60 subgenera (da Motta, 1991). These classifications were based solely on shell characters, whereas Tucker & Tenorio's (2009, 2013) completely novel classification was based on shell as well as radular characters. These authors divided cone snails into five families and 89 genera, while their restricted family Conidae was itself subdivided into four subfamilies. The robustness of this classification is questionable, when radular characters have been found to be a poor phylogenetic marker in other conoidean taxa (Castelin ; Kantor & Puillandre, 2012). In fact, cone snails, or Conus sensu lato, is in itself a concept with blurred limits. Depending on the authors, some marginal species may or may not be included in cone snails. This is the case of the Cryptoconidae sensu Tucker & Tenorio (2009), treated by some authors as cone snails (e.g. Cossmann, 1896), while excluded by others (Tucker & Tenorio, 2009). DNA data indicate that Genota, a genus sometimes included in this family, may be more closely related to the family Borsoniidae (Puillandre ). Similarly, Benthofascis was excluded from the Conidae by Puillandre . Consequently, the present study will not consider Cryptoconidae and Benthofascis as members of the ingroup, but will focus on the Conidae sensu Bouchet , i.e. Conidae, Conilithidae and Taranteconidae sensu Tucker & Tenorio (2009).

In the last decade, published molecular phylogenies (listed by Puillandre ) have suggested that cone snails actually encompass very deep lineages that diverged at least 33 Ma (Duda & Kohn, 2005), the three main ones being the single species Conus californicus, sister-group to all the other cones, the ‘Small Major Clade’ of Duda & Kohn (2005) (corresponding to the family Conilithidae of Tucker & Tenorio, 2009) and the ‘Large Major Clade’ of Duda & Kohn, 2005 (corresponding to the family Conidae of Tucker & Tenorio, 2009). Furthermore, Espiritu defined 21 molecular groups within the Large Major Clade. None of these molecular phylogenies was turned into a formal classification of cone snails. However, Bandyopadhyay and a few others after him (e.g. Kraus , 2012) did apply the name Conasprella to the Small Major Clade, thus leaving the name Conus for the Large Major Clade. Several other generic or subgeneric names have also been used sporadically (e.g. Jimenez, Olivera & Teichert, 2007; Holford ; Puillandre, Watkins & Olivera, 2010). Meanwhile, new names have continued to be established to this day, with no less than 127 genus-group names available to classify cones.

Puillandre recently published a phylogeny of the cone snails based on 330 species and sequences of three mitochondrial gene regions. In the present paper we utilized this molecular phylogeny as a foundation to establish a new genus- and subgenus-level classification of the Conidae, with four genera (Californiconus, Profundiconus, Conasprella and Conus) and 71 subgenera (Fig. 1). We also tentatively allocate all cone snail species currently considered as valid in WoRMS, but not represented in the molecular phylogeny, to genera and subgenera based on their morphological characters.

Figure 1.

Simplified version of the Bayesian tree based on the concatenation of sequences of three mitochondrial gene regions (COI, 16S, 12S) and published by Puillandre : fig. 2) showing the proposed classification. Genera and subgenera with multiple species are reduced to triangles, whose lengths are proportional to the branch lenghts. Posterior probabilities (>0.95) are shown for each node. Only (sub)genera with at least one sequenced representative are figured.

MATERIAL AND METHODS

Turning the phylogeny into a classification

Proposing a classification based on a phylogeny mainly consists of (1) identifying the groups that will be named; (2) attributing available names and, if necessary, establishing new ones, for the groups identified in (1); and (3) ranking these names.

Identifying groups to be named

As far as possible, only well supported (bootstrap probability >90%; Bayesian posterior probability >0.95) clades (and single-species lineages) were linked to names. When several alternatives were possible (e.g. one supported clade that includes two supported clades, both with available names attributable to them), other characters and properties, such as shell morphology, type of prey, bathymetric and/or geographical distribution, were considered in order to identify the most appropriate grouping to minimize within-group variability. Overall, a conservative approach was adopted, to minimize the number of supraspecific taxa, both named and unnamed. In two cases (Pyruconus and Cylinder) we attributed a name to a group we recognized to be non-monophyletic (although this non-monophyly is not supported), in order to avoid having to establish new names for many small clades. Such polyphyletic, but morphologically consistent, genera may correspond to grades and in future each of the included clades may be shown to deserve its own name.

Naming of groups

As far as possible, a genus-group name (i.e. a genus or subgenus) was applied based on the position of its type species in the tree. If the type species of a nominal genus or subgenus had not been sequenced, application of the name was determined by reference to the morphologically most similar species used in the molecular analysis. If more than one name was applicable for a clade, the valid name was determined by the rule of priority. In the classification below, OD refers to the fixation of type species by original designation, SD by subsequent designation and M by monotypy (as defined by ICZN, 1999: Art. 68, 69, respectively).

Ranking of groups

Tucker & Tenorio (2009, 2013) attributed family-rank names to the main lineages identified by molecular phylogenies (i.e. the Small and Large Major Clades) and genus-rank names to groups of species that roughly correspond to the same level of resolution as the 21 groups defined by Espiritu . However, contrary to the first two steps of the process, which are based on objective criteria (e.g. monophyly of the taxa) and nomenclatural rules (e.g. position of type species), the attribution of ranks to the named clades is subjective. Taxonomists generally place two species in two different genera or two different families (or any other rank) based on a certain level of divergence, but the threshold of divergence at which a given rank is recognized varies greatly depending on the taxa, the characters involved and who is making the classification (Avise & Liu, 2011). Even for a given taxon and character (e.g. a nucleotide sequence), it is difficult to provide an objective criterion, since ranks above the species level are not defined by any biological attribute or property that could define a threshold. (For the rank of species, on the other hand, objective criteria for delimitation have been proposed, as amply developed in the literature on integrative taxonomy, e.g. De Queiroz, 1998; Samadi & Barberousse, 2006.) Accepting that ranking is to a large extent a matter of opinion, we also took into consideration prevailing usage. In the vast literature dealing with conotoxins, ranking has been very conservative, with Conidae being the only accepted family-level taxon. We have also been guided by dated molecular phylogenies of other marine gastropod clades. For example, the mean age of the most recent common ancestors of each monophyletic genus of Muricidae (Neogastropoda) is about 20 Ma, with a minimum of 9.5 Ma and a maximum of 42.2 Ma, and the age of the subfamily Rapaninae is estimated at about 61 Ma (Claremont : fig. 2). The most recent common ancestors of the bursids Tutufa and Bursa (Littorinimorpha: Tonnoidea) are estimated to be about 26 and 35 Ma, respectively (Castelin : fig. 7). The ages of the genera Littoraria and Echinolittorina (Littorinimorpha: Littorinidae) and of Turbo (Vetigastropoda: Turbinidae) are estimated to be respectively 55 Ma (Reid, Dyal & Williams, 2012), 50 Ma (Reid ) and 60–117 Ma (Williams, 2007; Williams & Duda, 2008), respectively. To place these numbers in perspective, the oldest fossils attributed with certainty to cone snails are 55 Ma (Duda & Kohn, 2005), the minimum age for the divergence between the two main clades of cones is estimated to be 33 Ma (Duda & Kohn, 2005) and the species diversification within each of these two main clades is supposed to have started between 15 and 20 Ma (Duda & Palumbi, 1999). The main lineages within cone snails are thus roughly of the same age, or even younger, than the genera as defined in other groups of caenogastropods. In an attempt to assign ranks that are roughly comparable with those used for other gastropod taxa in terms of group age, we chose to rank the main lineages of cone snails at the level of genus. With this approach, all cone snails are placed in a single family (Conidae) and most (Large Major Clade) remain in the genus Conus (consistent with prevailing usage of this name). As stated above, based on the molecular clock calibrations of Duda & Kohn (2005), the lineages that ultimately gave rise to these genera presumably separated from each other at least 33 Ma. We further identified well-supported groupings within these genera and assigned the members of these groups to subgenera. Except for the difference in assigned rank, these subgenera are essentially comparable with the generic-level groups proposed by Tucker & Tenorio (2009, 2013).

Allocation of species to genus-level groups

All the 803 species of Conidae listed as valid in WoRMS (2014) were allocated to genera and subgenera, with two levels of confidence. Those printed in bold font were placed in the classification based on the DNA sequences of Puillandre . The others were classified based on their shell and/or radula characters, following Tucker & Tenorio (2013) and, for species not included or considered as synonyms by Tucker & Tenorio (2013), following Petuch (2013) or the advice of M. Tenorio (personal communication).

Species sometimes referred to as ‘cone snails’ but allocated to Artemidiconus (Conorbidae; Tucker & Tenorio, 2009; Bouchet ), Benthofascis (Conorbidae; Tucker & Tenorio, 2009; Bouchet ), Genota (Cryptoconidae in Tucker & Tenorio, 2009; or Borsoniidae in Bouchet ) and Genotina (Cryptoconidae in Tucker & Tenorio, 2009; or Mangeliidae in Bouchet ) were excluded. Within Conorbidae, only Benthofascis lozoueti has been sequenced and molecular analysis indicates that the family is separate from Conidae (Puillandre ). However, B. lozoueti is the only conorbid species that does not resorb the inner shell walls (Tucker, Tenorio & Stahlschmidt, 2011). It thus cannot be excluded that the other Conorbidae species—which resorb them—may in fact not be confamilial. Likewise, as indicated above, molecular data place Genota in the Borsoniidae (Puillandre ) and this clade is not further discussed here. Fossil taxa (Hemiconus Cossmann, 1889; Cryptoconus Koenen, 1867; Conorbis Swainson, 1840; Conilithes Swainson, 1840; Eoconus Tucker & Tenorio, 2009 and Plagioconus Tucker & Tenorio, 2009) are not discussed either. Consequently, only the Conidae, Conilithidae and Taranteconidae (sensu Tucker & Tenorio, 2009) are discussed below, i.e. the cone snails as defined by Bouchet .

The classification is summarized as a list of the valid Recent species with their genus-level assignment (Appendix Table 1) and as a list of nominal genus-group names with their current status (Appendix Table 2).

Appendix Table 1.

List of the valid Recent species of cones and their current classification.

Species	Genus (Subgenus)
abbas Hwass in Bruguière, 1792	Conus (Cylinder)
abbreviates Reeve, 1843	Conus (Virroconus)
abrolhosensis Petuch, 1987	Conus (Dauciconus)
achatinus Gmelin, 1791	Conus (Pionoconus)
aculeiformis (Reeve, 1844)	Conasprella (Fusiconus)
acutangulus Lamarck, 1810	Conus (Turriconus)
adami Wils, 1988	Conus (Plicaustraconus)
adamsonii Broderip, 1836	Conus (Textilia)
advertex (Garrard, 1961)	Conus (Plicaustraconus)
aemulus Reeve, 1844	Conus (Lautoconus)
africanus Kiener, 1848	Conus (Lautoconus)
alabaster Reeve, 1849	Conus (Phasmoconus)
alainallaryi Bozzetti & Monnier, 2009	Conus (Dauciconus)
albellus Röckel & Korn, 1990	Conus (Eremiconus)
albuquerquei Trovão, 1978	Conus (Lautoconus)
alconnelli da Motta, 1986	Conus (Splinoconus)
alexandrei (Limpalaër & Monnier, 2012)	Conus (Phasmoconus)
alexandremonteiroi (Cossignani, 2014)	Conasprella (Ximeniconus)
alexandrinus Kaicher, 1977	Conus (Lautoconus)
algoensis G. B. Sowerby I, 1834	Conus (Sciteconus)
alisi (Moolenbeek, Röckel & Richard, 1995)	Conasprella (Boucheticonus)
allamandi (Petuch, 2013)	Conasprella (Ximeniconus)
allaryi Bozzetti, 2008	Conus (Lautoconus)
amadis Gmelin, 1791	Conus (Leptoconus)
ambiguus Reeve, 1844	Conus (Monteiroconus)
ammiralis Linnaeus, 1758	Conus (Cylinder)
amphiurgus Dall, 1889	Conus (Dauciconus)
amplus Röckel & Korn, 1992	Conus (Phasmoconus)
anabathrum Crosse, 1865	Conus (Dauciconus)
anabelae Rolán & Röckel, 2001	Conus (Lautoconus)
anaglyptica (Crosse, 1865)	Conasprella (Ximeniconus)
andamanensis E. A. Smith, 1879	Conus (Phasmoconus)
andremenezi Olivera & Biggs, 2010	Conus (Turriconus)
anemone Lamarck, 1810	Conus (Floraconus)
angasi Tryon, 1884	Conus (Plicaustraconus)
angioiorum Röckel & Moolenbeek, 1992	Conus (Phasmoconus)
anosyensis Bozzetti, 2008	Conus (Rhizoconus)
anthonyi (Petuch, 1975)	Conus (Lautoconus)
antonioi (Cossignani, 2014)	Conus (Lautoconus)
antoniomonteiroi Rolán, 1990	Conus (Lautoconus)
aphrodite (Petuch, 1979)	Conasprella (Conasprella)
aplustre Reeve, 1843	Conus (Floraconus)
arafurensis (Monnier, Limpalaër & Robin, 2013)	Conus (Pionoconus)
araneosus [Lightfoot], 1786	Conus (Conus)
arangoi Sarasúa, 1977	Conus (Dauciconus)
arawak (Petuch & R. F. Myers, 2014)	Conasprella (Ximeniconus)
archon Broderip, 1833	Conus (Stephanoconus)
arcuata (Broderip & G. B. Sowerby I, 1829)	Conasprella (Kohniconus)
ardisiaceus Kiener, 1850	Conus (Floraconus)
arenatus Hwass in Bruguière, 1792	Conus (Puncticulis)
aristophanes G. B. Sowerby II, 1857	Conus (Virroconus)
armadillo Shikama, 1971	Conus (Phasmoconus)
armiger (Crosse, 1858)	Conasprella (Dalliconus)
articulata (G.B. Sowerby II, 1873)	Conasprella (Conasprella)
artoptus G. B. Sowerby I, 1833	Conus (Virgiconus)
asiaticus da Motta, 1985	Conus (Phasmoconus)
ateralbus Kiener, 1850	Conus (Kalloconus)
athenae Filmer, 2011	Conus (Strategoconus)
atimovatae (Bozzetti, 2012)	Conus (Pionoconus)
atlanticoselvagem Afonso & Tenorio, 2004	Conus (Kalloconus)
attenuatus Reeve, 1844	Conus (Attenuiconus)
augur [Lightfoot], 1786	Conus (Strategoconus)
aulicus Linnaeus, 1758	Conus (Darioconus)
aurantius Hwass in Bruguière, 1792	Conus (Stephanoconus)
auratinus da Motta, 1982	Conus (Darioconus)
aureonimbosus Petuch, 1987	Conus (Attenuiconus)
aureopunctatus Petuch, 1987	Conus (Dauciconus)
aureus Hwass in Bruguière, 1792	Conus (Cylinder)
auricomus Hwass in Bruguière, 1792	Conus (Darioconus)
aurisiacus Linnaeus, 1758	Conus (Pionoconus)
australis Holten, 1802	Conus (Phasmoconus)
austroviola Röckel & Korn, 1992	Conus (Virgiconus)
axelrodi Walls, 1978	Conus (Strategoconus)
babaensis Rolán & Röckel, 2001	Conus (Lautoconus)
baccata (G. B. Sowerby III, 1877)	Conasprella (Ximeniconus)
baeri Röckel & Korn, 1992	Conus (Plicaustraconus)
bahamensis Vink & Röckel, 1995	Conus (Lindaconus)
baileyi (Röckel & da Motta, 1979)	Conasprella (Conasprella)
bairstowi G. B. Sowerby III, 1889	Conus (Sciteconus)
bajanensis (Nowell-Usticke, 1968)	Conasprella (Dalliconus)
balabacensis Filmer, 2012	Conus (Phasmoconus)
balteatus G. B. Sowerby I, 1833	Conus (Floraconus)
bandanus Hwass in Bruguière, 1792	Conus (Conus)
barbieri G. Raybaudi Massilia, 1995	Conus (Cylinder)
barthelemyi Bernardi, 1861	Conus (Pionoconus)
bartschi G. D. Hanna & Strong, 1949	Conus (Stephanoconus)
bayani Jousseaume, 1872	Conus (Splinoconus)
bayeri Petuch, 1987	Conus (Dauciconus)
beatrix Tenorio, Poppe & Tagaro, 2007	Conus (Turriconus)
behelokensis Lauer, 1989	Conus (Darioconus)
belairensis Pin & Leung Tack in Pin, 1989	Conus (Lautoconus)
belizeanus (Petuch & Sargent, 2011)	Conus (Dauciconus)
bellocqae van Rossum, 1996	Conus (Monteiroconus)
bellulus Rolán, 1990	Conus (Lautoconus)
bengalensis (Okutani, 1968)	Conus (Cylinder)
berdulinus Veillard, 1972	Conus (Virgiconus)
bernardinoi (Cossignani, 2014)	Conus (Lautoconus)
berschaueri (Petuch & R. F. Myers, 2014)	Conasprella (Ximeniconus)
bessei Petuch, 1992	Conus (Dauciconus)
betulinus Linnaeus, 1758	Conus (Dendroconus)
biancae Bozzetti, 2010	Conus (Darioconus)
biliosus (Röding, 1798)	Conus (Splinoconus)
binghamae Petuch, 1987	Conus (Kellyconus)
biraghii (G. Raybaudi Massilia, 1992)	Conasprella (Lilliconus)
blanfordianus Crosse, 1867	Conus (Phasmoconus)
boavistensis Rolán & Fernandes, 1990	Conus (Lautoconus)
bocagei Trovão, 1978	Conus (Lautoconus)
boeticus Reeve, 1844	Conus (Splinoconus)
boholensis (Petuch, 1979)	Conasprella (Conasprella)
bondarevi Röckel & G. Raybaudi Massilia, 1992	Conus (Splinoconus)
bonfigliolii (Bozzetti, 2010)	Conus (Plicaustraconus)
borgesi Trovão, 1979	Conus (Lautoconus)
boschorum Moolenbeek & Coomans, 1993	Conus (Pseudolilliconus)
boucheti (Richard, 1983)	Conasprella (Conasprella)
boui da Motta, 1988	Conus (Dauciconus)
boutetorum Richard & Rabiller, 2013	Conus (Pionoconus)
bozzettii (Lauer, 1991)	Conasprella (Conasprella)
brianhayesi Korn, 2001	Conus (Sciteconus)
broderipii Reeve, 1844	Conus (Phasmoconus)
bruguieri Kiener, 1846	Conus (Lautoconus)
brunneobandatus Petuch, 1992	Conus (Dauciconus)
brunneofilaris Petuch, 1990	Conus (Dauciconus)
brunneus Wood, 1828	Conus (Stephanoconus)
bruuni Powell, 1958	Conus (Afonsoconus)
bulbus Reeve, 1843	Conus (Lautoconus)
bullatus Linnaeus, 1758	Conus (Textilia)
buniatus (Bozzetti, 2013)	Conus (Strategoconus)
burryae Clench, 1942	Conus (Dauciconus)
buxeus (Röding, 1798)	Conus (Dendroconus)
byssinus (Röding, 1798)	Conus (Kalloconus)
cabraloi (Cossignani, 2014)	Conus (Lautoconus)
cacao Ferrario, 1983	Conus (Lautoconus)
cagarralensis (Cossignani, 2014)	Conus (Lautoconus)
caillaudii Kiener, 1846	Conus (Splinoconus)
cakobaui (Moolenbeek, Röckel & Bouchet, 2008)	Profundiconus
calhetae Rolán, 1990	Conus (Lautoconus)
californicus (Reeve, 1844)	Californiconus
cancellatus Hwass in Bruguière, 1792	Conus (Dauciconus)
canonicus Hwass in Bruguière, 1792	Conus (Cylinder)
capitanellus Fulton, 1938	Conus (Splinoconus)
capitaneus Linnaeus, 1758	Conus (Rhizoconus)
capreolus Röckel, 1985	Conus (Splinoconus)
caracteristicus Fischer von Waldheim, 1807	Conus (Puncticulis)
carcellesi Martins, 1945	Conus (Lamniconus)
cardinalis Hwass in Bruguière, 1792	Conus (Dauciconus)
cargilei Coltro, 2004	Conus (Dauciconus)
carioca Petuch, 1986	Conus (Sandericonus)
carnalis G. B. Sowerby III, 1879	Conus (Pseudonoduloconus)
castaneus Kiener, 1848	Conus (Dauciconus)
catus Hwass in Bruguière, 1792	Conus (Pionoconus)
cavailloni Fenaux, 1942	Conus (Phasmoconus)
caysalensis L. Raybaudi & Prati, 1994	Conus (Dauciconus)
cebuensis Wils, 1990	Conus (Phasmoconus)
cedonulli Linnaeus, 1767	Conus (Stephanoconus)
centurio (Born, 1778)	Conasprella (Kohniconus)
cepasi Trovão, 1975	Conus (Lautoconus)
ceruttii Cargile, 1997	Conus (Dauciconus)
cervus Lamarck, 1822	Conus (Textilia)
chaldaeus (Röding, 1798)	Conus (Virroconus)
chiangi (Azuma, 1972)	Conus (Stephanoconus)
chiapponorum Lorenz, 2004	Conus (Textilia)
chytreus Tryon, 1884	Conus (Lautoconus)
ciderryi da Motta, 1985	Conus (Phasmoconus)
cinereus Hwass in Bruguière, 1792	Conus (Phasmoconus)
cingulatus Lamarck, 1810	Conus (Dauciconus)
circumactus Iredale, 1929	Conus (Strategoconus)
circumcisus Born, 1778	Conus (Pionoconus)
clarus E. A. Smith, 1881	Conus (Mamiconus)
claudiae Tenorio & Afonso, 2004	Conus (Lautoconus)
clerii Reeve, 1844	Conus (Lamniconus)
cloveri Walls, 1978	Conus (Lautoconus)
cocceus Reeve, 1844	Conus (Floraconus)
coccineus Gmelin, 1791	Conus (Rubroconus)
coelinae Crosse, 1858	Conus (Virgiconus)
coffeae Gmelin, 1791	Conus (Leporiconus)
collisus Reeve, 1849	Conus (Phasmoconus)
colmani Röckel & Korn, 1990	Conus (Eremiconus)
colombi (Monnier & Limpalaër, 2012)	Conus (Dauciconus)
colombianus Petuch, 1987	Conus (Dauciconus)
coltrorum (Petuch & R. F. Myers, 2014)	Conus (Dauciconus)
comatosa (Pilsbry, 1904)	Conasprella (Fusiconus)
compressus G. B. Sowerby II, 1866	Conus (Floraconus)
condei (Afonso & Tenorio, 2014)	Conus (Lautoconus)
consors G. B. Sowerby I, 1833	Conus (Pionoconus)
conspersus Reeve, 1844	Conus (Phasmoconus)
corallinus Kiener, 1847	Conus (Splinoconus)
cordigera G. B. Sowerby II, 1866	Conus (Eugeniconus)
coriolisi (Röckel, Richard & Moolenbeek, 1995)	Conasprella (Fusiconus)
coromandelica (E. A. Smith, 1894)	Conasprella (Pseudoconorbis)
coronatus Gmelin, 1791	Conus (Virroconus)
crioulus Tenorio & Afonso, 2004	Conus (Lautoconus)
crocatus Lamarck, 1810	Conus (Darioconus)
crotchii Reeve, 1849	Conus (Lautoconus)
cumingii Reeve, 1848	Conus (Rhizoconus)
cuna Petuch, 1998	Conus (Atlanticonus)
cuneolus Reeve, 1843	Conus (Lautoconus)
curassaviensis Hwass in Bruguière, 1792	Conus (Stephanoconus)
curralensis Rolán, 1986	Conus (Lautoconus)
cuvieri Crosse, 1858	Conus (Gastridium)
cyanostoma A. Adams, 1855	Conus (Mamiconus)
cylindraceus Broderip & G. B. Sowerby I, 1830	Conus (Turriconus)
dalli Stearns, 1873	Conus (Cylinder)
damasoi (Cossignani, 2007)	Conasprella (Ximeniconus)
damottai Trovão, 1979	Conus (Lautoconus)
dampierensis Coomans & Filmer, 1985	Conus (Phasmoconus)
danilai Röckel & Korn, 1990	Conus (Splinoconus)
daphne Boivin, 1864	Conus (Phasmoconus)
darkini Röckel, Korn & Richard, 1993	Conus (Embrikena)
daucus Hwass in Bruguière, 1792	Conus (Dauciconus)
dayriti Röckel & da Motta, 1983	Conus (Splinoconus)
decoratus Röckel, Rolán & Monteiro, 1980	Conus (Lautoconus)
dedonderi (Goethaels & D. Monsecour, 2013)	Conus (Strategoconus)
delanoyae Trovão, 1979	Conus (Lautoconus)
delessertii (Récluz, 1843)	Conasprella (incertae sedis)
denizi (Afonso & Tenorio, 2011)	Conus (Lautoconus)
derrubado Rolán & Fernandes, 1990	Conus (Lautoconus)
desidiosus A. Adams, 1853	Conus (Lautoconus)
deynzerorum Petuch, 1995	Conus (Dauciconus)
diadema G. B. Sowerby I, 1834	Conus (Lividoconus)
dianthus G. B. Sowerby III, 1882	Conus (Dauciconus)
dictator (Melvill, 1898)	Conasprella (Fusiconus)
diegoi (Cossignani, 2014)	Conus (Lautoconus)
dieteri (Moolenbeek, Zandbergen & Bouchet, 2008)	Conasprella (Fusiconus)
diminutus Trovão & Rolán, 1986	Conus (Lautoconus)
dispar G. B. Sowerby I, 1833	Conus (Dauciconus)
distans Hwass in Bruguière, 1792	Conus (Fraterconus)
dominicanus Hwass in Bruguière, 1792	Conus (Stephanoconus)
dondani (Kosuge, 1981)	Profundiconus
donnae Petuch, 1998	Conus (Dauciconus)
dorotheae Monnier & Limpalaër, 2010	Conus (Lautoconus)
dorreensis Péron, 1807	Conus (Virroconus)
duffyi Petuch, 1992	Conus (Stephanoconus)
dusaveli (H. Adams, 1872)	Conus (Textilia)
ebraeus Linnaeus, 1758	Conus (Virroconus)
eburneus Hwass in Bruguière, 1792	Conus (Tesselliconus)
echinophilus (Petuch, 1975)	Conus (Lautoconus)
echo Lauer, 1989	Conus (Darioconus)
edaphus Dall, 1910	Conus (Tesselliconus)
ednae (Petuch, 2013)	Conus (Sandericonus)
edwardpauli Petuch, 1998	Conus (Dauciconus)
eldredi Morrison, 1955	Conus (Gastridium)
elegans (G. B. Sowerby III, 1895)	Conasprella (Fusiconus)
eleutheraensis (Petuch, 2013)	Conus (Dauciconus)
elokismenos (Kilburn, 1975)	Conasprella (Fusiconus)
emaciatus Reeve, 1849	Conus (Virgiconus)
emarginata (Reeve, 1844)	Conasprella (Kohniconus)
emersoni (G. D. Hanna, 1963)	Profundiconus
empressae Lorenz, 2001	Conus (Splinoconus)
encaustus Kiener, 1845	Conus (Virroconus)
episcopatus da Motta, 1982	Conus (Darioconus)
ericmonnieri (Petuch & R. F. Myers, 2014)	Conasprella (Ximeniconus)
ermineus Born, 1778	Conus (Chelyconus)
ernesti Petuch, 1990	Conus (Dauciconus)
erythraeensis Reeve, 1843	Conus (Phasmoconus)
escondidai Poppe & Tagaro, 2005	Conus (Lividoconus)
estivali Moolenbeek & Richard, 1995	Conus (Klemaeconus)
eucoronata (G.B. Sowerby III, 1903)	Conasprella (Conasprella)
eugrammata (Bartsch & Rehder, 1943)	Conasprella (Conasprella)
evansi Bondarev, 2001	Conus (Phasmoconus)
eversoni Petuch, 1987	Conus (Attenuiconus)
evorai Monteiro, Fernandes & Rolán, 1995	Conus (Lautoconus)
excelsus G. B. Sowerby III, 1908	Conus (Turriconus)
exiguus Lamarck, 1810	Conus (Phasmoconus)
eximius Reeve, 1849	Conus (Lividoconus)
explorator Vink, 1990	Conus (Dauciconus)
fantasmalis Rolán, 1990	Conus (Lautoconus)
felitae Rolán, 1990	Conus (Lautoconus)
felix Fenzan, 2012	Conus (Plicaustraconus)
fenzani (Petuch & Sargent, 2011)	Conasprella (Kohniconus)
fergusoni G. B. Sowerby II, 1873	Conus (Pyruconus)
fernandesi Tenorio, Afonso & Rolán, 2008	Conus (Lautoconus)
ferrugineus Hwass in Bruguière, 1792	Conus (Strategoconus)
fiadeiroi (Tenorio, Afonso, Cunha & Rolán, 2014)	Conus (Lautoconus)
figulinus Linnaeus, 1758	Conus (Dendroconus)
fijiensis (Moolenbeek, Röckel & Bouchet, 2008)	Conasprella (Fusiconus)
fijisulcatus Moolenbeek, Röckel & Bouchet, 2008	Conus (Asprella)
filmeri Rolán & Röckel, 2000	Conus (Lautoconus)
fischoederi Röckel & da Motta, 1983	Conus (Pionoconus)
flamingo Petuch, 1980	Conus (Dauciconus)
flammeacolor Petuch, 1992	Conus (Dauciconus)
flavescens G. B. Sowerby I, 1834	Conus (Dauciconus)
flavidus Lamarck, 1810	Conus (Virgiconus)
flavus Röckel, 1985	Conus (Phasmoconus)
flavusalbus Rolán & Röckel, 2000	Conus (Lautoconus)
floccatus G. B. Sowerby I, 1841	Conus (Pionoconus)
floridulus A. Adams & Reeve, 1848	Conus (Lividoconus)
fluviamaris (Petuch & Sargent, 2011)	Conasprella (Ximeniconus)
fontonae Rolán & Trovão, 1990	Conus (Lautoconus)
fragilissimus Petuch, 1979	Conus (Gastridium)
franciscoi Rolán & Röckel, 2000	Conus (Lautoconus)
frausseni (Tenorio & Poppe, 2004)	Profundiconus
frigidus Reeve, 1848	Conus (Virgiconus)
fulmen Reeve, 1843	Conus (Pionoconus)
fumigatus Hwass in Bruguière, 1792	Conus (Splinoconus)
furnae Rolán, 1990	Conus (Lautoconus)
furvus Reeve, 1843	Conus (Calibanus)
fuscoflavus Röckel, Rolán & Monteiro, 1980	Conus (Lautoconus)
fuscolineatus G. B. Sowerby III, 1905	Conus (Lautoconus)
gabelishi da Motta & Ninomiya, 1982	Conus (Floraconus)
garciai da Motta, 1982	Conus (Dauciconus)
garywilsoni Lorenz & Morrison, 2004	Conus (Lividoconus)
gauguini Richard & Salvat, 1973	Conus (Pionoconus)
generalis Linnaeus, 1767	Conus (Strategoconus)
genuanus Linnaeus, 1758	Conus (Kalloconus)
geographus Linnaeus, 1758	Conus (Gastridium)
gibsonsmithorum Petuch, 1986	Conus (Dauciconus)
gigasulcatus Moolenbeek, Röckel & Bouchet, 2008	Conus (Asprella)
gilberti (Bozzetti, 2012)	Conus (Splinoconus)
gilvus Reeve, 1849	Conus (Phasmoconus)
giorossii Bozzetti, 2005	Conus (Phasmoconus)
gladiator Broderip, 1833	Conus (Monteiroconus)
glans Hwass in Bruguière, 1792	Conus (Leporiconus)
glaucus Linnaeus, 1758	Conus (Dendroconus)
glenni Petuch, 1993	Conus (Atlanticonus)
glicksteini Petuch, 1987	Conus (Dauciconus)
gloriakiiensis Kuroda & Itô, 1961	Conus (Splinoconus)
gloriamaris Chemnitz, 1777	Conus (Cylinder)
glorioceanus Poppe & Tagaro, 2009	Conus (Cylinder)
goajira Petuch, 1992	Conus (Dauciconus)
gondwanensis Röckel & Moolenbeek, 1995	Conus (Turriconus)
gonsaloi (Afonso & Tenorio, 2014)	Conus (Lautoconus)
gordyi (Röckel & Bondarev, 2000)	Conasprella (Conasprella)
goudeyi (Monnier & Limpalaër, 2012)	Conus (Phasmoconus)
gradatulus Weinkauff, 1875	Conus (Sciteconus)
gradatus W. Wood, 1828	Conus (Dauciconus)
grahami Röckel, Cosel & Burnay, 1980	Conus (Lautoconus)
granarius Kiener, 1847	Conus (Stephanoconus)
grangeri G. B. Sowerby III, 1900	Conus (Asprella)
granulatus Linnaeus, 1758	Conus (Atlanticonus)
granum Röckel & Fischöder, 1985	Conus (Leporiconus)
gratacapii Pilsbry, 1904	Conus (Turriconus)
grohi (Tenorio & Poppe, 2004)	Conasprella (Endemoconus)
guanche Lauer, 1993	Conus (Lautoconus)
gubernator Hwass in Bruguière, 1792	Conus (Pionoconus)
guidopoppei (G. Raybaudi Massilia, 2005)	Conasprella (Fusiconus)
guinaicus Hwass in Bruguière, 1792	Conus (Lautoconus)
guyanensis (Van Mol, 1973)	Conasprella (Dalliconus)
habui Lan, 2002	Conus (Phasmoconus)
hamamotoi Yoshiba & Koyama, 1984	Conus (Splinoconus)
hamanni Fainzilber & Mienis, 1986	Conus (Leptoconus)
hanshassi (Lorenz & Barbier, 2012)	Conus (Strategoconus)
harasewychi Petuch, 1987	Conus (Dauciconus)
harlandi Petuch, 1987	Conus (Stephanoconus)
havanensis Aguayo & Pérez Farfante, 1947	Conus (Dauciconus)
helgae Blöcher, 1992	Conus (Asprella)
henckesi (Coltro, 2004)	Conasprella (Ximeniconus)
hennequini Petuch, 1993	Conus (Dauciconus)
henriquei (Petuch & R. F. Myers, 2014)	Conasprella (Coltroconus)
herndli (Petuch & R. F. Myers, 2014)	Conasprella (Ximeniconus)
hieroglyphus Duclos, 1833	Conus (Ductoconus)
hilli Petuch, 1990	Conus (Dauciconus)
hirasei (Kuroda, 1956)	Conus (Klemaeconus)
hivanus Moolenbeek, Zandbergen & Bouchet, 2008	Conus (Strategoconus)
honkeri Petuch, 1988	Conus (Attenuiconus)
honkerorum (Petuch & R. F. Myers, 2014)	Conus (Dauciconus)
hopwoodi (Tomlin, 1937)	Conasprella (Fusiconus)
howelli (Iredale, 1929)	Conasprella (Endemoconus)
hyaena Hwass in Bruguière, 1792	Conus (Rhizoconus)
hybridus Kiener, 1847	Conus (Lautoconus)
hypochlorus (Tomlin, 1937)	Conasprella (Fusiconus)
iansa (Petuch, 1979)	Conasprella (Coltroconus)
ichinoseana (Kuroda, 1956)	Conasprella (Fusiconus)
ignotus Cargile, 1998	Conus (Kellyconus)
ikedai (Ninomiya, 1987)	Profundiconus
immelmani Korn, 1998	Conus (Leptoconus)
imperialis Linnaeus, 1758	Conus (Stephanoconus)
inconstans E. A. Smith, 1877	Conus (Dauciconus)
infinitus Rolán, 1990	Conus (Lautoconus)
infrenatus Reeve, 1848	Conus (Sciteconus)
inscriptus Reeve, 1843	Conus (Phasmoconus)
insculpta (Kiener, 1847)	Conasprella (Fusiconus)
iodostoma Reeve, 1843	Conus (Asprella)
ione (Fulton, 1938)	Conasprella (Endemoconus)
irregularis G. B. Sowerby II, 1858	Conus (Lautoconus)
isabelarum Tenorio & Afonso, 2004	Conus (Lautoconus)
jacarusoi Petuch, 1998	Conus (Dauciconus)
janowskyae (Tucker & Tenorio, 2011)	Conasprella (Kohniconus)
janus Hwass in Bruguière, 1792	Conus (Phasmoconus)
jaspidea (Gmelin, 1791)	Conasprella (Ximeniconus)
jeanmartini (G. Raybaudi Massilia, 1992)	Profundiconus
jickelii Weinkauff, 1873	Conus (Phasmoconus)
joliveti (Moolenbeek, Röckel & Bouchet, 2008)	Conasprella (Fusiconus)
jorioi (Petuch, 2013)	Conus (Dauciconus)
josephinae Rolán, 1980	Conus (Lautoconus)
joserochoi (Cossignani, 2014)	Conus (Lautoconus)
jourdani da Motta, 1984	Conus (Lautoconus)
jucundus G. B. Sowerby III, 1887	Conus (Dauciconus)
judaeus Bergh, 1895	Conus (Virroconus)
julieandreae Cargile, 1995	Conus (Stephanoconus)
julii Lienard, 1870	Conus (Textilia)
kaiserae (Tenorio, Tucker & Chaney, 2012)	Conus (Dauciconus)
kalafuti da Motta, 1987	Conus (Dauciconus)
kanakinus (Richard, 1983)	Profundiconus
kawamurai Habe, 1962	Conus (Leptoconus)
kermadecensis Iredale, 1913	Conus (Lividoconus)
kersteni Tenorio, Afonso & Rolán, 2008	Conus (Lautoconus)
kerstitchiWalls, 1978	Conus (Dauciconus)
kevani Petuch, 1987	Conus (Dauciconus)
kiicumulus Azuma, 1982	Conus (Phasmoconus)
kimioi (Habe, 1965)	Conasprella (Boucheticonus)
kinoshitai Kuroda, 1956	Conus (Afonsoconus)
kintoki Habe & Kosuge, 1970	Conus (Virgiconus)
kirkandersi Petuch, 1987	Conus (Dauciconus)
klemae (Cotton, 1953)	Conus (Klemaeconus)
kohni (McLean & Nybakken, 1979)	Conasprella (Kohniconus)
korni G. Raybaudi Massilia, 1993	Conus (Pseudolilliconus)
kostini Filmer, Monteiro, Lorenz & Verdasca, 2012	Conus (Embrikena)
koukae (Monnier, Limpalaër & Robin, 2013)	Conus (Pionoconus)
kremerorum Petuch, 1988	Conus (Dauciconus)
kuiperi Moolenbeek, 2006	Conus (Pseudolilliconus)
kulkulcan Petuch, 1980	Conus (Dauciconus)
kuroharai (Habe, 1965)	Conus (Phasmoconus)
lamberti Souverbie, 1877	Conus (Darioconus)
lani (Crandall, 1979)	Profundiconus
largilliertii Kiener, 1847	Conus (Dauciconus)
laterculatus G. B. Sowerby II, 1870	Conus (Phasmoconus)
laueri (Monnier & Limpalaër, 2013)	Conus (Darioconus)
lecourtorum (Lorenz, 2011)	Conus (Virroconus)
leekremeri Petuch, 1987	Conus (Dauciconus)
legatus Lamarck, 1810	Conus (Cylinder)
lemniscatus Reeve, 1849	Conus (Lamniconus)
lenavati da Motta & Röckel, 1982	Conus (Splinoconus)
lenhilli (Cargile, 1998)	Conasprella (Dalliconus)
lentiginosa (Reeve, 1844)	Conasprella (Fusiconus)
leobottonii Lorenz, 2006	Conus (Pionoconus)
leobrerai da Motta & Martin, 1982	Conus (Phasmoconus)
leopardus (Röding, 1798)	Conus (Lithoconus)
levis (Bozzetti, 2012)	Conus (Pseudolilliconus)
levistimpsoni (Tucker, 2013)	Conus (Dauciconus)
leviteni (Tucker, Tenorio & Chaney, 2011)	Conus (Darioconus)
lienardi Bernardi & Crosse, 1861	Conus (Phasmoconus)
lightbourni Petuch, 1986	Conus (Bermudaconus)
limpusi Röckel & Korn, 1990	Conus (Eremiconus)
lindae Petuch, 1987	Conus (Lindaconus)
lineopunctatus Kaicher, 1977	Conus (Lautoconus)
lischkeanus Weinkauff, 1875	Conus (Lividoconus)
litoglyphus Hwass in Bruguière, 1792	Conus (Strategoconus)
litteratus Linnaeus, 1758	Conus (Elisaconus)
lividus Hwass in Bruguière, 1792	Conus (Lividoconus)
lizardensis Crosse, 1865	Conus (Eremiconus)
lizarum (Raybaudi Massilia & da Motta, 1992)	Conasprella (Fusiconus)
lobitensis Kaicher, 1977	Conus (Lautoconus)
locumtenens Blumenbach, 1791	Conus (Leptoconus)
lohri Kilburn, 1972	Conus (Darioconus)
longilineus Röckel, Rolán & Monteiro, 1980	Conus (Lautoconus)
longurionis (Kiener, 1847)	Conasprella (Fusiconus)
lorenzi Monnier & Limpalaër, 2012	Conasprella (Conasprella)
loyaltiensis (Röckel & Moolenbeek, 1995)	Profundiconus
lozeti Richard, 1980	Conus (Plicaustraconus)
lucasi (Bozzetti, 2010)	Conus (Textilia)
lucaya Petuch, 2000	Conus (Dauciconus)
luciae Moolenbeek, 1986	Conus (Turriconus)
lucidus (W. Wood, 1828)	Conasprella (Ximeniconus)
lugubris Reeve, 1849	Conus (Lautoconus)
luquei Rolán & Trovão, 1990	Conus (Lautoconus)
luteus G. B. Sowerby I, 1833	Conus (Leporiconus)
lynceus G. B. Sowerby II, 1858	Conus (Phasmoconus)
maculiferus G. B. Sowerby I, 1833	Conus (Dauciconus)
madagascariensis G. B. Sowerby II, 1858	Conus (Darioconus)
madecassinus (Bozzetti, 2012)	Conus (Phasmoconus)
magellanicus Hwass in Bruguière, 1792	Conus (Dauciconus)
magnificus Reeve, 1843	Conus (Darioconus)
magnottei Petuch, 1987	Conus (Dauciconus)
magus Linnaeus, 1758	Conus (Pionoconus)
mahogani (Reeve, 1843)	Conasprella (Ximeniconus)
maioensis Trovão, Rolán & Félix-Alves, 1990	Conus (Lautoconus)
malacanus Hwass in Bruguière, 1792	Conus (Splinoconus)
maldivus Hwass in Bruguière, 1792	Conus (Strategoconus)
mappa [Lightfoot], 1786	Conus (Stephanoconus)
marchionatus Hinds, 1843	Conus (Eugeniconus)
marielae Rehder & Wilson, 1975	Conus (Phasmoconus)
marmoreus Linnaeus, 1758	Conus (Conus)
martensi E. A. Smith, 1884	Conus (Splinoconus)
martinianus Reeve, 1844	Conus (Phasmoconus)
maya (Petuch & Sargent, 2011)	Conus (Dauciconus)
mazei (Deshayes, 1874)	Conasprella (Dalliconus)
mcbridei Lorenz, 2005	Conus (Harmoniconus)
mcgintyi (Pilsbry, 1955)	Conasprella (Dalliconus)
medoci Lorenz, 2004	Conus (Dendroconus)
melissae Tenorio, Afonso & Rolán, 2008	Conus (Lautoconus)
melvilli G. B. Sowerby III, 1879	Conus (Quasiconus)
memiae (Habe & Kosuge, 1970)	Conasprella (Conasprella)
mercator Linnaeus, 1758	Conus (Lautoconus)
messiasi Rolán & Fernandes, 1990	Conus (Lautoconus)
micropunctatus Rolán & Röckel, 2000	Conus (Lautoconus)
miles Linnaeus, 1758	Conus (Rhizoconus)
milesi E. A. Smith, 1887	Conus (Turriconus)
miliaris Hwass in Bruguière, 1792	Conus (Virroconus)
milneedwardsi Jousseaume, 1894	Conus (Leptoconus)
mindana (Hwass in Bruguière, 1792)	Conasprella (Ximeniconus)
miniexcelsus Olivera & Biggs, 2010	Conus (Turriconus)
minnamurra (Garrard, 1961)	Conus (Eremiconus)
miruchae Röckel, Rolán & Monteiro, 1980	Conus (Lautoconus)
mitratus Hwass in Bruguière, 1792	Conus (Turriconus)
moluccensis Küster, 1838	Conus (Phasmoconus)
monachus Linnaeus, 1758	Conus (Pionoconus)
moncuri Filmer, 2005	Conus (Elisaconus)
monile Hwass in Bruguière, 1792	Conus (Strategoconus)
monilifer Broderip, 1833	Conus (Dauciconus)
moolenbeeki Filmer, 2011	Conus (Phasmoconus)
mordeirae Rolán & Trovão, 1990	Conus (Lautoconus)
moreleti Crosse, 1858	Conus (Virgiconus)
morrisoni G. Raybaudi Massilia, 1991	Conus (Pionoconus)
mozambicus Hwass in Bruguière, 1792	Conus (Floraconus)
mucronatus Reeve, 1843	Conus (Phasmoconus)
mulderi Fulton, 1936	Conus (Phasmoconus)
muriculatus G. B. Sowerby I, 1833	Conus (Lividoconus)
mus Hwass in Bruguière, 1792	Conus (Monteiroconus)
musicus Hwass in Bruguière, 1792	Conus (Harmoniconus)
mustelinus Hwass in Bruguière, 1792	Conus (Rhizoconus)
namocanus Hwass in Bruguière, 1792	Conus (Rhizoconus)
nanus G. B. Sowerby I, 1833	Conus (Harmoniconus)
naranjus Trovão, 1975	Conus (Lautoconus)
natalaurantius (S. G. Veldsman, 2013)	Conus (Darioconus)
natalis G. B. Sowerby II, 1858	Conus (Leptoconus)
navarroi Rolán, 1986	Conus (Lautoconus)
negroides Kaicher, 1977	Conus (Lautoconus)
neptunus Reeve, 1843	Conus (Phasmoconus)
niederhoeferi (Monnier, Limpalaër & Lorenz, 2012)	Conus (Phasmoconus)
nielsenae Marsh, 1962	Conus (Splinoconus)
nigromaculatus Röckel & Moolenbeek, 1992	Conus (Pionoconus)
nigropunctatus G. B. Sowerby II, 1858	Conus (Pionoconus)
nimbosus Hwass in Bruguière, 1792	Conus (Phasmoconus)
nobilis Linnaeus, 1758	Conus (Eugeniconus)
nobrei Trovão, 1975	Conus (Lautoconus)
nocturnus [Lightfoot], 1786	Conus (Conus)
nodulosus G. B. Sowerby II, 1864	Conus (Cylinder)
nucleus Reeve, 1848	Conus (Splinoconus)
nussatella Linnaeus, 1758	Conus (Hermes)
nux Broderip, 1833	Conus (Harmoniconus)
nybakkeni (Tenorio, Tucker & Chaney, 2012)	Conus (Dauciconus)
obscurus G. B. Sowerby I, 1833	Conus (Gastridium)
ochroleucus Gmelin, 1791	Conus (Phasmoconus)
ogum (Petuch & R. F. Myers, 2014)	Conasprella (Ximeniconus)
oishii (Shikama, 1977)	Conus (Phasmoconus)
olgiatii Bozzetti, 2007	Conus (Splinoconus)
omaria Hwass in Bruguière, 1792	Conus (Darioconus)
orbignyi (Audouin, 1831)	Conasprella (Fusiconus)
orion Broderip, 1833	Conus (Dauciconus)
ortneri Petuch, 1998	Conus (Dauciconus)
ostrinus (Tucker & Tenorio, 2011)	Conus (Dauciconus)
otohimeae (Kuroda & Itô, 1961)	Conasprella (Conasprella)
pacei (Petuch, 1987)	Conasprella (Dalliconus)
pacificus (Moolenbeek & Röckel, 1996)	Profundiconus
pagoda (Kiener, 1847)	Conasprella (Conasprella)
papilliferus G. B. Sowerby I, 1834	Conus (Floraconus)
papuensis Coomans & Moolenbeek, 1982	Conus (Splinoconus)
paraguana Petuch, 1987	Conus (Dauciconus)
parascalaris Petuch, 1987	Conus (Dauciconus)
parius Reeve, 1844	Conus (Phasmoconus)
parvatus Walls, 1979	Conus (Harmoniconus)
paschalli Petuch, 1998	Conus (Dauciconus)
patae Abbott, 1971	Conus (Kellyconus)
patglicksteinae Petuch, 1987	Conus (Dauciconus)
patriceae (Petuch & R. F. Myers, 2014)	Conus (Lamniconus)
patricius Hinds, 1843	Conus (Pyruconus)
paukstisi (Tucker, Tenorio & Chaney, 2011)	Conus (Harmoniconus)
paulae Petuch, 1988	Conus (Dauciconus)
pauperculus G. B. Sowerby I, 1834	Conus (Splinoconus)
peli Moolenbeek, 1996	Conus (Rhizoconus)
penchaszadehi Petuch, 1986	Conus (Dauciconus)
pennaceus Born, 1778	Conus (Darioconus)
pepeiu (Moolenbeek, Zandbergen & Bouchet, 2008)	Conasprella (Fusiconus)
pergrandis (Iredale, 1937)	Conus (Embrikena)
perplexa (G. B. Sowerby II, 1857)	Conasprella (Ximeniconus)
pertusus Hwass in Bruguière, 1792	Conus (Rhizoconus)
petergabrieli Lorenz, 2006	Conus (Phasmoconus)
philippii Kiener, 1847	Conus (Dauciconus)
pica A. Adams & Reeve, 1848	Conus (Phasmoconus)
pictus Reeve, 1843	Conus (Sciteconus)
pineaui Pin & Leung Tack, 1989	Conus (Lautoconus)
planorbis Born, 1778	Conus (Strategoconus)
plinthis Richard & Moolenbeek, 1988	Conus (Klemaeconus)
polongimarumai Kosuge, 1980	Conus (Stephanoconus)
poormani Berry, 1968	Conus (Dauciconus)
poremskii (Petuch & R. F. Myers, 2014)	Conasprella (Ximeniconus)
portobeloensis Petuch, 1990	Conus (Dauciconus)
poulosi Petuch, 1993	Conus (Attenuiconus)
praecellens A. Adams, 1855	Conus (Turriconus)
pretiosus G. Nevill & H. Nevill, 1874	Conus (Phasmoconus)
primus Röckel & Korn, 1990	Conus (Phasmoconus)
princeps Linnaeus, 1758	Conus (Ductoconus)
priscai (Bozzetti, 2012)	Conus (Cylinder)
profundorum (Kuroda, 1956)	Profundiconus
proximus G. B. Sowerby II, 1860	Conus (Phasmoconus)
pseudaurantius Vink & Cosel, 1985	Conus (Stephanoconus)
pseudimperialis Moolenbeek, Zandbergen & Bouchet, 2008	Conus (Stephanoconus)
pseudocardinalis Coltro, 2004	Conus (Dauciconus)
pseudocuneolus Röckel, Rolán & Monteiro, 1980	Conus (Lautoconus)
pseudokimioi da Motta & Martin, 1982	Conasprella (Boucheticonus)
pseudonivifer Monteiro, Tenorio & Poppe, 2004	Conus (Kalloconus)
pseudorbignyi (Röckel & Lan, 1981)	Conasprella (Fusiconus)
pulcher [Lightfoot], 1786	Conus (Kalloconus)
pulcherrima Brazier, 1896	INCERTAE SEDIS (Kenyonia)
pulicarius Hwass in Bruguière, 1792	Conus (Puncticulis)
puncticulata (Hwass in Bruguière, 1792)	Conasprella (Ximeniconus)
purissimus Filmer, 2011	Conus (Splinoconus)
purpurascens G. B. Sowerby I, 1833	Conus (Chelyconus)
pusio (Hwass in Bruguière, 1792)	Conasprella (Ximeniconus)
queenslandis da Motta, 1984	Conus (Splinoconus)
quercinus [Lightfoot], 1786	Conus (Lividoconus)
quiquandoni Lorenz & Barbier, 2008	Conus (Phasmoconus)
rachelae (Petuch, 1988)	Conasprella (Kohniconus)
radiatus Gmelin, 1791	Conus (Phasmoconus)
rainesae (McGinty, 1953)	Conasprella (Dalliconus)
ranonganus da Motta, 1978	Conus (Phasmoconus)
raoulensis (Powell, 1958)	Conasprella (Endemoconus)
rattus Hwass in Bruguière, 1792	Conus (Rhizoconus)
raulsilvai Rolán, Monteiro & Fernandes, 1998	Conus (Lautoconus)
rawaiensis da Motta, 1978	Conus (Rhizoconus)
recluzianus Bernardi, 1853	Conus (Splinoconus)
recurvus Broderip, 1833	Conus (Dauciconus)
reductaspiralis Walls, 1979	Conus (Splinoconus)
regius Gmelin, 1791	Conus (Stephanoconus)
regonae Rolán & Trovão, 1990	Conus (Lautoconus)
regularis G. B. Sowerby I, 1833	Conus (Dauciconus)
retifer Menke, 1829	Conus (Cylinder)
richardbinghami Petuch, 1993	Conus (Dauciconus)
richardsae Röckel & Korn, 1992	Conus (Splinoconus)
richeri Richard & Moolenbeek, 1988	Conus (Phasmoconus)
ritae Petuch, 1995	Conus (Atlanticonus)
rizali Olivera & Biggs, 2010	Conus (Turriconus)
roatanensis (Petuch & Sargent, 2011)	Conasprella (Ximeniconus)
roberti (Richard, 2009)	Conasprella (Dalliconus)
robini (Limpalaër & Monnier, 2012)	Conus (Pionoconus)
roeckeli Rolán, 1980	Conus (Lautoconus)
rolani Röckel, 1986	Conus (Asprella)
rosalindensis Petuch, 1998	Conus (Dauciconus)
rosemaryae Petuch, 1990	Conus (Dauciconus)
roseorapum G. Raybaudi & da Motta, 1990	Conus (Splinoconus)
rouxi (Monnier, Limpalaër & Robin, 2013)	Conus (Pionoconus)
royaikeni (S. G. Veldsman, 2010)	Conus (Leptoconus)
rufimaculosus Macpherson, 1959	Conus (Klemaeconus)
ruthae (Monnier & Limpalaër, 2013)	Conus (Splinoconus)
rutila (Menke, 1843)	Conasprella (Parviconus)
saecularis (Melvill, 1898)	Conasprella (Fusiconus)
sagei (Korn & G. Raybaudi Massilia, 1993)	Conasprella (Lilliconus)
sahlbergi da Motta & Harland, 1986	Conus (Dauciconus)
salletae (Cossignani, 2014)	Conus (Lautoconus)
salreiensis Rolán, 1980	Conus (Lautoconus)
salzmanni G. Raybaudi Massilia & Rolán, 1997	Conus (Phasmoconus)
sanderi Wils & Moolenbeek, 1979	Conus (Sandericonus)
sandwichensis Walls, 1978	Conus (Tesselliconus)
sanguineus Kiener, 1850	Conus (Stephanoconus)
sanguinolentus Quoy & Gaimard, 1834	Conus (Lividoconus)
santanaensis (Afonso & Tenorio, 2014)	Conus (Lautoconus)
santinii (Monnier & Limpalaër, 2014)	Conus (Phasmoconus)
saragasae Rolán, 1986	Conus (Lautoconus)
sargenti (Petuch, 2013)	Conasprella (Ximeniconus)
sartii Korn, Niederhöfer & Blöcher, 2002	Conus (Phasmoconus)
sauros (Garcia, 2006)	Conasprella (Dalliconus)
sazanka Shikama, 1970	Conus (Splinoconus)
scalaris Valenciennes, 1832	Conus (Dauciconus)
scalarispira (Bozzetti, 2012)	Conus (Pseudolilliconus)
scalarissimus da Motta, 1988	Conus (Dauciconus)
scalptus Reeve, 1843	Conus (Phasmoconus)
scaripha (Dall, 1910)	Conasprella (Kohniconus)
scopulicola (Okutani, 1972)	Profundiconus
scopulorum Van Mol, Tursch & Kempf, 1971	Conus (Brasiliconus)
scottjordani (Poppe, Monnier & Tagaro, 2012)	Conus (Cylinder)
sculletti Marsh, 1962	Conus (Papyriconus)
sculpturatus Röckel & da Motta, 1986	Conus (Phasmoconus)
sennottorum Rehder & Abbott, 1951	Conus (Dauciconus)
serranegrae Rolán, 1990	Conus (Lautoconus)
sertacinctus Röckel, 1986	Conus (Phasmoconus)
shaskyi (Tenorio, Tucker & Chaney, 2012)	Conus (Dauciconus)
shikamai Coomans, Moolenbeek & Wils, 1985	Conus (Splinoconus)
sieboldii (Reeve, 1848)	Conasprella (Endemoconus)
silviae (Cossignani, 2014)	Conus (Lautoconus)
simonei (Petuch & R. F. Myers, 2014)	Conasprella (Ximeniconus)
skoglundae (Tenorio, Tucker & Chaney, 2012)	Conus (Dauciconus)
smirna (Bartsch & Rehder, 1943)	Profundiconus
sogodensis (Poppe, Monnier & Tagaro, 2012)	Conus (Phasmoconus)
solangeae Bozzetti, 2004	Conus (Textilia)
solidus Gmelin, 1791	Conus (Stephanoconus)
solomonensis Delsaerdt, 1992	Conus (Phasmoconus)
somalica (Bozzetti, 2013)	Conasprella (Endemoconus)
spectrum Linnaeus, 1758	Conus (Phasmoconus)
sphacelatus G. B. Sowerby I, 1833	Conus (Dauciconus)
spiceri Bartsch & Rehder, 1943	Conus (Virgiconus)
spirofilis (Habe & Kosuge, 1970)	Conasprella (Endemoconus)
splendidulus G. B. Sowerby I, 1833	Conus (Strategoconus)
sponsalis Hwass in Bruguière, 1792	Conus (Harmoniconus)
spurius Gmelin, 1791	Conus (Lindaconus)
stahlschmidti (Tenorio & Tucker, 2014)	Profundiconus
stanfieldi Petuch, 1998	Conus (Dauciconus)
stearnsii (Conrad, 1869)	Conasprella (Ximeniconus)
stercusmuscarum Linnaeus, 1758	Conus (Pionoconus)
stimpsoni Dall, 1902	Conus (Dauciconus)
stocki (Coomans & Moolenbeek, 1990)	Conasprella (Fusiconus)
stramineus Lamarck, 1810	Conus (Phasmoconus)
straturatus G. B. Sowerby II, 1865	Conus (Phasmoconus)
striatellus Link, 1807	Conus (Strategoconus)
striatus Linnaeus, 1758	Conus (Pionoconus)
striolatus Kiener, 1848	Conus (Pionoconus)
stupa (Kuroda, 1956)	Conus (Turriconus)
stupella (Kuroda, 1956)	Conus (Turriconus)
suduirauti Raybaudi Massilia, 2004	Conus (Strategoconus)
sugimotonis Kuroda, 1928	Conus (Klemaeconus)
sukhadwalai Röckel & da Motta, 1983	Conus (Splinoconus)
sulcatus Hwass in Bruguière, 1792	Conus (Asprella)
sulcocastaneus Kosuge, 1981	Conus (Asprella)
sunderlandi Petuch, 1987	Conus (Dauciconus)
suratensis Hwass in Bruguière, 1792	Conus (Dendroconus)
sutanorcum Moolenbeek, Röckel & Bouchet, 2008	Conus (Phasmoconus)
suturatus Reeve, 1844	Conus (Tesselliconus)
swainsoni Estival & Cosel, 1986	Conus (Strategoconus)
swinneni (Tenorio, Afonso, Cunha & Rolán, 2014)	Conus (Lautoconus)
sydneyensis G. B. Sowerby III, 1887	Conus (Mamiconus)
tabidus Reeve, 1844	Conus (Monteiroconus)
tacomae Boyer & Pelorce, 2009	Conus (Lautoconus)
taeniatus Hwass in Bruguière, 1792	Conus (Virroconus)
tagaroae (Limpalaër & Monnier, 2013)	Conus (Cylinder)
taslei Kiener, 1850	Conus (Lautoconus)
telatus Reeve, 1848	Conus (Cylinder)
tenuilineatus Rolán & Röckel, 2001	Conus (Lautoconus)
tenuistriatus G. B. Sowerby II, 1858	Conus (Leporiconus)
teodorae Rolán & Fernandes, 1990	Conus (Lautoconus)
teramachii (Kuroda, 1956)	Profundiconus
terebra Born, 1778	Conus (Virgiconus)
terryni Tenorio & Poppe, 2004	Conus (Pseudopterygia)
tessulatus Born, 1778	Conus (Tesselliconus)
tethys (Petuch & Sargent, 2011)	Conus (Virgiconus)
textile Linnaeus, 1758	Conus (Cylinder)
thalassiarchus G. B. Sowerby I, 1834	Conus (Calibanus)
theodorei Petuch, 2000	Conus (Dauciconus)
therriaulti (Petuch, 2013)	Conus (Lindaconus)
thevenardensis da Motta, 1987	Conus (Splinoconus)
thomae Gmelin, 1791	Conus (Darioconus)
tiaratus G. B. Sowerby I, 1833	Conus (Virroconus)
tiki (Moolenbeek, Zandbergen & Bouchet, 2008)	Conasprella (Fusiconus)
timorensis Hwass in Bruguière, 1792	Conus (Textilia)
tinianus Hwass in Bruguière, 1792	Conus (Floraconus)
tirardi Röckel & Moolenbeek, 1996	Conus (Rhizoconus)
tisii T. C. Lan, 1978	Conus (Embrikena)
tornata (G. B. Sowerby I, 1833)	Conasprella (Ximeniconus)
tostesi Petuch, 1986	Conus (Lamniconus)
traceyi (Tucker & Stahlschmidt, 2010)	Conasprella (Pseudoconorbis)
traillii A. Adams, 1855	Conus (Pseudolilliconus)
traversiana (E. A. Smith, 1875)	Conasprella (Fusiconus)
trencarti Nolf & Verstraeten, 2008	Conus (Lautoconus)
tribblei Walls, 1977	Conus (Splinoconus)
trigonus Reeve, 1848	Conus (Plicaustraconus)
tristensis Petuch, 1987	Conus (Dauciconus)
trochulus Reeve, 1844	Conus (Kalloconus)
troendlei Moolenbeek, Zandbergen & Bouchet, 2008	Conus (Splinoconus)
trovaoi Rolán & Röckel, 2000	Conus (Lautoconus)
tuberculosus (Tomlin, 1937)	Profundiconus
tulipa Linnaeus, 1758	Conus (Gastridium)
tuticorinensis Röckel & Korn, 1990	Conus (Quasiconus)
typhon Kilburn, 1975	Conus (Splinoconus)
unifasciatus Kiener, 1850	Conus (Lautoconus)
urashimanus Kuroda & Itô, 1961	Conus (Splinoconus)
vanhyningi (Rehder, 1944)	Conasprella (Ximeniconus)
vanvilstereni (Moolenbeek & Zandbergen, 2013)	Conus (Splinoconus)
vappereaui Monteiro, 2009	Conus (Phasmoconus)
variegatus Kiener, 1848	Conus (Lautoconus)
varius Linnaeus, 1758	Conus (Strategoconus)
vaubani (Röckel, Richard & Moolenbeek, 1995)	Profundiconus
vautieri Kiener, 1847	Conus (Puncticulis)
vayssierei Pallary, 1906	Conus (Lautoconus)
velaensis Petuch, 1993	Conus (Dauciconus)
venezuelanus Petuch, 1987	Conus (Dauciconus)
ventricosus Gmelin, 1791	Conus (Lautoconus)
venulatus Hwass in Bruguière, 1792	Conus (Kalloconus)
verdensis Trovão, 1979	Conus (Lautoconus)
vexillum Gmelin, 1791	Conus (Rhizoconus)
victoriae Reeve, 1843	Conus (Cylinder)
vicweei Old, 1973	Conus (Textilia)
vidua Reeve, 1843	Conus (Conus)
vikingorum Petuch, 1993	Conus (Dauciconus)
villepinii P. Fischer & Bernardi, 1857	Conus (Dauciconus)
viminea (Reeve, 1849)	Conasprella (Fusiconus)
viola Cernohorsky, 1977	Conus (Splinoconus)
violaceus Gmelin, 1791	Conus (Virgiconus)
virgatus Reeve, 1849	Conus (Dauciconus)
virgo Linnaeus, 1758	Conus (Virgiconus)
visagenus Kilburn, 1974	Conus (Plicaustraconus)
visseri Delsaerdt, 1990	Conus (Pseudolilliconus)
vittatus Hwass in Bruguière, 1792	Conus (Dauciconus)
voluminalis Reeve, 1843	Conus (Splinoconus)
vulcanus Tenorio & Afonso, 2004	Conus (Lautoconus)
wakayamaensis (Kuroda, 1956)	Conasprella (Conasprella)
wallacei (Lorenz & Morrison, 2004)	Conus (Pseudolilliconus)
wallangra (Garrard, 1961)	Conus (Plicaustraconus)
wandae (Cossignani, 2014)	Conus (Lautoconus)
wendrosi (Tenorio & Afonso, 2013)	Conasprella (Ximeniconus)
wilsi Delsaerdt, 1998	Conus (Rhizoconus)
wittigi Walls, 1977	Conus (Phasmoconus)
xanthicus Dall, 1910	Conus (Dauciconus)
xanthocinctus Petuch, 1986	Conus (Lamniconus)
xicoi Röckel, 1987	Conus (Lautoconus)
ximenes (Gray, 1839)	Conasprella (Ximeniconus)
yemenensis Bondarev, 1997	Conus (Phasmoconus)
zandbergeni Filmer & Moolenbeek, 2010	Conus (Phasmoconus)
zapatosensis Röckel, 1987	Conus (Phasmoconus)
zebra Lamarck, 1810	Conus (Phasmoconus)
zebroides Kiener, 1848	Conus (Lautoconus)
zeylanicus Gmelin, 1791	Conus (Puncticulis)
ziczac Mühlfeld, 1816	Conus (Dauciconus)
zinhoi (Cossignani, 2014)	Conus (Lautoconus)
zonatus Hwass in Bruguière, 1792	Conus (Stephanoconus)
zylmanae Petuch, 1998	Conus (Dauciconus)

Appendix Table 2.

Nominal genus-group names of Recent Conidae and their current status in the present classification.

Name	Status
Afonsoconus Tucker & Tenorio, 2013	Conus (Afonsoconus)
Africonus Petuch, 1975	synonym of Conus (Lautoconus)
ArubaconusPetuch, 2013	synonym of Conus (Ductoconus)
Asprella Schaufuss, 1869	Conus (Asprella)
Atlanticonus Petuch & Sergent, 2012	Conus (Atlanticonus)
Attenuiconus Petuch, 2013	Conus (Attenuiconus)
Austroconus Tucker & Tenorio, 2009	synonym of Conus (Mamiconus)
Bathyconus Tucker & Tenorio, 2009	synonym of Conasprella (Fusiconus)
Bermudaconus Petuch, 2013	Conus (Bermudaconus)
Boucheticonus Tucker & Tenorio, 2013	Conasprella (Boucheticonus)
Brasiliconus Petuch, 2013	Conus (Brasiliconus)
Calamiconus Tucker & Tenorio, 2009	synonym of Conus (Lividiconus)
Calibanus da Motta, 1991	Conus (Calibanus)
Californiconus Tucker & Tenorio, 2009	Californiconus
Cariboconus Petuch, 2003	synonym of Conus (Dauciconus)
Chelyconus Mörch, 1852	Conus (Chelyconus)
Cleobula Iredale, 1930	synonym of Conus (Dendroconus)
Coltroconus Petuch, 2013	Conasprella (Coltroconus)
Conasprella Thiele, 1929	Conasprella (Conasprella)
Conasprelloides Tucker & Tenorio, 2009	synonym of Conus (Dauciconus)
Continuconus Tucker & Tenorio, 2013	synonym of Conus (Klemaeconus)
Conus Linnaeus, 1758	Conus (Conus)
Cornutoconus Suzuki, 1972	synonym of Conus (Taranteconus)
Coronaxis Swainson, 1840	synonym of Conus (Conus)
Cucullus Röding, 1798	synonym of Conus (Conus)
Cylinder Montfort, 1810	synonym of Conus (Cylindrus)
Cylindrella Swainson, 1840	synonym of Conus (Asprella)
Cylindrus Batsch, 1789	Conus (Cylindrus)
Dalliconus Tucker & Tenorio, 2009	Conasprella (Dalliconus)
Darioconus Iredale, 1930	Conus (Darioconus)
Dauciconus Cotton, 1945	Conus (Dauciconus)
Dendroconus Swainson, 1840	Conus (Dendroconus)
Ductoconus da Motta, 1991	Conus (Ductoconus)
Duodenticonus Tucker & Tenorio, 2013	synonym of Conasprella (Conasprella)
Dyraspis Iredale, 1949	synonym of Conus (Virroconus)
Elisaconus Tucker & Tenorio, 2013	Conus (Elisaconus)
Embrikena Iredale, 1937	Conus (Embrikena)
Endemoconus Iredale, 1931	Conasprella (Endemoconus)
Eremiconus Tucker & Tenorio, 2009	Conus (Eremiconus)
Erythroconus da Motta, 1991	synonym of Conus (Darioconus)
Eugeniconus da Motta, 1991	Conus (Eugeniconus)
Floraconus Iredale, 1930	Conus (Floraconus)
Fraterconus Tucker & Tenorio, 2013	Conus (Fraterconus)
Fulgiconus da Motta, 1991	synonym of Conus (Phasmoconus)
Fumiconus da Motta, 1991	synonym of Conasprella (Fusiconus)
Fusiconus da Motta, 1991	Conasprella (Fusiconus)
Gastridium Modeer, 1793	Conus (Gastridium)
Genuanoconus Tucker & Tenorio, 2009	synonym of Conus (Kalloconus)
Gladioconus Tucker & Tenorio, 2009	synonym of Conus (Monteiroconus)
Globiconus Tucker & Tenorio, 2009	synonym of Conasprella (Ximeniconus)
Gradiconus da Motta, 1991	synonym of Conus (Dauciconus)
Graphiconus da Motta, 1991	synonym of Conus (Phasmoconus)
Harmoniconus da Motta, 1991	Conus (Harmoniconus)
Hermes Montfort, 1810	Conus (Hermes)
Heroconus da Motta, 1991	synonym of Conus (Pionoconus)
Isoconus Tucker & Tenorio, 2013	synonym of Conus (Splinoconus)
Jaspidiconus Petuch, 2003	synonym of Conasprella (Ximeniconus)
Kalloconus da Motta, 1991	Conus (Kalloconus)
Kellyconus Petuch, 2013	Conus (Kellyconus)
Kenyonia Brazier, 1896	Conus (Kenyonia)
Kermasprella Powell, 1958	synonym of Conasprella (Endemoconus)
Ketyconus da Motta, 1991	synonym of Conus (Floraconus)
Kioconus da Motta, 1991	synonym of Conus (Splinoconus)
Klemaeconus Tucker & Tenorio, 2013	Conus (Klemaeconus)
Kohniconus Tucker & Tenorio, 2009	Conasprella (Kohniconus)
Kurodaconus Shikama & Habe, 1968	synonym of Conus (Turriconus)
Lamniconus da Motta, 1991	Conus (Lamniconus)
Lautoconus Monterosato, 1923	Conus (Lautoconus)
Leporiconus Iredale, 1930	Conus (Leporiconus)
Leptoconus Swainson, 1840	Conus (Leptoconus)
Lilliconus Raybaudi Massilia, 1994	Conasprella (Lilliconus)
Lindaconus Petuch, 2002	Conus (Lindaconus)
Lithoconus Mörch, 1852	Conus (Lithoconus)
Lividoconus Wils, 1970	Conus (Lividoconus)
Lizaconus da Motta, 1991	synonym of Conus (Profundiconus)
Magelliconus da Motta, 1991	synonym of Conus (Purpuriconus)
Mamiconus Cotton & Godfrey, 1932	Conus (Mamiconus)
Miliariconus Tucker & Tenorio, 2009	synonym of Conus (Virroconus)
Mitraconus Tucker & Tenorio, 2013	synonym of Conus (Turriconus)
Monteiroconus da Motta, 1991	Conus (Monteiroconus)
Nataliconus Tucker & Tenorio, 2009	synonym of Conus (Leptoconus)
Nimboconus Tucker & Tenorio, 2013	synonym of Conus (Phasmoconus)
Nitidoconus Tucker & Tenorio, 2013	synonym of Conus (Splinoconus)
Ongoconus da Motta, 1991	synonym of Conus (Kioconus)
Papyriconus Tucker & Tenorio, 2013	Conus (Papyriconus)
Parviconus Cotton & Godfrey, 1932	Conasprella (Parviconus)
Perplexiconus Tucker & Tenorio, 2009	synonym of Conasprella (Ximeniconus)
Phasmoconus Mörch, 1852	Conus (Phasmoconus)
Pionoconus Mörch, 1852	Conus (Pionoconus)
Plicaustraconus Moolenbeek, 2008	Conus (Plicaustraconus)
Poremskiconus Petuch, 2013	synonym of Conus (Dauciconus)
Profundiconus Kuroda, 1956	Profundiconus
Protoconus da Motta, 1991	synonym of Conus (Tenorioconus)
Protostrioconus Tucker & Tenorio, 2009	synonym of Conus (Gastridium)
Pseudoconorbis Tucker & Tenorio, 2009	Conasprella (Pseudoconorbis)
Pseudohermes Tucker & Tenorio, 2013	synonym of Conus (Virgiconus)
Pseudolilliconus Tucker & Tenorio, 2009	Conus (Pseudolilliconus)
Pseudonoduloconus Tucker & Tenorio, 2009	Conus (Pseudonoduloconus)
Pseudopterygia Tucker & Tenorio, 2013	Conus (Pseudopterygia)
Puncticulis Swainson, 1840	Conus (Puncticulis)
Purpuriconus da Motta, 1991	synonym of Conus (Dauciconus)
Pyruconus Olsson, 1967	Conus (Pyruconus)
Quasiconus Tucker & Tenorio, 2009	Conus (Quasiconus)
Regiconus Iredale, 1930	synonym of Conus (Darioconus)
Rhizoconus Mörch, 1852	Conus (Rhizoconus)
Rhombiconus Tucker & Tenorio, 2009	synonym of Conus (Stephanoconus)
Rhombus Montfort, 1810	synonym of Conus (Rhombiconus)
Rolaniconus Tucker & Tenorio, 2009	synonym of Conus (Strategoconus)
Rollus Montfort, 1810	synonym of Conus (Gastridium)
Rubroconus Tucker & Tenorio, 2013	Conus (Rubroconus)
Sandericonus Petuch, 2013	Conus (Sandericonus)
Sciteconus da Motta, 1991	Conus (Sciteconus)
Seminoleconus Petuch, 2003	synonym of Conus (Protoconus)
Socioconus da Motta, 1991	synonym of Conus (Pionoconus)
Splinoconus da Motta, 1991	Conus (Splinoconus)
Spuriconus Petuch, 2003	synonym of Conus (Lindaconus)
Stellaconus Tucker & Tenorio, 2009	synonym of Conus (Splinoconus)
Stephanoconus Mörch, 1852	Conus (Stephanoconus)
Strategoconus da Motta, 1991	Conus (Strategoconus)
Strioconus Thiele, 1929	synonym of Conus (Pionoconus)
Sulciconus Bielz, 1869	synonym of Conus (Asprella)
Taranteconus Azuma, 1972	synonym of Conus (Stephanoconus)
Tenorioconus Petuch & Drohlshagen, 2011	synonym of Conus (Stephanoconus)
Tesselliconus da Motta, 1991	Conus (Tesselliconus)
Textilia Swainson, 1840	Conus (Textilia)
Thalassiconus Tucker & Tenorio, 2013	synonym of Conus (Calibanus)
Theliconus Swainson, 1840	synonym of Conus (Hermes)
Thoraconus da Motta, 1991	synonym of Conus (Fulgiconus)
Trovaoconus Tucker & Tenorio, 2009	synonym of Conus (Kalloconus)
Tuckericonus Petuch, 2013	synonym of Conus (Dauciconus)
Tuliparia Swainson, 1840	synonym of Conus (Gastridium)
Turriconus Shikama & Habe, 1968	Conus (Turriconus)
Utriculus Schumacher, 1817	synonym of Conus (Gastridium)
Varioconus da Motta, 1991	synonym of Conus (Lautoconus)
Viminiconus Tucker & Tenorio, 2009	synonym of Conasprella (Fusiconus)
Virgiconus Cotton, 1945	Conus (Virgiconus)
Virroconus Iredale, 1930	Conus (Virroconus)
Vituliconus da Motta, 1991	synonym of Conus (Strategoconus)
Ximeniconus Emerson & Old, 1962	Conasprella (Ximeniconus)
Yeddoconus Tucker & Tenorio, 2009	synonym of Conasprella (Endemoconus)

SYSTEMATICS

Superfamily CONOIDEA Fleming, 1822

Family Conidae Fleming, 1822

[Synonyms: Conilithidae Tucker & Tenorio, 2009, n. syn. Taranteconidae Tucker & Tenorio, 2009, n. syn. Puncticuliinae Tucker & Tenorio, 2009, n. syn.]

Genus

Type species: Reeve, 1844; OD

Included species:

(Reeve, 1844)

Remarks: Conus californicus has always been considered a unique species within cone snails, because of its molecular (including toxicological: Biggs ; Elliger ) and morphological singularities and also because of its diet, since it is able to prey indifferently on fish, molluscs and worms (Kohn, 1966).

Genus

Type species: Chelyconus profundorum Kuroda, 1956; OD

[Synonym: Lizaconus da Motta, 1991; type species: Kuroda, 1956; OD]

Included species:

cakobaui (Moolenbeek, Röckel & Bouchet, 2008)

dondani (Kosuge, 1981) n. comb.

emersoni (G. D. Hanna, 1963)

frausseni (Tenorio & Poppe, 2004)

ikedai (Ninomiya, 1987)

jeanmartini (G. Raybaudi Massilia, 1992)

(Richard, 1983) n. comb.

lani (Crandall, 1979)

loyaltiensis (Röckel & Moolenbeek, 1995)

pacificus (Moolenbeek & Röckel, 1996) n. comb.

(Kuroda, 1956)

scopulicola (Okutani, 1972)

(Bartsch & Rehder, 1943)

stahlschmidti (Tenorio & Tucker, 2014)

(Kuroda, 1956)

tuberculosus (Tomlin, 1937) n. comb.

(Röckel, Richard & Moolenbeek, 1995)

Remarks: Puillandre did not sequence the type species of Profundiconus; their sequenced material was identified as P. aff. profundorum. However, Tucker & Tenorio (2009, 2013) found Profundiconus to be a morphologically well supported group and we are thus confident in applying this name to the clade containing P. aff. profundorum.

Genus

Subgenus

Type species: Conus pagoda Kiener, 1847; SD, Tucker & Tenorio (2009: 140) (under ICZN, 1999: Art. 70.3.2).

[Synonym: Duodenticonus Tucker & Tenorio 2013; type species: Habe & Kosuge, 1970; OD; n. syn.]

Included species:

(Petuch, 1979)

(G.B. Sowerby II, 1873)

(Röckel & da Motta, 1979)

(Petuch, 1979) n. comb.

(Richard, 1983) n. comb.

bozzettii (Lauer, 1991)

(G.B. Sowerby III, 1903)

(Bartsch & Rehder, 1943)

gordyi (Röckel & Bondarev, 2000)

lorenzi Monnier & Limpalaër, 2012

(Habe & Kosuge, 1970) n. comb.

(Kuroda & Itô, 1961) n. comb.

(Kiener, 1847)

wakayamaensis (Kuroda, 1956)

Subgenus

Type species: Moolenbeek, Röckel & Richard, 1995; OD

Included species: Remarks: The species C. kimioi was placed in the subgenus Boucheticonus (following Tucker & Tenorio 2013), even if the corresponding clade was not highly supported.

(Moolenbeek, Röckel & Richard, 1995) n. comb.

(Habe, 1965) n. comb.

pseudokimioi da Motta & Martin, 1982 n. comb.

Subgenus

Type species: Conus iansa Petuch, 1979; OD

Included species:

henriquei (Petuch & R. F. Myers, 2014) n. comb.

iansa (Petuch, 1979) n. comb.

Subgenus

Type species: Conus mcgintyi Pilsbry, 1955; OD

Included species: Subgenus

armiger (Crosse, 1858) n. comb.

bajanensis (Nowell-Usticke, 1968) n. comb.

guyanensis (van Mol, 1973) n. comb.

lenhilli (Cargile, 1998) n. comb.

(Deshayes, 1874) n. comb.

mcgintyi (Pilsbry, 1955) n. comb.

pacei (Petuch, 1987) n. comb.

rainesae (McGinty, 1953) n. comb.

roberti (Richard, 2009) n. comb.

sauros (Garcia, 2006) n. comb.

Type species: Conus howelli Iredale, 1929; OD

[Synonyms: Kermasprella Powell, 1958; type species: Conus raoulensis Powell, 1958; OD. Yeddoconus Tucker & Tenorio, 2009; type species: Reeve, 1848; OD]

Included species:

grohi (Tenorio & Poppe, 2004) n. comb.

howelli (Iredale, 1929) n. comb.

(Fulton, 1938) n. comb.

raoulensis (Powell, 1958) n. comb.

(Reeve, 1848) n. comb.

somalica (Bozzetti, 2013) n. comb.

spirofilis (Habe & Kosuge, 1970) n. comb.

Subgenus

Type species: Kiener, 18547; OD

[Synonyms: Bathyconus Tucker & Tenorio, 2009; type species: Audouin, 1831; OD; n. syn. Fumiconus da Motta, 1991; type species: Conus traversianus E.A. Smith, 1875, OD; n. syn. Viminiconus Tucker & Tenorio, 2009; type species: Reeve, 1849; OD; n. syn.]

Included species: Subgenus

aculeiformis (Reeve, 1844) n. comb.

(Pilsbry, 1904) n. comb.

dictator (Melvill, 1898) n. comb.

dieteri (Moolenbeek, Zandbergen & Bouchet, 2008) n. comb.

elegans (G. B. Sowerby III, 1895) n. comb.

(Kilburn, 1975) n. comb.

fijiensis (Moolenbeek, Röckel & Bouchet, 2008) n. comb.

(G. Raybaudi Massilia, 2005) n. comb.

hypochlorus (Tomlin, 1937) n. comb.

(Tomlin, 1937) n. comb.

(Kuroda, 1956) n. comb.

insculpta (Kiener, 1847) n. comb.

lentiginosa (Reeve, 1844) n. comb.

(Moolenbeek, Röckel & Bouchet, 2008) n. comb.

(Röckel, Richard & Moolenbeek, 1995) n. comb.

lizarum (Raybaudi Massilia & da Motta, 1992) n. comb.

(Kiener, 1847) n. comb.

(Audouin, 1831) n. comb.

pepeiu (Moolenbeek, Zandbergen & Bouchet, 2008) n. comb.

(Röckel & Lan, 1981) n. comb.

saecularis (Melvill, 1898) n. comb.

stocki (Coomans & Moolenbeek, 1990) n. comb.

tiki (Moolenbeek, Zandbergen & Bouchet, 2008) n. comb.

traversiana (E. A. Smith, 1875) n. comb.

(Reeve, 1849) n. comb.

Type species: Conus emarginatus Reeve, 1844; OD

Included species: Remarks: Tucker & Tenorio (2009) included both Conasprella delesserti and Conasprella arcuata in Kohniconus. In our analysis, the two species do not cluster together. The name Kohniconus could have been applied to the lineage that includes the species C. delessertii, but because of the closer morphological resemblance of Conus emarginatus with C. arcuata, we have applied Kohniconus to the lineage that includes C. arcuata.

(Broderip & G. B. Sowerby I, 1829) n. comb.

centurio (Born, 1778) n. comb.

emarginata (Reeve, 1844) n. comb.

fenzani (Petuch & Sargent, 2011) n. comb.

janowskyae (Tucker & Tenorio, 2011) n. comb.

kohni (McLean & Nybakken, 1979) n. comb.

rachelae (Petuch, 1988) n. comb.

scaripha (Dall, 1910) n. comb.

Subgenus

Type species: Conus biraghii G. Raybaudi Massilia, 1992; OD

Included species: Subgenus

biraghii (G. Raybaudi Massilia, 1992) n. comb.

sagei (Korn & G. Raybaudi Massilia, 1993) n. comb.

Type species: Conus rutilus Menke, 1843; OD

Included species:

rutila (Menke, 1843) n. comb.

Subgenus

Type species: Conorbis coromandelicus E. A. Smith, 1894; OD

Included species:

coromandelica (E. A. Smith, 1894) n. comb.

traceyi (Tucker & Stahlschmidt, 2010) n. comb.

Subgenus

Type species: Gray, 1839; OD

[Synonyms: Globiconus Tucker & Tenorio, 2009; type species: G.B. Sowerby I, 1833; OD; n. syn. Jaspidiconus Petuch, 2003; type species: Gmelin, 1791; OD; n. syn. Perplexiconus Tucker & Tenorio, 2009; type species: G. B. Sowerby II, 1857; OD; n. syn.]

Included species:

alexandremonteiroi (Cossignani, 2014) n. comb.

allamandi (Petuch, 2013) n. comb.

anaglyptica (Crosse, 1865) n. comb.

arawak (Petuch & R. F. Myers, 2014) n. comb.

baccata (G. B. Sowerby III, 1877) n. comb.

berschaueri (Petuch & R. F. Myers, 2014) n. comb.

damasoi (Cossignani, 2007) n. comb.

ericmonnieri (Petuch & R. F. Myers, 2014) n. comb.

fluviamaris (Petuch & Sargent, 2011) n. comb.

henckesi (Coltro, 2004) n. comb.

herndli (Petuch & R. F. Myers, 2014) n. comb.

(Gmelin, 1791) n. comb.

(W. Wood, 1828) n. comb.

(Reeve, 1843) n. comb.

(Hwass in Bruguière, 1792) n. comb.

ogum (Petuch & R. F. Myers, 2014) n. comb.

(G. B. Sowerby II, 1857) n. comb.

pfluegeri (Petuch, 2003) n. comb.

poremskii (Petuch & R. F. Myers, 2014) n. comb.

(Hwass in Bruguière, 1792) n. comb.

pusio (Hwass in Bruguière, 1792) n. comb.

roatanensis (Petuch & Sargent, 2011) n. comb.

sargenti (Petuch, 2013) n. comb.

simonei (Petuch & R. F. Myers, 2014) n. comb.

(Conrad, 1869) n. comb.

(G. B. Sowerby I, 1833) n. comb.

vanhyningi (Rehder, 1944) n. comb.

wendrosi (Tenorio & Afonso, 2013) n. comb.

(Gray, 1839) n. comb.

Incertae sedis

(Récluz, 1843) n. comb.

Genus

Subgenus

Type species: Linnaeus, 1758; SD, Children (1823: 69)

[Synonyms: Cucullus Röding, 1798; type species: Linnaeus, 1758; SD, Winckworth (1945: 139). Coronaxis Swainson, 1840; type species: Hwass, 1792; M]

Included species:

[Lightfoot], 1786

Hwass in Bruguière, 1792

Linnaeus, 1758

nocturnus [Lightfoot], 1786

vidua Reeve, 1843

Subgenus

Type species: Kuroda, 1956; OD

Included species:

Powell, 1958

Kuroda, 1956

Subgenus

Type species: Conus asper Lamarck, 1810, by typification of replaced name. Asprella was established as a substitute name for Cylindrella Swainson, 1840 (see below). Conus asper was not among the 11 species included in Asprella by Schaufuss (1869: 43–44), and Wenz (1940) cited Conus sulcatus Bruguière, 1792 (a species originally included by Schaufuss) as the type species; Conus sulcatus and C. asper are subjective synonyms.

[Synonyms: Cylindrella Swainson, 1840; type species: Conus asper Lamarck, 1810; subjective synonym of Hwass, 1792; M. Cylindrella Swainson, 1840 (Conidae), is a homonym of Cylindrella Swainson, 1840 (Urocoptidae) and has been placed on the Official Index of Rejected and Invalid Names by ICZN, 1999: Opinion 1030. Sulciconus Bielz, 1869; type species (here designated): Hwass in Bruguière, 1792, n. syn. The names Asprella and Sulciconus were both published in 1869 and their exact dates of publication are not known in order to establish priority. Under ICZN (1999): Art. 24.2, we act here as First Revisers and give precedence to the name Asprella over Sulciconus.]

Included species:

fijisulcatus Moolenbeek, Röckel & Bouchet, 2008

gigasulcatus Moolenbeek, Röckel & Bouchet, 2008

G. B. Sowerby III, 1900

helgae Blöcher, 1992

iodostoma Reeve, 1843

Röckel, 1986

Hwass in Bruguière, 1792

sulcocastaneus Kosuge, 1981

Subgenus

Type species: Conus granulatus Linnaeus, 1758; OD

Included species:

cuna Petuch, 1998

glenni Petuch, 1993

granulatus Linnaeus, 1758

ritae Petuch, 1995

Subgenus

Type species: Conus attenuatus Reeve, 1844; OD

Included species: Subgenus

attenuatus Reeve, 1844

aureonimbosus Petuch, 1987

eversoni Petuch, 1987

honkeri Petuch, 1988

poulosi Petuch, 1993

Type species: Conus cyanostoma A. Adams, 1855; OD

[Possible senior synonym: Mamiconus Cotton & Godfrey, 1932; type species: Conus superstes Hedley, 1911; OD. The identity of C. superstes is uncertain; it is listed as a nomen dubium by Tucker & Tenorio (2013)]

Included species:

clarus E. A. Smith, 1881

cyanostoma A. Adams, 1855

sydneyensis G. B. Sowerby III, 1887

Subgenus

Type species: Conus lightbourni Petuch, 1986; OD

Included species:

lightbourni Petuch, 1986

Subgenus

Type species: Conus scopulorum Van Mol, Tursch & Kempf, 1971; OD

Included species:

scopulorum Van Mol, Tursch & Kempf, 1971

Subgenus

Type species: Reeve, 1843; OD

[Synonym: Thalassiconus Tucker & Tenorio, 2013; type species: G.B. Sowerby I, 1834, n. syn.; OD]

Included species:

Reeve, 1843

G. B. Sowerby I, 1834

Subgenus

Type species: Conus testudinarius Hwass, 1792 [synonym of Born, 1778]; SD, Cossmann (1896: 160)

Included species:

Born, 1778

G. B. Sowerby I, 1833

Subgenus

Type species: Linnaeus, 1758; OD. Dubois & Bour (2010) revived the name Cylindrus as from Batsch, 1789 and designated Conus textile as its type species. This has the consequence of destabilizing the pulmonate genus name Cylindrus Fitzinger, 1833, which is in current use. The ICZN will be petitioned to maintain usage of Cylinder Montfort, 1810, and Cylindrus Fitzinger, 1833, in their accustomed sense.]

Included species:

abbas Hwass in Bruguière, 1792

Linnaeus, 1758

Hwass in Bruguière, 1792

barbieri G. Raybaudi Massilia, 1995

(Okutani, 1968)

Hwass in Bruguière, 1792

Stearns, 1873

Chemnitz, 1777

glorioceanus Poppe & Tagaro, 2009

Lamarck, 1810

nodulosus G. B. Sowerby II, 1864

priscai (Bozzetti, 2012) n. comb.

Menke, 1829

scottjordani (Poppe, Monnier & Tagaro, 2012) n. comb.

tagaroae (Limpalaër & Monnier, 2013) n. comb.

telatus Reeve, 1848

Linnaeus, 1758

Reeve, 1843

Remarks: Tucker & Tenorio (2009) described two ‘body plans’ in their genus Cylinder, citing Conus textile and C. gloriamaris as examples of each group; they may correspond to the two groups found in the molecular phylogeny. Tucker & Tenorio (2009) relied on their similar radulae and diets to maintain them in a single genus: we have followed them and place all the species in a single subgenus. Among the sequenced species, C. ammiralis, C. canonicus and C. dalli cluster with the type species C. textile; the other species constitute a separate clade.

Subgenus

Type species: Hwass, 1792; OD

[Synonyms: Regiconus Iredale, 1930; type species: Linnaeus, 1758; OD; the simultaneously published Darioconus was given precedence over Regiconus by First Reviser Kira (1954: 75). Erythroconus da Motta, 1991; type species: Lamarck, 1810; OD]

Included species:

Linnaeus, 1758

auratinus da Motta, 1982

Hwass in Bruguière, 1792

behelokensis Lauer, 1989

biancae Bozzetti, 2010

Lamarck, 1810

echo Lauer, 1989

da Motta, 1982

lamberti Souverbie, 1877

laueri (Monnier & Limpalaër, 2013) n. comb.

leviteni (Tucker, Tenorio & Chaney, 2011) n. comb.

Kilburn, 1972

madagascariensis G. B. Sowerby II, 1858

Reeve, 1843

natalaurantius (S. G. Veldsman, 2013) n. comb.

Hwass in Bruguière, 1792

Born, 1778

Gmelin, 1791

Remarks: The species C. thomae was placed in the subgenus Darioconus (following Tucker & Tenorio 2013), even though the corresponding clade was not highly supported.

Subgenus

Type species: Hwass, 1792; OD

[Synonyms: Gradiconus da Motta, 1991; type species: W. Wood, 1828; OD; n. syn.; Magelliconus da Motta, 1991; type species: Conus magellanicus Hwass in Bruguière, 1792; OD; n. syn.; Purpuriconus da Motta, 1991; type species: Conus cardinalis Hwass in Bruguière, 1792; OD; n. syn.; Cariboconus Petuch, 2003; type species: Conus amphiurgus Dall, 1889; OD; Conasprelloides Tucker & Tenorio, 2009; type species: Hwass in Bruguière, 1792; OD; n. syn.; Poremskiconus Petuch, 2013; type species: Conus ziczac Mühlfeld, 1816; OD; n. syn.; Tuckericonus Petuch 2013; type species: G. B. Sowerby I, 1834; OD; n. syn.]

Included species:

abrolhosensis Petuch, 1987

alainallaryi Bozzetti & Monnier, 2009

Dall, 1889

Crosse, 1865

Sarasúa, 1977

aureopunctatus Petuch, 1987

bayeri Petuch, 1987

belizeanus (Petuch & Sargent, 2011) n. comb.

bessei Petuch, 1992

boui da Motta, 1988

brunneobandatus Petuch, 1992

brunneofilaris Petuch, 1990

burryae Clench, 1942

dianthus G. B. Sowerby III, 1882

dispar G. B. Sowerby I, 1833

donnae Petuch, 1998

edwardpauli Petuch, 1998

eleutheraensis (Petuch, 2013) n. comb.

ernesti Petuch, 1990

explorator Vink, 1990

flamingo Petuch, 1980

flammeacolor Petuch, 1992

G. B. Sowerby I, 1834

garciai da Motta, 1982

gibsonsmithorum Petuch, 1986

glicksteini Petuch, 1987

goajira Petuch, 1992

W. Wood, 1828

harasewychi Petuch, 1987

havanensis Aguayo & Pérez Farfante, 1947

hennequini Petuch, 1993

hilli Petuch, 1990

honkerorum (Petuch & R. F. Myers, 2014) n. comb.

inconstans E. A. Smith, 1877

Petuch, 1998

jorioi (Petuch, 2013) n. comb.

G. B. Sowerby III, 1887

kaiserae (Tenorio, Tucker & Chaney, 2012) n. comb.

kalafuti da Motta, 1987

kerstitchi Walls, 1978

kevani Petuch, 1987

kirkandersi Petuch, 1987

kremerorum Petuch, 1988

kulkulcan Petuch, 1980

largilliertii Kiener, 1847

leekremeri Petuch, 1987

levistimpsoni (Tucker, 2013) n. comb.

lucaya Petuch, 2000

maculiferus G. B. Sowerby I, 1833

magellanicus Hwass in Bruguière, 1792

magnottei Petuch, 1987

maya (Petuch & Sargent, 2011) n. comb.

monilifer Broderip, 1833

Hwass in Bruguière, 1792

cardinalis Hwass in Bruguière, 1792

cargilei Coltro, 2004

castaneus Kiener, 1848

caysalensis L. Raybaudi & Prati, 1994

ceruttii Cargile, 1997

cingulatus Lamarck, 1810

colombi (Monnier & Limpalaër, 2012) n. comb.

colombianus Petuch, 1987

coltrorum (Petuch & R. F. Myers, 2014) n. comb.

Hwass in Bruguière, 1792

deynzerorum Petuch, 1995

nybakkeni (Tenorio, Tucker & Chaney, 2012) n. comb.

Broderip, 1833

ortneri Petuch, 1998

ostrinus (Tucker & Tenorio, 2011) n. comb.

paraguana Petuch, 1987

parascalaris Petuch, 1987

paschalli Petuch, 1998

patglicksteinae Petuch, 1987

paulae Petuch, 1988

penchaszadehi Petuch, 1986

Kiener, 1847

Berry, 1968

portobeloensis Petuch, 1990

pseudocardinalis Coltro, 2004

Broderip, 1833

regularis G. B. Sowerby I, 1833

richardbinghami Petuch, 1993

rosalindensis Petuch, 1998

rosemaryae Petuch, 1990

sahlbergi da Motta & Harland, 1986

scalaris Valenciennes, 1832

scalarissimus da Motta, 1988

sennottorum Rehder & Abbott, 1951

shaskyi (Tenorio, Tucker & Chaney, 2012) n. comb.

skoglundae (Tenorio, Tucker & Chaney, 2012) n. comb.

sphacelatus G. B. Sowerby I, 1833

stanfieldi Petuch, 1998

stimpsoni Dall, 1902

sunderlandi Petuch, 1987

theodorei Petuch, 2000

tristensis Petuch, 1987

velaensis Petuch, 1993

venezuelanus Petuch, 1987

vikingorum Petuch, 1993

P. Fischer & Bernardi, 1857

Reeve, 1849

Hwass in Bruguière, 1792

xanthicus Dall, 1910

ziczac Mühlfeld, 1816

Petuch, 1998

Subgenus

Type species: Linnaeus, 1758; SD, Herrmannsen (1847: 177)

[Synonym: Cleobula Iredale, 1930; type species: Linnaeus, 1758; OD]

Included species: Remarks: The species C. medoci was placed in the subgenus Dendroconus (following Tucker and Tenorio 2013), even if the corresponding clade was not highly supported

Linnaeus, 1758

(Röding, 1798)

Linnaeus, 1758

glaucus Linnaeus, 1758

Lorenz, 2004

suratensis Hwass in Bruguière, 1792

Subgenus

Type species: Linnaeus, 1758; OD

[Synonym: Arubaconus Petuch, 2013; type species: Duclos, 1833; OD; n. syn.]

Included species:

Duclos, 1833

Linnaeus, 1758

Subgenus

Type species: Linnaeus, 1758; OD

Included species:

Linnaeus, 1758

Subgenus

Type species: Iredale, 1937; M

Included species:

darkini Röckel, Korn & Richard, 1993

kostini Filmer, Monteiro, Lorenz & Verdasca, 2012

moncuri Filmer, 2005

(Iredale, 1937)

tisii T. C. Lan, 1978

Subgenus

Type species: Conus minnamurra (Garrard, 1961); OD

Included species:

albellus Röckel & Korn, 1990

colmani Röckel & Korn, 1990

limpusi Röckel & Korn, 1990

lizardensis Crosse, 1865

minnamurra (Garrard, 1961)

Subgenus

Type species: Linnaeus, 1758; OD

Included species: Subgenus

cordigera G. B. Sowerby II, 1866

marchionatus Hinds, 1843

Linnaeus, 1758

Type species: Lamarck, 1810; OD

[Synonym: Ketyconus da Motta, 1991; type species: Hwass in Bruguière, 1792; OD; n. syn.]

Included species:

Lamarck, 1810

aplustre Reeve, 1843

Kiener, 1850

G. B. Sowerby I, 1833

cocceus Reeve, 1844

G. B. Sowerby II, 1866

gabelishi da Motta & Ninomiya, 1982

Hwass in Bruguière, 1792

papilliferus G. B. Sowerby I, 1834

Hwass in Bruguière, 1792

Subgenus

Type species: Hwass in Bruguière, 1792; OD

Included species:

Hwass in Bruguière, 1792

Subgenus

Type species: Linnaeus, 1758; SD, Winckworth (1935: 322)

[Synonyms: Rollus Montfort, 1810; type species: Linnaeus, 1758; OD. Utriculus Schumacher, 1817; type species: Linnaeus, 1758; M. Tuliparia Swainson, 1840; type species: Coronaxis nebulosa Swainson, 1840 [synonym of ]; M. Protostrioconus Tucker & Tenorio, 2009; type species: G.B. Sowerby I, 1833; OD; n. syn.]

Included species:

Crosse, 1858

eldredi Morrison, 1955

fragilissimus Petuch, 1979

Linnaeus, 1758

G. B. Sowerby I, 1833

Linnaeus, 1758

Subgenus

Type species: Hwass in Bruguière, 1792; OD

Included species:

mcbridei Lorenz, 2005

Hwass in Bruguière, 1792

G. B. Sowerby I, 1833

Broderip, 1833

Walls, 1979

paukstisi (Tucker, Tenorio & Chaney, 2011) n. comb.

Hwass in Bruguière, 1792

Subgenus

Type species: Linnaeus, 1758; OD

[Synonym: Theliconus Swainson, 1840; type species: Linnaeus, 1758; OD]

Included species:

Linnaeus, 1758

Subgenus

Type species: [Lightfoot], 1786; OD

[Synonyms: Genuanoconus Tucker & Tenorio, 2009; type species: Linnaeus, 1758; OD; n. syn. Trovaoconus Tucker & Tenorio, 2009; type species: Hwass in Bruguière, 1792; OD; n. syn.]

Included species:

Kiener, 1850

atlanticoselvagem Afonso & Tenorio, 2004

(Röding, 1798)

Linnaeus, 1758

Monteiro, Tenorio & Poppe, 2004

[Lightfoot], 1786

Reeve, 1844

Hwass in Bruguière, 1792

Subgenus

Type species: Conus patae Abbott, 1971; OD

Included species:

binghamae Petuch, 1987

ignotus Cargile, 1998

patae Abbott, 1971

Subgenus

Type species: Cotton, 1953; OD

[Synonym: Continuconus Tucker & Tenorio, 2013; type species: Richard & Moolenbeek, 1988; OD; n. syn.]

Included species:

estivali Moolenbeek & Richard, 1995

(Kuroda, 1956)

(Cotton, 1953)

Richard & Moolenbeek, 1988

rufimaculosus Macpherson, 1959

Kuroda, 1928

Subgenus

Type species: Conus clerii Reeve, 1844; OD

Included species:

carcellesi Martins, 1945

clerii Reeve, 1844

lemniscatus Reeve, 1849

patriceae (Petuch & R. F. Myers, 2014) n. comb.

tostesi Petuch, 1986

xanthocinctus Petuch, 1986

Subgenus

Type species: Conus mediterraneus Hwass in Bruguière, 1792 [synonym of Gmelin, 1791]; OD

[Synonyms: Africonus Petuch, 1975; type species: Reeve, 1843; OD; n. syn. Varioconus da Motta, 1991; type species: Conus variegatus Kiener, 1848 OD; n. syn.]

Included species:

aemulus Reeve, 1844

africanus Kiener, 1848

albuquerquei Trovão, 1978

alexandrinus Kaicher, 1977

allaryi Bozzetti, 2008

anabelae Rolán & Röckel, 2001

anthonyi (Petuch, 1975)

antonioi (Cossignani, 2014) n. comb.

antoniomonteiroi Rolán, 1990

babaensis Rolán & Röckel, 2001

belairensis Pin & Leung Tack in Pin, 1989

bellulus Rolán, 1990

bernardinoi (Cossignani, 2014) n. comb.

Rolán & Fernandes, 1990

bocagei Trovão, 1978

Trov ão, 1979

bruguieri Kiener, 1846

bulbus Reeve, 1843

cabraloi (Cossignani, 2014) n. comb.

Ferrario, 1983

cagarralensis (Cossignani, 2014) n. comb.

Monteiro, Fernandes & Rolán, 1995

Rolán, 1990

Rolán, 1990

fernandesi Tenorio, Afonso & Rolán, 2008

fiadeiroi (Tenorio, Afonso, Cunha & Rolán, 2014) n. comb.

filmeri Rolán & Röckel, 2000

flavusalbus Rolán & Röckel, 2000

Rolán & Trovão, 1990

Rolán & Röckel, 2000

furnae Rolán, 1990

Röckel, Rolán & Monteiro, 1980

fuscolineatus G. B. Sowerby III, 1905

gonsaloi (Afonso & Tenorio, 2014) n. comb.

Röckel, Cosel & Burnay, 1980

Lauer, 1993

Hwass in Bruguière, 1792

Kiener, 1847

Rolán, 1990

G. B. Sowerby II, 1858

isabelarum Tenorio & Afonso, 2004

Rolán, 1980

joserochoi (Cossignani, 2014) n. comb.

jourdani da Motta, 1984

kersteni Tenorio, Afonso & Rolán, 2008

lineopunctatus Kaicher, 1977

lobitensis Kaicher, 1977

Röckel, Rolán & Monteiro, 1980

Reeve, 1849

Rolán & Trovão, 1990

Trovão, Rolán & Félix-Alves, 1990

melissae Tenorio, Afonso & Rolán, 2008

Linnaeus, 1758

Rolán & Fernandes, 1990

micropunctatus Rolán & Röckel, 2000

Rolán, 1990

cepasi Trovão, 1975

chytreus Tryon, 1884

claudiae Tenorio & Afonso, 2004

cloveri Walls, 1978

condei (Afonso & Tenorio, 2014) n. comb.

crioulus Tenorio & Afonso, 2004

Reeve, 1849

Reeve, 1843

curralensis Rolán, 1986

Trovão, 1979

Röckel, Rolán & Monteiro, 1980

Trovão, 1979

denizi (Afonso & Tenorio, 2011) n. comb.

Rolán & Fernandes, 1990

desidiosus A. Adams, 1853

diegoi (Cossignani, 2014) n. comb.

Trovão& Rolán, 1986

Monnier & Limpalaër, 2010

echinophilus (Petuch, 1975)

Röckel, Rolán & Monteiro, 1980

Rolán & Trovão, 1990

naranjus Trovão, 1975

Rolán, 1986

negroides Kaicher, 1977

nobrei Trovão, 1975

Pin & Leung Tack, 1989

Röckel, Rolán & Monteiro, 1980

Rolán, Monteiro & Fernandes, 1998

Rolán & Trovão, 1990

roeckeli Rolán, 1980

salletae (Cossignani, 2014) n. comb.

Rolán, 1980

santanaensis (Afonso & Tenorio, 2014) n. comb.

saragasae Rolán, 1986

Rolán, 1990

silviae (Cossignani, 2014) n. comb.

swinneni (Tenorio, Afonso, Cunha & Rolán, 2014) n. comb.

tacomae Boyer & Pelorce, 2009

taslei Kiener, 1850

tenuilineatus Rolán & Röckel, 2001

Rolán & Fernandes, 1990

trencarti Nolf & Verstraeten, 2008

trovaoi Rolán & Röckel, 2000

unifasciatus Kiener, 1850

variegatus Kiener, 1848

vayssierei Pallary, 1906

Gmelin, 1791

verdensis Trovão, 1979

vulcanus Tenorio & Afonso, 2004

wandae (Cossignani, 2014) n. comb.

Röckel, 1987

zebroides Kiener, 1848

zinhoi (Cossignani, 2014) n. comb.

Subgenus

Type species: Hwass in Bruguière, 1792; OD

Included species:

Gmelin, 1791

Hwass in Bruguière, 1792

Röckel & Fischöder, 1985

G. B. Sowerby I, 1833

G. B. Sowerby II, 1858

Subgenus

Type species: Conus amadis Gmelin, 1791; SD, Herrmannsen (1847: 584)

[Synonym: Nataliconus Tucker & Tenorio, 2009; type species: G. B. Sowerby II, 1858; OD; n. syn.]

Included species:

amadis Gmelin, 1791

hamanni Fainzilber & Mienis, 1986

Korn, 1998

kawamurai Habe, 1962

Blumenbach, 1791

milneedwardsi Jousseaume, 1894

G. B. Sowerby II, 1858

royaikeni (S. G. Veldsman, 2010) n. comb.

Subgenus

Type species: Conus lindae Petuch, 1987; OD

[Synonym: Spuriconus Petuch, 2003; type species: Gmelin, 1791; OD]

Included species:

bahamensis Vink & Röckel, 1995

lindae Petuch, 1987

Gmelin, 1791

therriaulti (Petuch, 2013) n. comb.

Subgenus

Type species: Conus millepunctatus Lamarck, 1822 [synonym of Röding, 1798]; SD, Cossmann (1896: 186)

Included species:

Röding, 1798

Subgenus

Type species: Hwass in Bruguière, 1792; OD

[Synonym: Calamiconus Tucker & Tenorio, 2009; type species: Conus Weinkauff, 1875; OD; n. syn.]

Included species:

G. B. Sowerby I, 1834

escondidai Poppe & Tagaro, 2005

Reeve, 1849

A. Adams & Reeve, 1848

garywilsoni Lorenz & Morrison, 2004

kermadecensis Iredale, 1913

Weinkauff, 1875

Hwass in Bruguière, 1792

G. B. Sowerby I, 1833

[Lightfoot], 1786

Quoy & Gaimard, 1834

Subgenus

Type species: Conus ambiguus Reeve, 1844; OD

[Synonym: Gladioconus Tucker & Tenorio, 2009; type species: Broderip, 1833; OD; n. syn.]

Included species:

ambiguus Reeve, 1844

bellocqae van Rossum, 1996

Broderip, 1833

Hwass in Bruguière, 1792

Reeve, 1844

Subgenus

Type species: Conus sculletti Marsh, 1962; OD

Included species:

sculletti Marsh, 1962

Subgenus

Type species: Gmelin, 1791; SD, Cotton (1945: 250)

[Synonyms: Fulgiconus da Motta, 1991; type species: Küster, 1838; OD; n. syn.; Graphiconus da Motta, 1991; type species: Reeve, 1843; OD; n. syn.; Thoraconus da Motta, 1991; type species: Conus exiguus Lamarck, 1810; OD; n. syn. Nimboconus Tucker & Tenorio, 2013; type species: Hwass in Bruguière, 1792; OD; n. syn.]

Included species: Subgenus

alabaster Reeve, 1849

alexandrei (Limpalaër & Monnier, 2012) n. comb.

amplus Röckel & Korn, 1992

E. A. Smith, 1879

angioiorum Röckel & Moolenbeek, 1992

armadillo Shikama, 1971

asiaticus da Motta, 1985

Holten, 1802

balabacensis Filmer, 2012

Crosse, 1867

broderipii Reeve, 1844

cavailloni Fenaux, 1942

cebuensis Wils, 1990

ciderryi da Motta, 1985

Hwass in Bruguière, 1792

collisus Reeve, 1849

G. B. Sowerby II, 1870

leobrerai da Motta & Martin, 1982

lienardi Bernardi & Crosse, 1861

G. B. Sowerby II, 1858

(Bozzetti, 2012) n. comb.

marielae Rehder & Wilson, 1975

martinianus Reeve, 1844

Küster, 1838

moolenbeeki Filmer, 2011

Reeve, 1843

mulderi Fulton, 1936

Reeve, 1843

niederhoeferi (Monnier, Limpalaër & Lorenz, 2012) n. comb.

Hwass in Bruguière, 1792

Gmelin, 1791

oishii (Shikama, 1977)

Reeve, 1844

petergabrieli Lorenz, 2006

pica A. Adams & Reeve, 1848

pretiosus G. Nevill & H. Nevill, 1874

primus Röckel & Korn, 1990

G. B. Sowerby II, 1860

quiquandoni Lorenz & Barbier, 2008

Gmelin, 1791

conspersus Reeve, 1844

dampierensis Coomans & Filmer, 1985

daphne Boivin, 1864

erythraeensis Reeve, 1843

evansi Bondarev, 2001

exiguus Lamarck, 1810

Röckel, 1985

gilvus Reeve, 1849

giorossii Bozzetti, 2005

goudeyi (Monnier & Limpalaër, 2012) n. comb.

habui Lan, 2002

Reeve, 1843

Hwass in Bruguière, 1792

jickelii Weinkauff, 1873

kiicumulus Azuma, 1982

kuroharai (Habe, 1965)

ranonganus da Motta, 1978

Richard & Moolenbeek, 1988

salzmanni G. Raybaudi Massilia & Rolán, 1997

santinii (Monnier & Limpalaër, 2014) n. comb.

sartii Korn, Niederhöfer & Blöcher, 2002

scalptus Reeve, 1843

sculpturatus Röckel & da Motta, 1986

sertacinctus Röckel, 1986

sogodensis (Poppe, Monnier & Tagaro, 2012) n. comb.

solomonensis Delsaerdt, 1992

Linnaeus, 1758

stramineus Lamarck, 1810

straturatus G. B. Sowerby II, 1865

Moolenbeek, Röckel & Bouchet, 2008

vappereaui Monteiro, 2009

wittigi Walls, 1977

yemenensis Bondarev, 1997

zandbergeni Filmer & Moolenbeek, 2010

zapatosensis Röckel, 1987

zebra Lamarck, 1810

Type species: Linnaeus, 1758; SD, Cossmann (1896: 160)

[Synonyms: Heroconus da Motta, 1991; type species: Conus poehlianus G. B. Sowerby III, 1887 [synonym of G. B. Sowerby I, 1833]; OD. Strioconus Thiele, 1929; type species: Linnaeus, 1758; OD. Socioconus da Motta, 1991; type species: G. B. Sowerby I, 1833; OD]

Included species:

Gmelin, 1791

arafurensis (Monnier, Limpalaër & Robin, 2013) n. comb.

atimovatae (Bozzetti, 2012) n. comb.

Linnaeus, 1758

Bernardi, 1861

boutetorum Richard & Rabiller, 2013

Hwass in Bruguière, 1792

Born, 1778

G. B. Sowerby I, 1833

fischoederi Röckel & da Motta, 1983

G. B. Sowerby I, 1841

fulmen Reeve, 1843

Richard & Salvat, 1973

Linnaeus, 1758

Linnaeus, 1758

Hwass in Bruguière, 1792

koukae (Monnier, Limpalaër & Robin, 2013) n. comb.

leobottonii Lorenz, 2006

Linnaeus, 1758

monachus Linnaeus, 1758

morrisoni G. Raybaudi Massilia, 1991

nigromaculatus Röckel & Moolenbeek, 1992

nigropunctatus G. B. Sowerby II, 1858

robini (Limpalaër & Monnier, 2012) n. comb.

rouxi (Monnier, Limpalaër & Robin, 2013) n. comb.

Kiener, 1848

Subgenus

Type species: Conus advertex (Garrard, 1961); OD

Included species:

adami Wils, 1988

advertex (Garrard, 1961)

angasi Tryon, 1884

baeri Röckel & Korn, 1992

bonfigliolii (Bozzetti, 2010) n. comb.

felix Fenzan, 2012

Richard, 1980

trigonus Reeve, 1848

visagenus Kilburn, 1974

wallangra (Garrard, 1961)

Subgenus

Type species: Conus boschorum Moolenbeek & Coomans, 1993; OD

Included species:

boschorum Moolenbeek & Coomans, 1993

korni G. Raybaudi Massilia, 1993

kuiperi Moolenbeek, 2006

levis (Bozzetti, 2012) n. comb.

scalarispira (Bozzetti, 2012) n. comb.

traillii A. Adams, 1855

visseri Delsaerdt, 1990

wallacei (Lorenz & Morrison, 2004)

Subgenus

Type species: Conus carnalis G. B. Sowerby III, 1879; OD

Included species:

carnalis G. B. Sowerby III, 1879

Subgenus

Type species: Conus terryni Tenorio & Poppe, 2004; OD

Included species:

terryni Tenorio & Poppe, 2004

Subgenus

Type species: Hwass in Bruguière, 1792; M.

Included species: Subgenus

Hwass in Bruguière, 1792

Hwass in Bruguière, 1792

vautieri Kiener, 1847

Fischer von Waldheim, 1807

Gmelin, 1791

Type species: Hinds, 1843; OD

Included species:

G. B. Sowerby II, 1873

Hinds, 1843

Remarks: Even though the two species are not in the same group in the molecular phylogeny, and to avoid the creation of a new subgeneric name, we have followed Tucker & Tenorio (2009), who placed C. fergusoni and C. patricius in the same genus, Pyruconus (here ranked as subgenus), based on radula and shell characters.

Subgenus

Type species: G. B. Sowerby III, 1879; OD

Included species:

G. B. Sowerby III, 1879

tuticorinensis Röckel & Korn, 1990

Subgenus

Type species: Linnaeus, 1758; SD, Cossmann (1896: 162)

Included species:

anosyensis Bozzetti, 2008

Linnaeus, 1758

cumingii Reeve, 1848

hyaena Hwass in Bruguière, 1792

Linnaeus, 1758

Hwass in Bruguière, 1792

Hwass in Bruguière, 1792

peli Moolenbeek, 1996

Hwass in Bruguière, 1792

Hwass in Bruguière, 1792

rawaiensis da Motta, 1978

tirardi Röckel & Moolenbeek, 1996

Gmelin, 1791

wilsi Delsaerdt, 1998

Subgenus

Type species: Conus coccineus Gmelin, 1791; OD

Included species:

coccineus Gmelin, 1791

Subgenus

Type species: Conus sanderi Wils & Moolenbeek, 1979; OD

Included species:

carioca Petuch, 1986

ednae (Petuch, 2013) n. comb.

sanderi Wils & Moolenbeek, 1979

Subgenus

Type species: Conus algoensis G. B. Sowerby I, 1834; OD

Included species:

algoensis G. B. Sowerby I, 1834

bairstowi G. B. Sowerby III, 1889

brianhayesi Korn, 2001

gradatulus Weinkauff, 1875

Reeve, 1848

Reeve, 1843

Subgenus

Type species: Röding, 1798; OD

[Synonyms: Kioconus da Motta, 1991; type species: Bernardi, 1853; OD; n. syn. Ongoconus da Motta, 1991; type species: Reeve, 1843; OD; Precedence of Splinoconus (established at the rank of genus) over simultaneously established Kioconus and Ongoconus (established at the rank of subgenus) determined by ICZN (1999) Art. 24.1. Stellaconus Tucker & Tenorio, 2009; type species: Conus malacanus Hwass in Bruguière, 1792; OD; n. syn. Isoconus Tucker & Tenorio, 2013; type species: Kiener, 1847; OD; n. syn. Nitidoconus Tucker & Tenorio, 2013; type species: Reeve, 1844; OD; n. syn.]

Included species:

da Motta, 1986

bayani Jousseaume, 1872

(Röding, 1798)

Reeve, 1844

bondarevi Röckel & G. Raybaudi Massilia, 1992

caillaudii Kiener, 1846

Fulton, 1938

capreolus Röckel, 1985

Kiener, 1847

danilai Röckel & Korn, 1990

Röckel & da Motta, 1983

empressae Lorenz, 2001

Hwass in Bruguière, 1792

gilberti (Bozzetti, 2012) n. comb

gloriakiiensis Kuroda & Itô, 1961

Yoshiba & Koyama, 1984

da Motta & Röckel, 1982

malacanus Hwass in Bruguière, 1792

thevenardensis da Motta, 1987

Walls, 1977

troendlei Moolenbeek, Zandbergen & Bouchet, 2008

typhon Kilburn, 1975

martensi E. A. Smith, 1884

nielsenae Marsh, 1962

Reeve, 1848

olgiatii Bozzetti, 2007

papuensis Coomans & Moolenbeek, 1982

pauperculus G. B. Sowerby I, 1834

purissimus Filmer, 2011

da Motta, 1984

recluzianus Bernardi, 1853

reductaspiralis Walls, 1979

richardsae Röckel & Korn, 1992

G. Raybaudi & da Motta, 1990

ruthae (Monnier & Limpalaër, 2013) n. comb.

Shikama, 1970

Coomans, Moolenbeek & Wils, 1985

sukhadwalai Röckel & da Motta, 1983

urashimanus Kuroda & Itô, 1961

vanvilstereni (Moolenbeek & Zandbergen, 2013) n. comb.

Cernohorsky, 1977

Reeve, 1843

Subgenus

Type species: Conus leucostictus Gmelin, 1791 [= Gmelin, 1791]; SD, Wenz (1943: 1469)

[Synonyms: Rhombus Montfort, 1810 (invalid; junior homonym of Rhombus Walbaum, 1792); type species: Linnaeus, 1758; OD. Cornutoconus Suzuki, 1972; type species: Cornutoconus lamellatus Suzuki, 1972 (= Conus chiangi (Azuma, 1972)); OD. Taranteconus Azuma, 1972; type species: (Azuma, 1972); OD; n. syn. Protoconus da Motta, 1991 (invalid; junior homonym of Protoconus Yu, 1979 and Protoconus Stinchcomb, 1986); type species: Linnaeus, 1767; OD. Seminoleconus Petuch, 2003; type species: †Conus violetae Petuch 1988 (Pliocene); OD. Rhombiconus Tucker & Tenorio, 2009; type species: Linnaeus, 1758; OD; n. syn. Tenorioconus Petuch & Drohlshagen, 2011; type species: Linnaeus, 1767; OD; n. syn.]

Included species:

Broderip, 1833

aurantius Hwass in Bruguière, 1792

G. D. Hanna & Strong, 1949

Wood, 1828

Linnaeus, 1767

(Azuma, 1972)

Hwass in Bruguière, 1792

dominicanus Hwass in Bruguière, 1792

duffyi Petuch, 1992

harlandi Petuch, 1987

granarius Kiener, 1847

Linnaeus, 1758

julieandreae Cargile, 1995

[Lightfoot], 1786

polongimarumai Kosuge, 1980

pseudaurantius Vink & Cosel, 1985

pseudimperialis Moolenbeek, Zandbergen & Bouchet, 2008

Gmelin, 1791

sanguineus Kiener, 1850

solidus Gmelin, 1791

Hwass in Bruguière, 1792

Subgenus

Type species: Linnaeus, 1767; OD

[Synonyms: Vituliconus da Motta, 1991; type species: Conus vitulinus Hwass in Bruguière, 1792 (= Conus planorbis Born, 1778); OD; n. syn. Rolaniconus Tucker & Tenorio, 2009; type species: Linnaeus, 1758; OD; n. syn.]

Included species:

athenae Filmer, 2011

[Lightfoot], 1786

axelrodi Walls, 1978

buniatus (Bozzetti, 2013) n. comb.

Linnaeus, 1767

hanshassi (Lorenz & Barbier, 2012) n. comb.

hivanus Moolenbeek, Zandbergen & Bouchet, 2008

Hwass in Bruguière, 1792

maldivus Hwass in Bruguière, 1792

Iredale, 1929

dedonderi (Goethaels & D. Monsecour, 2013) n. comb.

Hwass in Bruguière, 1792

Hwass in Bruguière, 1792

Born, 1778

splendidulus G. B. Sowerby I, 1833

suduirauti Raybaudi Massilia, 2004

Link, 1807

swainsoni Estival & Cosel, 1986

Linnaeus, 1758

Subgenus

Type species: Born, 1778; OD

Included species: Subgenus

Hwass in Bruguière, 1792

edaphus Dall, 1910

sandwichensis Walls, 1978

Reeve, 1844

Born, 1778

Type species: Linnaeus, 1758; SD, Cotton (1945: 261)

Included species: Subgenus

adamsonii Broderip, 1836

Linnaeus, 1758

Lamarck, 1822

chiapponorum Lorenz, 2004

(H. Adams, 1872)

julii Lienard, 1870

lucasi (Bozzetti, 2010) n. comb.

solangeae Bozzetti, 2004

timorensis Hwass in Bruguière, 1792

vicweei Old, 1973

Type species: Turriconus nakayasui Shikama & Habe, 1968 [synonym of G. B. Sowerby III, 1908]; OD

[Synonyms: Kurodaconus Shikama & Habe, 1968; type species: Kuroda, 1956; OD; n. syn. Mitraconus Tucker & Tenorio, 2013; type species: Hwass in Bruguière, 1792; OD; n. syn.]

Included species:

Lamarck, 1810

Olivera & Biggs, 2010

beatrix Tenorio, Poppe & Tagaro, 2007

Broderip & G. B. Sowerby I, 1830

G. B. Sowerby III, 1908

Röckel & Moolenbeek, 1995

gratacapii Pilsbry, 1904

Moolenbeek, 1986

milesi E. A. Smith, 1887

Olivera & Biggs, 2010

Hwass in Bruguière, 1792

A. Adams, 1855

rizali Olivera & Biggs, 2010

(Kuroda, 1956)

stupella (Kuroda, 1956)

Subgenus

Type species: Linnaeus, 1758; OD

[Synonym: Pseudohermes Tucker & Tenorio 2013; type species: Conus austroviola Röckel & Korn, 1992; OD; n. syn.]

Included species:

artoptus G. B. Sowerby I, 1833

austroviola Röckel & Korn, 1992

berdulinus Veillard, 1972

Crosse, 1858

Reeve, 1849

Lamarck, 1810

Reeve, 1848

Habe & Kosuge, 1970

Crosse, 1858

spiceri Bartsch & Rehder, 1943

Born, 1778

tethys (Petuch & Sargent, 2011) n. comb.

Gmelin, 1791

Linnaeus, 1758

Subgenus

Type species: Linnaeus, 1758; OD

[Synonyms: Dyraspis Iredale, 1949; type species: Péron, 1807; OD, n. syn. Miliariconus Tucker & Tenorio, 2009; type species: Hwass in Bruguière, 1792; OD, n. syn.]

Included species: Incertae Sedis

Reeve, 1843

G. B. Sowerby II, 1857

(Röding, 1798)

Gmelin, 1791

Péron, 1807

Linnaeus, 1758

encaustus Kiener, 1845

Bergh, 1895

lecourtorum (Lorenz, 2011) n. comb.

Hwass in Bruguière, 1792

Hwass in Bruguière, 1792

G. B. Sowerby I, 1833

Genus

Type species: Kenyonia pulcherrima Brazier, 1896; M

Included species:

pulcherrima Brazier, 1896