Literature DB >> 25903813

Signals from the brain induce variation in avian facial shape.

Diane Hu1, Nathan M Young1, Qiuping Xu2, Heather Jamniczky3, Rebecca M Green3, Washington Mio2, Ralph S Marcucio1, Benedikt Hallgrimsson3.   

Abstract

BACKGROUND: How developmental mechanisms generate the phenotypic variation that is the raw material for evolution is largely unknown. Here, we explore whether variation in a conserved signaling axis between the brain and face contributes to differences in morphogenesis of the avian upper jaw. In amniotes, including both mice and avians, signals from the brain establish a signaling center in the ectoderm (the Frontonasal ectodermal zone or "FEZ") that directs outgrowth of the facial primordia.
RESULTS: Here we show that the spatial organization of this signaling center differs among avians, and these correspond to Sonic hedgehog (Shh) expression in the basal forebrain and embryonic facial shape. In ducks this basal forebrain domain is present almost the entire width, while in chickens it is restricted to the midline. When the duck forebrain is unilaterally transplanted into stage matched chicken embryos the face on the treated side resembles that of the donor.
CONCLUSIONS: Combined with previous findings, these results demonstrate that variation in a highly conserved developmental pathway has the potential to contribute to evolutionary differences in avian upper jaw morphology. Developmental Dynamics 244:1133-1143, 2015.
© 2015 Wiley Periodicals, Inc. © 2015 Wiley Periodicals, Inc.

Entities:  

Keywords:  Sonic hedgehog (Shh); craniofacial development; forebrain; frontonasal ectodermal zone (FEZ); variation

Year:  2015        PMID: 25903813      PMCID: PMC4618284          DOI: 10.1002/dvdy.24284

Source DB:  PubMed          Journal:  Dev Dyn        ISSN: 1058-8388            Impact factor:   3.780


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