Literature DB >> 35874066

A checklist of spiders of Nepal (Arachnida; Araneae).

Sanskar Subedi1, Ritu Joshi1, Samir Karki1, Shila Gurung1.   

Abstract

Spiders (order Araneae) have a worldwide distribution. As of June 17, 2022, the World spider catalog, Version 23.0, enlists 50,188 species of spiders from throughout the world. Except for the open sea and air, spiders live in every ecological environment. They prefer densely vegetated environments and are the world's seventh most diverse group of creatures in terms of species diversity. The checklist of spiders of south Asia mentions 222 spider species from Nepal, grouped into 23 families. However, the official database of the Nepalese government only lists 175 species of Nepalese spiders. This checklist is a vital update to the diversity of the spider fauna of Nepal. The goal is to compile a thorough list of all the spiders found in Nepal. We have enlisted 386 different spider species from Nepal, belonging to 135 different genera and 34 different families, after reviewing previous scientific publications, computer databases of the Nepalese government, and the World spider catalog, Version 23.0. It adds a total of 211 new spiders to Nepal's biodiversity database. Linyphiidae and Salticidae spiders dominate accounting for 27.46 percent and 17.36 percent of total species, respectively, on the checklist. Corrections to previous misidentifications are also included in this checklist, as well as taxonomy revisions. Synonymous species are sorted out to avoid recurrence. The trends in Nepalese spider discovery and dispersion have also been studied.
© 2022 The Author(s).

Entities:  

Keywords:  Endemic; Nepalese; Species; Spider

Year:  2022        PMID: 35874066      PMCID: PMC9304739          DOI: 10.1016/j.heliyon.2022.e09927

Source DB:  PubMed          Journal:  Heliyon        ISSN: 2405-8440


Introduction

Spiders are found all over the planet and have mastered all natural situations, excluding the open sea and the air [1]. The World spider catalog, Version 23.0, has 50,188 species of spiders from all across the world as of June 17,2022 [2]. The amount of extant species of spiders has been estimated to be over 170,000 [3] although it could be much higher. Spiders are without a doubt the largest taxonomic group that is totally made up of predators. Part of their success can be attributed to their skill to colonize almost all terrestrial habitats, from marine intertidal zones [4]to high altitude areas, as evidenced by the champion Euophrys omnisuperstes [5] which can be found at 6700 m altitude in the Himalayas [6]. Spider study is growing, thanks to new taxonomic discoveries, neuronal properties of spider venom, and the versatility of spider silk. Brignoli [7, 8, 9, 10], Wunderlich [11, 12, 13], Ono [14, 15, 16, 17, 18, 19], Zabka [20, 21, 22, 23], Bohdanowicz [24, 25], Thapa [26], and Buchar [27, 28, 29] were among the significant contributors to the study of Nepalese spiders in the mid-twentieth century [30]. Several studies have been based on Prof. J. Martens' collections from the Himalayan expeditions. Buchar introduced 7 new Lycosid spiders from Nepal in 1976, 1978, and 1984. Bohdanowicz (1979, 1987) described new Synagelides spiders from Nepal in his studies. Ono (1979, 1980, and 1985) used numerous thomisid spiders from Nepal in his research. Other researches by Ono (1983, 2006) and Jocqué (1992) describe endemic Zodariidae spiders from Nepal. Nishikawa's study from 1980 introduced 2 new Agelena spiders from Nepal's Khumbu region. Zabka discovered 11 distinct salticid spiders in Nepal during his research. Other 2 spider species belonging to the genus Suffasia were also discovered from eastern Nepal [31]. Two hersilid spiders from Nepal were introduced by Baehr & Baehr in 1993. There were 144 species of spiders in Nepal, according Thapa’s book; “Enumeration of Spiders of Nepal” [26]. Jastrzebski [32, 33, 34, 35, 36, 37, 38, 39] cites 15 different salticid spiders from Nepal in his various researches. A study on rice field spiders was also conducted in Nepal's Chitwan district [40]. Jäger [41, 42, 43, 44, 45] describes the family Sparassidae, which includes 33 species from Nepal. Wang discovered 7 new Himalcoelotes species in his research [46]. There are 37 new spiders of the genus Draconarius in the revised list of ceolotine spiders from Nepal [47]. Different Studies by Tanasevitch [48, 49, 50, 51, 52, 53, 54, 55, 56] Tanasevitch and Saaristo [57], and Wunderlich [11, 12, 13] enlist many linyphid spiders from Nepal. The “Nepal biodiversity resource book” featured a checklist of spiders of Nepal (Annex 2.1) with 175 species of spiders belonging to 22 distinct families, based on data from earlier publications [26, 40, 58]. Similarly, the checklist of south Asian spiders identified 222 spider species belonging to 79 genera. 176 of those species were only found in Nepal [30]. Wang and Zhu discovered 3 new species of the genus Himalmartensus in Nepal [59]. 38 different Nepalese spiders of the genus Draconarius are described in a study based on J. Marten's collection from Himalayan trips [47]. Four new Camptoscaphiella species were discovered in Nepal by Baehr and Ubick in 2010. Platnick et al. found 3 new species of Brignolia in Nepal [60]. Huber described 2 species of genus Pholcus from Nepal in 2011. Jager found a new species of genus Ctenus in 2012. Four species of the genus Himalayana from Nepal are included in Grismado’s study [61]. Nepal's official spider species count is 175 as published in “Nepal's Sixth National Report to The Convention on Biological Diversity” (2018), released by the Ministry of Forest and Environment, Government of Nepal [62]. Furthermore, the study by Xu et al. documented 5 new species of the genus Leclercera from Nepal [63]. Chang and Li added another Leclercera spider to the list [64]. For the first time in Nepal, Shrestha and Dorr announced the finding of the genus Lactrodectus in 2020 [65]. Nine new Asian salticid spider species were discovered during a field study in Nepal's Chitwan National Park [66]. A new genus, Himalafurca, is described in a recent study including two species from Nepal [52]. A different study reports 7 new Pimoa species from Nepal [67]. Furthermore research on Nepalese spiders has been ongoing since last inventory, and this update to the Nepalese spider checklist is critical. The purpose of this article is to compile a list of all the spider species discovered in Nepal, update their taxonomy and reveal the current state of spider research in Nepal.

Study area

Nepal, a southeast Asian country that makes up around 0.1 percent of the world's landmass, is home to 118 different ecosystems [68]. It is a biogeographical transition zone between the Paleotropic and Palaearctic biogeographical realms in the south and the Palaearctic biogeographical realms in the north [69]. Nepal is divided into three ecological regions; Mountain, hills and terai. The climate ranges from tropical to arctic in a short distance of 180 km [68]. Extreme height fluctuation (70–8848.86 m) and precipitation (up to 5500 mm yearly) [70], divergent temperatures, aspect, and humidity form a complex mosaic of ecosystems and habitat ranging from tropical forest through alpine highlands in Nepal [68]. The classification by Stainton identified 35 various forest types in Nepal [71]. Nepal is home to a vast range of flora and animals. According to Nepal's sixth national report to the Convention on Biological Diversity (2018), the country is home to over 13,000 species of flora and over 17,000 species of fauna [62]. Politically, Nepal is divided into 77 districts and 7 federal provinces (see Figure 1).
Figure 1

Map of Nepal; Geographical and political. The small rectangular box displays Nepal's geographic location inside Asia. Nepal is a landlocked nation in Southeast Asia that shares borders with both China and India. Lying below is the enlarged political map of Nepal. There, it makes reference of Nepal's federal provinces and districts.

Map of Nepal; Geographical and political. The small rectangular box displays Nepal's geographic location inside Asia. Nepal is a landlocked nation in Southeast Asia that shares borders with both China and India. Lying below is the enlarged political map of Nepal. There, it makes reference of Nepal's federal provinces and districts.

Materials and methods

Data collection

This checklist was created using information from previous publications in international journals, books, a computer database, and scientific records from “The World Spider Catalog” (Version-23.0). Using Google Scholar's all-in-title feature, we looked for literature discussing Nepalese spiders using keywords with Boolean operators “Spider” OR “Spiders”, “Nepal”. There were a total of 21 search results, with 14 articles and 7 citations. To find the papers listed, these citations were copied and searched in full scale in Google Chrome. Then, using the terms “Checklist of Nepalese spiders,” “List of Nepalese spiders,” “Nepalese spiders,” “Nepali spiders,” “Spiders from Nepal,” “Spiders in Nepal,” and “Spiders of Nepal”, multiple full-scale Google searches for more literature were conducted. Digital data on Nepalese spiders was collected from a government database. The World spider catalog was browsed through in order to include every article on Nepalese spiders in this study. For the last inspection, snowball referencing was used. This resulted in 94 published articles about Nepalese spiders in total. In addition, other publications, reports, and catalogs were scrutinized for accuracy.

Analysis

We investigated the trend in discovery of spiders from Nepal through measure of Karl Pearson’s coefficient of correlation [72]. Also, we compared our findings and examined the spider biodiversity within and outside Nepalese territory.

Representation

The findings were then organized as a checklist in a tabular fashion and classified according to the spider species' taxonomic family. The species' location, global distribution and bibliography are also included. The spider taxonomy is based on Version 23.0 of the World Spider Catalog (2022). The information is also displayed using descriptive statistics such as graphs and charts. The map was plotted in ArcMap version 10.4.1.

Results

We found 386 different spider species in Nepal, divided into 135 genera and 34 families (see Tables 1 and 2). Out of these, 251 species are endemic. This is a more than twofold increase in Nepal's spider biodiversity database. Linyphiidae and Salticidae spiders dominate accounting for 27.46 percent and 17.36 percent of total species, respectively (Figure 2, Figure 3). Nepalese spider study appears to be centered in the mountain ecological zone (63%) followed by the hills (31%) and the Terai region (6%) (Figure 4). The bulk of enlisted spiders are found in Province 1, while Sudurpaschim and Madhesh Provinces each have only one spider described (Figure 5). About 94 different articles on Nepalese spiders have been published since 1910. Karl Pearson’s correlation analysis revealed a slightly positive trend in discovery of spiders from Nepal (r =0.228). Maximum discoveries (44 species) have been been made in 2021 (Figure 6).
Table 1

Nepalese spider genera and species by family.

SNFamilyNo.of GeneraNo. of SpeciesNo. of endemic speciesGuild Structures [75]
1.Agelenidae45046Sheet web builders
2.Amaurobiidae133Sheet web builders
3.Anapidae122Orb weavers
4.Araneidae490Orb weavers
5.Ctenidae111Ground/other hunters
6.Deinopidae110Ambush hunters
7.Dictynidae110Space web builder
8.Eresidae110Sheet web builders
9.Gnaphosidae241Ground hunters
10.Hahniidae222Sheet web builders
11.Hersilidae131Sensing web builder
12.Linyphiidae4110682Web builders/wandering
13.Lycosidae7184Ground hunters
14.Mysmenidae111Space web builder
15.Nesticidae110Space web builder
16.Oonopidae51612Ground hunters
17.Oxyopidae130Stalkers
18.Pholcidae120Space web builder
19.Pimoidae199Sheet web builders
20.Pisauridae110Ambush hunter
21.Psechridae231Sheet web builders
22.Psilodercidae177-
23.Salticidae306727Stalkers
24.Scytodidae110Hunters
25.Selenopidae110Ambush hunters
26.Sparassidae33332Wandering spiders
27.Symphytognathidae111Orb weavers
28.Tetrablemmidae222Sheet web builders
29.Tetragnathidae220Orb weavers
30.Theraphosidae111Sensing web builder
31.Theridiidae221Space web builder
32.Thomisidae7237Ambush hunters
33.Titanoecidae111Space web builder
34.Zodariidae387Specialists
Figure 2

(A) Total species per family of Nepalese spiders. (B) Percentage of each family in Nepal’s endemic species.

Figure 3

No of genera in each family of Nepalese spiders.

Figure 4

Nepalese spider species by ecological regions.

Summary of the spider fauna of Nepal.

Number of Families: 34.

Number of Genera: 135.

Number of Species: 386.

Number of Endemic Species: 251.

Figure 5

Nepalese spider species by federal provinces.

Figure 6

Trend of spider discoveries in Nepal.

Nepalese spider genera and species by family. Spiders of Nepal and their distribution by family (A) Total species per family of Nepalese spiders. (B) Percentage of each family in Nepal’s endemic species. No of genera in each family of Nepalese spiders. Nepalese spider species by ecological regions. Summary of the spider fauna of Nepal. Number of Families: 34. Number of Genera: 135. Number of Species: 386. Number of Endemic Species: 251. Nepalese spider species by federal provinces. Trend of spider discoveries in Nepal. This checklist contains taxonomic upgrades as well as corrections to past misidentifications. To avoid recurrence, synonymous species are sorted. Seven Tetrablemmidae species (Tetrablemma elongata, Tetrablemma laboriosa, Tetrablemma mandibulata, Tetrablemma maxillosa, Tetrablemma phulchoki, Tetrablemma straminea, Tetrablemma virescens, Theridiosoma sp) and two Synagelides species (Synagelides wangdicus and Synagelides wuermlii) enlisted earlier [30, 58] are omitted in this checklist due to lack of published references or collected specimens. Similarly, some enlistments from earlier lists are excluded due to unidentified specific trait. The checklist has also taken into account previous misidentifications, synonyms, and taxonomic transfers of several spider species.

Discussion

With 386 species of spiders, Nepal has about five times more species-to-area ratio (0.00262) than its neighbouring countries; China (0.000546) and India (0.000512) having 5249 and 1686 spider species each [73, 74]. It accounts for 16.79% of spiders of South Asia (2299 species) and 0.77% of total spiders in the world [2, 30]. The family Linyphiidae dominates the spider inventory of Nepal, although Salticids lead the Chinese and Indian catalogues. With 63% of total spiders enlisted, maximum expeditions have been focused on Mountain ecological region of Nepal. In Nepalese spiders, there are conspicuous Himalayan radiations. Deeply separated valleys and a plethora of mountain ranges preventing ground-dwelling arthropods from spreading quickly from one valley chain to the next, has resulted in the evolution of several species [47]. The diversity of coelotine spiders in Nepal astounded the authors; Wang and Martens [47]. Local species of genera Draconarius, Pseudopoda, and Bhutaniella have particularly striking traits [43, 47]. The existence of Euophrys omnisuperstes amid snow and stony debris at a height of 6700 m above sea level is intriguing. The study of Nepal's endemic Himalayan spider species has got a good attention, but the lush lower vegetation has been overlooked. Vast swaths of biologically significant land have remained mostly unexplored. Out of 77 districts of Nepal, 39 have not been explored a bit for spider diversity. Since 1910, there have been only 94 publications on Nepalese spiders. There is a weak positive Karl Pearson’s coefficient of correlation (r = 0.228) between years and new spiders discovered in Nepal. A simple keyword search {allintitle: spider “Country name”} yields about 40 times less results on google scholar for Nepal than those for China and India. These clearly indicate a significant research gap. Scholars from around the world appear to be curious but Nepalese have played a modest role in spider studies. Also, a 100% research focus has been on baseline surveys. With growing global interests on spider webs, ecology and venom, other thematic areas should be covered as well. Nepal thus seems a promising land for spider diversity. Further explorations might significantly boost global spider inventory. The authors thus invite and encourage researchers from all around the world to investigate Nepalese spiders.

Declarations

Author contribution statement

All authors listed have significantly contributed to the development and the writing of this article.

Funding statement

This research did not receive any specific grant from funding agencies in the public, commercial, or not-for-profit sectors.

Data availability statement

Data included in article/supplementary material/referenced in article.

Declaration of interests statement

The authors declare no conflict of interest.

Additional information

No additional information is available for this paper.
Table 2

Spiders of Nepal and their distribution by family

S.NGeneraSpecies with BibliographyLocation (District/Province)Distribution
I. FAMILY AGELENIDAE C.L. Koch, 1837
1.Agelena Walekenaer, 1837Agelena lukla [76]Solukhumbu/ Province 1Nepal and China
Agelena Sherpa [76]Endemic
2.Draconarius Ovtchinnikov, 1999Draconarius beloniformis [47]Mustang/ Gandaki ProvinceEndemic
Draconarius bifarius [47]Terhathum/ Province 1Endemic
Draconarius brevikarenos [47]Ilam/ Province 1Endemic
Draconarius capitellus [47]Myagdi/ Gandaki ProvinceEndemic
Draconarius communis [47]Parbat/ Gandaki ProvinceEndemic
Draconarius condocephalus [47]Taplejung/ Province 1Endemic
Draconarius confusus [47]Dolakha/ Bagmati ProvinceEndemic
Draconarius contiguus [47]Dolpa/ Karnali ProvinceEndemic
Draconarius cylindratus [47]Taplejung/ Province 1Endemic
Draconarius dapaensis [47]Mustang/ Gandaki ProvinceEndemic
Draconarius distinctus [47]Panchthar / Province 1Endemic
Draconarius dorsicephalus [47]Dolakha/Bagmati ProvinceEndemic
Draconarius gorkhaensis [47]Gorkha/ Gandaki ProvinceEndemic
Draconarius gurkha [47]Solukhumbu/ Province 1Endemic
Draconarius latiforus [47]Sankhuwasabha / Province 1Endemic
Draconarius meganiger [47]Myagdi/ Gandaki ProvinceEndemic
Draconarius microcoelotes [47]Endemic
Draconarius panchtharensis [47]Panchthar/ Province 1Endemic
Draconarius paraepisomos [47]Mustang/ Gandaki ProvinceEndemic
Draconarius phulchokiensis [47]Lalitpur/ Bagmati ProvinceEndemic
Draconarius pseudogurkha [47]Solukhumbu/ Province 1Endemic
Draconarius pseudomeganiger[47]Lalitpur/ Bagmati ProvinceEndemic
Draconarius sacculus [47]Taplejung/ Province 1Endemic
Draconarius schawalleri [47]Gorkha/ Gandaki ProvinceEndemic
Draconarius semicirculus [47]Dolakha/ Bagmati ProvinceEndemic
Draconarius seorsus [47]Dolakha/ Bagmati ProvinceEndemic
Draconarius simplicifolis [47]Myagdi/ Gandaki ProvinceEndemic
Draconarius spinosus [47]Mustang/ Gandaki ProvinceEndemic
Draconarius subconfusus [47]Panchthar/ Province 1Endemic
Draconarius subepisomos [47]Solukhumbu/ Province 1Endemic
Draconarius subrotundus [47]Sankhuwasabha / Province 1Endemic
Draconarius taplejungensis [47]Taplejung/ Province 1Endemic
Draconarius testudinatus [47]Taplejung/ Province 1Endemic
Draconarius tinjuraensis [47]Terhathum/ Province 1Endemic
Draconarius tritos [47]Taplejung/ Province 1Endemic
Draconarius volutobursarius[47]Dolpa/ Karnali ProvinceEndemic
Draconarius wuermlii [47]Taplejung/ Province 1Nepal and Bhutan
Draconarius yadongensis [47]Nepal and China
3.Himalcoelotes Wang ,2002Himalcoelotes aequoreus [46]Mustang/ Gandaki ProvinceEndemic
Himalcoelotes bursarius [46]Sindhupalchowk/ Bagmati P.Endemic
Himalcoelotes diatropos [46]Rasuwa/ Bagmati P.Endemic
Himalcoelotes gyirongensis[46]Parbat/ Gandaki P.Nepal and China
Himalcoelotes martensi [46]Kaski/ Gandaki ProvinceEndemic
Himalcoelotes pirum [46]Manang/ Gandaki P.Endemic
Himalcoelotes sherpa [46]Solukhumbu/ Province 1Endemic
Himalcoelotes subsherpa [46]Ramechhap/ Bagmati P.Endemic
Himalcoelotes syntomos [46]Lalitpur/ Bagmati P.Endemic
4.
TegenariaLatreille, 1804
Tegenaria lunakensis[77]
Taplejung/ Province 1
Endemic
II. FAMILY AMAUROBIIDAE Thorell, 1870
1.
HimalmartensusWang and Zhu, 2008
Himalmartensus ausobskyi [59]Dolakha/ Bagmati P.Endemic
Himalmartensus martensi [59]Kathmandu/ Bagmati P.Endemic
Himalmartensus nepalensis [59]
Rasuwa/ Bagmati P.
Endemic
III. FAMILY ANAPIDAE Simon, 1895
1.
MetanapisBrignoli, 1981
Metanapis montisemodi [7]Myagdi/ Gandaki P.Endemic
Metanapis tectimundi [7]
Rasuwa/ Bagmati P.
Endemic
IV. FAMILY ARANEIDAE Clerck, 1757
1.GasteracanthaSundevall, 1833Gasteracantha kuhli [78]Banke/ Lumbini P.Nepal, India, Japan
Gasteracantha sanguinolenta [78] [[7878]]Nepal, Africa, Yemen
2.HypsosingaAusserer, 1871Hypsosinga pygmaea [58]NepalNepal, North America
Hypsosinga sanguinea [58]NepalNepal, Europe, North Africa
3.MacracanthaSimon, 1864Macracantha hasselti [78]∗transferred from genus GasteracanthaBanke/ Lumbini P.Nepal, Pakistan, India, China
4.
NeosconaSimon, 1864
Neoscona arabesca [58]NepalNepal, Israel, India
Neoscona nautical [58]NepalAsia and pacific islands
Neoscona scylla [58]NepalNepal, Russia, China, Korea
Neoscona theisi [58]
Nepal
Nepal, Australia, India
V. FAMILY CTENIDAE Keyserling, 1877
1.
CtenusWalckenaer, 1805
Ctenus martensi [42]
Parbat/ Gandaki Province
Endemic
VI. FAMILY DEINOPIDAE C.L. Koch, 1850
1
AsianopisLin and Li, 2020
Asianopis goalparaensis [58]∗Transferred from genus Deinopis
Nepal
Nepal and India
VII. FAMILY DICTYNIDAE O. Pickard-Cambridge, 1871
1
NigmaLehtinen, 1967
Nigma shiprai [58]∗Transferred from genus Dictyna
Nepal
Nepal and India
VIII. FAMILY ERESIDAE C.L. Koch, 1851
1.
StegodyphusSimon, 1873
Stegodyphus sarasinorum [2]
Nepal
India, Sri Lanka, Nepal (Endemic to South Asia)
IX. FAMILY GNAPHOSIDAE Pocock, 1898
1.DrassodesWestring, 1851Drassodes lutescens [79]∗synonym: Phaeocedus mosambaensisSolukhumbu/ Province 1Nepal, Ukraine, Caucasus
Drassodes phagduaensis [77]Taplejung/ Province 1Endemic
2.
GnaphosaLatreille, 1804
Gnaphosa mandschurica [80]Mustang/ Gandaki PMongolia, China, Russia
Gnaphosa moerens [80]
Dolpa/ Karnali Province
China and Nepal
X. FAMILY HAHNIIDAE Bertkau, 1878
1.HahniaC.L Koch, 1841Hahnia alini [77]Taplejung/ Province 1Endemic
2.
NeoantisteaGertsch, 1934
Neoantistea janetscheki [81]
Solukhumbu/ Province 1
Endemic
XI. FAMILY HERSILIIDAE Thorell, 1870
1.
HersiliaAudouin, 1826
Hersilia martensi [82]Gorkha/ Gandaki ProvinceNepal and Thailand
Hersilia nepalensis [82]Endemic
Hersilia savignyi [82]
Dhading/ Bagmati P.
Nepal, India, Pakistan
XII. FAMILY LINYPHIIDAE Blackwall, 1859
1.AgynetaHull, 1911Agyneta bueko [11]Dolpa/ Karnali ProvinceEndemic
Agynetahimalaya [56]Panchthar, Province 1Endemic
Agyneta jiriensis [11]Dolakha/ Province 1Endemic
Agyneta pakistanica [52]Dailekh/ Karnali ProvinceNepal and Pakistan
Agyneta pseudofuscipalpis [11]Dolpa/ Karnali ProvinceEndemic
Agyneta yulungiensis [11]Endemic
2.AnguliphantesSaaristo &Tanasevitch, 1966Anguliphantes nepalensis [50]∗Transferred from genus LepthyphantesMyagdi/ Gandaki P.Nepal, Pakistan, India
3.AscetophantesTanasevitch & Saaristo, 2006Ascetophantes asceticus [48]∗Transferred from genus LepthyphantesIlam/ Province 1Endemic
4.AsthenargusSimon and Fage, 1922Asthenargus thaleri [11]Baglung/ Gandaki ProvinceEndemic
5.BathyphantesMenge, 1866Bathyphantes paracymbialis [52]Sankhuwasabha /Province 1Nepal, China, Laos, Myanmar, Thailand
6.CaviphantesOi, 1960Caviphantes pseudosaxetorum[52]Nepal, China , Japan
7.ClaviphantesTanasevitch & Saaristo, 2006Claviphantes bifurcatoides [52]∗Transferred from genus LepthyphantesEndemic
Claviphantes bifurcatus [48]∗Transferred from genus LepthyphantesParbat/ Gandaki ProvinceEndemic
8.ErigoneAudouin, 1826Erigone acuta [49]Sankhuwasabha /Province 1Endemic
Erigone atra [11]Mustang/ Gandaki P.Nepal, China, Russia
Erigone nepalensis [11]Sindhupalchowk/ BagmatiEndemic
Erigone prominens [52]Synonym: Erigone ouraniaDailekh/ Karnali ProvinceNepal to China, Australia
9.FistulaphantesTanasevitch & Saaristo, 2006Fistulaphantes canalis [52]Sankhuwasabha /Province 1Endemic
10.GnathoriumKarsch, 1881Gnathorium gibberum [52]Taplejung/ Province 1Nepal, China, Korea, Japan, Russia
11.GongylidiellumSimon, 1884Gongylidiellum kathmanduense[11]Baglung/ Gandaki P.Endemic
Gongylidiellum koshi [49]Sankhuwasabha/Province1Endemic
Gongylidiellum nepalense [50]Mustang/ Gandaki P.Nepal and India
12.HaloratesHull, 1911Halorates crassipalpis [52]previously misidentified as Collinsia japonicaMyagdi/ Gandaki ProvinceNepal and Pakistan
13.HelsdingeniaSaaristo and Tanasevitch, 2003Helsdingenia ceylonica [83]Lalitpur/ Bagmati ProvinceNepal, Sri Lanka (Endemic to South Asia)
14.HeterolinyphiaWunderlich, 1973Heterolinyphia tarakotensis [12]Dolpa/ Karnali ProvinceIndia and Nepal (Endemic to South Asia)
15.HilairaSimon, 1884Hilaira dapaensis [52]Taplejung/ Province 1Endemic
16.HimalafurcaTanasevitch, 2021Himalafurca martensi [52]Sankhuwasabha / Province 1Endemic
Himalafurca schawalleri [52]Taplejung/ Province 1Endemic
17.HimalaphantesTanasevitch, 1992Himalaphantes grandiculus [52]Panchthar/ Province 1Endemic
Himalaphantes magnus [48]Rasuwa/ Bagmati ProvinceEndemic
Himalaphantes martensi [48]Mustang, Gandaki ProvinceIndia and Nepal
18.HubertellaPlatnick, 1989Hubertella montana [52]Sindhupalchowk/ Bagmati P.Endemic
Hubertella orientalis[58]NepalEndemic
Hubertella thankurensis [11]Baglung/ Gandaki ProvinceEndemic
19.IndophantesSaaristo and Tanasevitch, 2003Indophantes agamus [57]Panchthar/ Province 1Endemic
Indophantes digitulus [48]Mustang/ Gandaki ProvinceNepal, Pakistan and India
20.LinyphiaLatreille, 1804Linyphia nepalensis [11]Baglung/ Gandaki ProvinceEndemic
21.MartensinusWunderlich, 1973Martensinus annulatus [12]Baglung/ Gandaki ProvinceEndemic
Martensinus micronetiformis [11]Mustang/ Gandaki ProvinceEndemic
22.MegalepthyphantesWunderlich, 1994Megalepthyphantesnebulosoides [48]∗transferred from genus LepthyphantesMustang/ Gandaki ProvinceCentral Asia, Iran
23.MughiphantesSaaristo & Tanasevitch, 1999Mughiphantes alticola [48]∗transferred from genus LepthyphantesMustang/ Gandaki ProvinceEndemic
Mughiphantes anachoretus [48]∗transferred from genus LepthyphantesEndemic
Mughiphantes ancoriformis [52]∗transferred from genus LepthyphantesTaplejung/ Province 1Endemic
Mughiphantes bicornis [57]Endemic
Mughiphantes cuspidatus [57]Endemic
Mughiphantes falxus [57]Endemic
Mughiphantes faustus [48]∗transferred from genus LepthyphantesRamechhap/ Bagmati ProvinceEndemic
Mughiphantes inermus [57]Sankhuwasabha / Province 1Endemic
Mughiphantes longiproper [57]Taplejung/ Province 1Endemic
Mughiphantes numilionis [48]∗transferred from genus LepthyphantesMustang/ Gandaki ProvinceEndemic
Mughiphantes occultus [48]∗transferred from genus LepthyphantesSolukhumbu/ Province 1Endemic
Mughiphantes restrictus [57]Sankhuwasabha / Province 1Endemic
Mughiphantes rotundatus [48]∗transferred from genus LepthyphantesMyagdi/ Gandaki P.Endemic
Mughiphantes setifer [48]∗transferred from genus LepthyphantesDolpa/ Karnali ProvinceEndemic
Mughiphantes setosus [57]Taplejung/ Province 1Endemic
Mughiphantes sherpa [48]Dolpa/ Karnali ProvinceEndemic
Mughiphantes yeti [48]∗ transferred from genus LepthyphantesSolukhumbu/ Province 1Endemic
24.NasoonaLocket, 1982Nasoona asocialis [52]∗ transferred from genus Oedothorax∗Previously published as Gorbothorax ungibbusKathmandu/ Bagmati ProvinceNepal, China, India
Nasoona comata [53]∗ transferred from genus GorbothoraxPanchthar/ Province 1Endemic
Nasoona conica [53]∗ transferred from genus GorbothoraxTaplejung/ Province 1Endemic
Nasoona setifera [53]∗ transferred from genus GorbothoraxTerathum/ Province 1Endemic
Nasoona wunderlichi [13]∗ transferred from genus GorbothoraxDolakha/ Bagmati ProvinceEndemic
25.NematogmusSimon, 1886Nematogmus dentimanus [52]Sankhuwasabha / Province 1Nepal, Srilanka, Malaysia, Indonesia
26.NerieneBlackwall, 1833Neriene oidedicata [52]∗ transferred from genus LinyphiaPanchthar/ Province 1Nepal, China, Russia, Korea, Japan
27.NesionetaMillidge, 1991Nesioneta muriensis [11]∗transferred from genus AgynetaMyagdi/ Gandaki ProvinceEndemic
28.OedothoraxBertkau, in Förster&Bertkau, 1883Oedothorax angelus [55]Panchthar/ Province 1Endemic
Oedothorax annulatus [13]Dolakha/ Bagmati ProvinceEndemic
Oedothorax assuetus [55]Kathmandu/ Bagmati P.Endemic
Oedothorax clypeellum [55]Endemic
Oedothorax coronatus [55]Ilam/ Province 1Endemic
Oedothorax cruciferoides [54]Endemic
Oedothorax dismodicoides [52]Myagdi/ Gandaki ProvinceEndemic
Oedothorax elongatus [52]Kavre/ Bagmati ProvinceEndemic
Oedothorax falcifer [55]Ilam/ Province 1Endemic
Oedothorax hirsutus [13]Lalitpur/ Bagmati ProvinceEndemic
Oedothorax kathmandu [54]Kathmandu/ Province 1Endemic
Oedothorax lineatus [13]Dolakha/ Bagmati ProvinceEndemic
Oedothorax lucidus [13]Endemic
Oedothorax malearmatus [55]Panchthar/ Province 1Endemic
Oedothorax mangsima [54]Sankhuwasabha / Province 1Endemic
Oedothorax modestus [55]Panchthar/ Province 1Endemic
Oedothorax savigniformis [55]Taplejung/ Province 1Endemic
Oedothorax sexoculatus [13]Dolakha/ Bagmati ProvinceEndemic
Oedothorax sexoculorum [55]Terhathum/ Province 1Endemic
Oedothorax simplicithorax [55]Ilam/ Province 1Endemic
Oedothorax tholusus [55]Kaski/ Gandaki ProvinceEndemic
Oedothorax triceps [54]Sindhupalchowk/ Bagmati P.Endemic
Oedothorax unicolor [13]Lalitpur/ Bagmati ProvinceEndemic
29.OiaWunderlich, 1973Oia Kathmandu [52]Sindhupalchowk/ Bagmati P.Endemic
Oia sororia [52]Myagdi/ Gandaki ProvinceNepal and India
30.PalliduphantesSaaristo & Tanasevitch, 2001Palliduphantes theosophicus [48]∗ transferred from genus LepthyphantesLalitpur/ Bagmati ProvinceEndemic
31.ParagongylidiellumWunderlich, 1973Paragongylidiellum caliginosum [52]Mustang/ Gandaki P.Nepal and India
32.ParbatthoraxTanasevitch, 2019Parbatthorax unicornis [51]Parbat/ Gandaki ProvinceEndemic
33.PiniphantesSaaristo & Tanasevitch, 1996Piniphantes himalayensis [48]Mustang/ Gandaki ProvinceNepal and Pakistan
34.PorrhommaSimon, 1884Porrhomma marphaense [11]∗nomen dubiumMustang/ Gandaki ProvinceEndemic
35.SalocaSimon, 1926Saloca gorapaniensis [11]Mustang/ Gandaki ProvinceEndemic
Saloca khumbuensis [11]Solukhumbu/ Bagmati P.Endemic
36.ScotargusSimon, 1913Scotargus pilosus [11]Mustang/ Gandaki P.Nepal, Europe, Algeria, Russia, Central Asia
37.SpiralophantesTanasevitch & Saaristo, 2006Spiralophantes mirabilis [57]Sankhuwasabha / Province 1Endemic
38.TapinocybaSimon, 1884Tapinocyba montivaga [52]Sankhuwasabha / Province 1Endemic
Tapinocyba altimontanus [57]Endemic
39.TenuiphantesSaaristo &Tanasevitch, 1996Tenuiphantes crassus [57]Taplejung / Province 1Endemic
Tenuiphantes plumipes [48]Gorkha/ Gandaki ProvinceEndemic
40.TisoSimon, 1884Tiso aestivus [52]Taplejung / Province 1Nepal, Canada, Japan
Tiso indianus [52]Nepal and India
41.
Walckenaeria Blackwall, 1833
Walckenaeria martensi [50]synonym: Walckenaeria nepalensis
Solukhumbu, Province 1
Nepal and India
XIII. FAMILY LYCOSIDAE Sundevall, 1833
1.AcantholycosaDahl, 1908Acantholycosa baltoroi [29]Solukhumbu/ Province 1Nepal, India, China
2.ArctosaC.L. Koch, 1847Arctosa janetscheki [27]Kavre/ Bagmati ProvinceEndemic
Arctosa raptor [84]Dolpa/ Karnali ProvinceRussia, Nepal, USA, Canada
3.HippasaSimon, 1885Hippasa greenalliae [58]NepalNepal, India, Sri Lanka
4.HylyphantesSimon, 1884∗senior synonym of genus ErigonidiumHylyphantes graminicola [58]NepalNepal, Europe, Russia, China
5.LycosaGravely, 1924Lycosa kempi [27]Dolakha/ Bagmati ProvinceNepal, Pakistan, India, China
6.PardosaC.L. Koch, 1847Pardosa bifasciata [27]previously published as Pardosa thaleriSolukhumbu/ Province 1Nepal, Europe, Turkey, Russia, China
Pardosa birmanica [27]Solukhumbu/ Province 1Nepal, Myanmar
Pardosa fletcheri [29]Myagdi/ Gandaki P.Nepal, Pakistan, India
Pardosa martensi [29]Dolpa/ Karnali ProvinceEndemic
Pardosa mongolica [28]Nepal, Russia, Mongolia,China
Pardosa orealis [28]Endemic
Pardosa pseudoannulata [58]∗Synonym: Lycosa pseudoannulataNepalNepal, Pakistan, China, India, Bhutan, Japan,Indonesia
Pardosa pusiola [2]NepalNepal, Bhutan, India
Pardosa sumatrana [27]Solukhumbu/ Province 1Nepal, Bhutan, India
Pardosa sutherlandi [29]Parbat/ Gandaki P.Nepal, Bhutan, India
Pardosa tridentis [27]Solukhumbu/ Province 1Nepal, India, Kashmir
7.
TrochosaC.L. Koch, 1847
Trochosa gravelyi [27]
Kavre/ Bagmati Province
Endemic
XIV. FAMILY MYSMENIDAE Petrunkevitch, 1928
1.
IardinisSimon, 1899
Iardinis martensi [7]
Dolakha/ Bagmati P.
Endemic
XV. FAMILY NESTICIDAE Simon, 1894
1.
NesticellaLehtinen &Saaristo, 1980
Nesticella nepalensis [85]
Dolakha/ Bagmati P.
Nepal, China, India
XVI. FAMILY OONOPIDAE Simon, 1890
1.BrignoliaDumitrescu and Georgescu, 1983Brignolia ankhu [60]Dhading/ Bagmati P.Endemic
Brignolia assam [60]Nuwakot/ Bagmati P.Nepal and India
Brignolia sukna [60]Ilam/ Province 1Nepal and India
2.CamptoscaphiellaCaporiacco, 1934Camptoscaphiella gunsa [86]Taplejung/ Province 1Nepal and India
Camptoscaphiella martensi [86]Mustang/ Gandaki P.Endemic
Camptoscaphiella nepalensis [86]Parbat/ Gandaki P.Endemic
Camptoscaphiella panchthar [86]Panchthar/ Province 1Endemic
Camptoscaphiella silens [86]Solukhumbu/ Province 1Endemic
Camptoscaphiella strepens [86]Gorkha/ Gandaki P.Endemic
Camptoscaphiella taplejung [86]Taplejung/ Province 1Endemic
3.HimalayanaGrismado, 2014Himalayana castanopsis [61]Ilam/ Province 1Endemic
Himalayana kathmandu [61]Kathmandu/ Bagmati P.Endemic
Himalayana martensi [61]Manang/ Gandaki P.Endemic
Himalayana parbat [61]Parbat/ Gandaki P.Endemic
4.PrethopalpusBaehr et al., 2012Prethopalpus ilam [87]Ilam/ Province 1Endemic
5.
TrilacunaTong & Li, 2007
Trilacuna bangla [61]
Sindhupalchowk/ Bagmati Province
Nepal and India
XVII. FAMILY OXYOPIDAE Thorell, 1869
1
OxyopesLatreille, 1804
Oxyopes javanus [58]NepalNepal, China, India
Oxyopes lineatus [58]NepalNepal, Europe, Turkey ,Russia
Oxyopes sertatus [58]
Nepal
Nepal, China, Korea, Japan
XVIII. FAMILY PHOLCIDAE C.L. Koch, 1850
1.
PholcusWalckenaer, 1805
Pholcus calligaster [88]Parsa/ Madhesh ProvinceNepal and Myanmar
Pholcus zham [88]
Sankhuwasabha/Province1
Nepal and China
XIX. FAMILY PIMOIDAE Wunderlich, 1986
1.
PimoaChamberlin & Ivie, 1943
Pimoa daman [67]Makwanpur/ Bagmati P.Endemic
Pimoa khaptad [67]Bajhang/ Sudurpashim P.Endemic
Pimoa koshi [67]Sankhuwasabha/Province1Endemic
Pimoa mechi [67]Taplejung/ Province 1Endemic
Pimoa mude [67]Sindhupalchowk/BagmatiEndemic
Pimoa nematoides [88]Dolakha/ Bagmati P.Endemic
Pimoa phaplu [67]Solukhumbu/ Province 1Endemic
Pimoa rara [67]Mugu/ Karnali ProvinceEndemic
Pimoa sinuosa [88]
Kaski/ Gandaki Province
Endemic
XX. FAMILY PISAURIDAE Simon, 1890
1.
PerenethisL. Koch, 1878
Perenethis sindica [89]
Taplejung/ Province 1
India, Sri Lanka, Nepal, China
XXI. FAMILY PSECHRIDAE Simon, 1890
1.PsechrusThorell, 1878Psechrus himalayanus [90]Rolpa/ Lumbini ProvinceIndia, Nepal
Psechrus marsyandi [45]Lamjung/ Gandaki P.Endemic
2.FeceniaSimon, 1887Fecenia protensa [58]∗Synonym: Facenia nicobarensisNepalNepal, Thailand, Vietnam, Brunei, Malaysia, India
Fecinia nicobarensis was transferred from genus Psechrus (Thorell, 1878)
XXII. FAMILY PSILOCERCIDAE Machado, 1951
1.LeclerceraDeeleman-Reinhold, 1995Leclercera ekteenensis [64]Panchthar/ Province 1Endemic
Leclercera machadoi [9]Baglung/ Gandaki P.Endemic
Leclercera mulcata [9]transferred from genus PsilodercesKathmandu/ Bagmati P.Endemic
Leclercera nagarjunensis [63]Endemic
Leclercera niuqu [63]Panchthar/ Province 1Endemic
Leclercera sidai [63]Ilam/ Province 1Endemic
Leclercera zhaoi [63]Endemic
Psilocercidae, a sub family of Ochyroceratidae was raised to family by Wunderlich (2008)
XXIII. FAMILY SALTICIDAE Blackwall, 1841
1.AsemoneaO Pickard-Cambridge, 1869Asamonea tenuipes [66]Chitwan/ Bagmati Province.Nepal, Sri Lanka, India, Myanmar, Vietnam, Singapore
2.BianorPeckham and Peckham, 1886Bianor albobimaculatus [91]Manang/ Gandaki P.Nepal, Iran, Pakistan India
Bianor tortus [37]Ilam/ Province 1Nepal and India
3.BrettusThorell, 1895Brettus anchorum[36]Gorkha/ Gandaki ProvinceNepal and India
4.CarrhotusThorell, 1891Carrhotus assam [91]Kaski/ Gandaki ProvinceNepal and India
Carrhotus catagraphus [34]Gorkha/ Gandaki ProvinceEndemic
Carrhotus erus [91]Kaski/ Gandaki P.Nepal and India
Carrhotus operosus [34]Mustang/ Gandaki P.Endemic
Carrhotus s-bulbosus [32]Sankhuwasabha/Province1Endemic
Carrhotus sannio [91]Myagdi/ GandakiNepal, China, India
Carrhotus viduus [34]Nepal, China. India, Iran
5.ChalcoscirtusBertkau, 1880Chalcoscirtus jiricus [22]transferred from genus EuophrysDolakha/ Bagmati ProvinceEndemic
Chalcoscirtus martensi [22]Mustang/ Gandaki P.Nepal, India and China
6.ChinattusLogunov, 1999Chinattus chichila [92]Sankhuwasabha/Province1Endemic
Chinattus validus [93]Myagdi/ Gandaki P.Nepal,Bhutan,China
7.ChrysillaThorell, 1887Chrysilla volupe [66]Chitwan/ Bagmati ProvinceNepal, Bhutan, India,Sri Lanka
8.EpeusPeckham and Peckham, 1886Epeus exdomus [94]Kathmandu/ Bagmati P.Endemic
Epeus indicus [35]Nuwakot/ Bagmati P.Nepal and India
9.EpocillaThorell, 1887Epocilla aurantiaca [66]Chitwan/ Bagmati Province.Nepal, Sri Lanka, Malaysia, Vietnam, India
10.EuophrysC. L. Koch, 1834Euophrys dhaulagirica [22]Mustang/ Gandaki P.Endemic
Euophrys nepalica [22]Myagdi/ Gandaki P.Nepal and China
Euophrys omnisuperstes [5]Sankhuwasabha/Province1Nepal and India
Euophrys yulungensis [22]Dolpa/ Karnali ProvinceChina and Nepal
11.HabrocestoidesPrószyn'ski, 1992Habrocestoides phulchokiensis [95]Lalitpur/ Bagmati ProvinceEndemic
12.HarmochirusSimon, 1885Harmochirus zabkai [96]Kathmandu/ Bagmati P.India, Nepal, Vietnam
13.HyllusC.L. Koch, 1846Hyllus semicupreus [66]Chitwan/ Bagmati Province.Nepal, Sri Lanka, India
14.IciusSimon, 1876Icius alboterminus [66]Nepal and India
15.NepalicusBlackwall, 1841Nepalicius nepalicus [97]∗Transferred from genus PseudiciusKathmandu/ Bagmati ProvinceNepal and India
16.OrientattusCaleb, 2020Orientattus minutes [23]O.minutes was transferred from genus PancoriusGorkha/ Gandaki ProvinceNepal
17.PancoriusSimon, 1902Pancorius armatus [39]Parbat/ Gandaki ProvinceEndemic
Pancorius cadus [39]Taplejung/ Province 1Endemic
Pancorius kaskiae [23]Kaski/ Gandaki ProvinceEndemic
Pancorius magnus [39]Ilam/ Province 1Nepal, China, India
Pancorius urnus [39]Ilam/ Province 1Endemic
18.PhaeaciusSimon, 1900Phaeacius fimbriatus [36]Sankhuwasabha/Province1Nepal, Indonesia, Java
Phaeacius saxicola [98]Taplejung/ Province 1Endemic
Phaeacius wanlessi [36]Sankhuwasabha/Province1Nepal, Sri Lanka
19.PhintellaStrand, 1906Phintella suavis [2]NepalNepal to Malaysia
Phintella vittata [66]Chitwan/ Bagmati P.Nepal, China, India
20.Plexippoides Prószyn'ski, 1984Plexippoides tristis [99]Mustang/ Gandaki P.Endemic
21.PlexippusC.L. Koch, 1846Plexippus paykulli [23]Myagdi/ Gandaki P.Asia, Africa, America, Europe
Plexippus petersi [23]Kaski/ Gandaki ProvinceAsia, Africa and Pacific islands
Plexippus pokharae [23]Endemic
22.PortiaKarsch, 1878Portia fimbriata [36]Kathmandu/ Bagmati P.Nepal, Sri Lanka, Taiwan to Australia
23.PtocasiusSimon, 1885Ptocasius nepalicus [20]Synonym: Yaginumaella nepalicaMustang/ Gandaki ProvinceNepal and China
Ptocasius tenzingi [20]Synonym: Yaginumaella tenzingiSolukhumbu/ Province 1Endemic
Ptocasius thakkholaicus [20]Synonym: Yaginumaella thakkholaicaMustang/ Gandaki ProvinceNepal and China
24.RheneThorell, 1869Rhene flavicomans [33]Sankhuwasabha/Province1Nepal, Bhutan, India, Thailand
Rhene phuntsholingensis [33]Nepal, Bhutan
25.SilerSimon, 1889Siler cupreus [66]Chitwan/ Bagmati P.Nepal, China, Taiwan, Korea, Japan
26.SitticusSimon 1901Sitticus niveosignatus [21]Dolpa/ Karnali ProvinceNepal to China
27.StenaelurillusSimon,1886Stenaelurillus triguttatus [100]Narayangadh/ Bagmati P.Nepal and China
28.SynagelidesStrand, 1906Synagelides bagmaticus [101]Bhaktapur/ Bagmati P.Endemic
Synagelides gosainkundicus [101]Rasuwa/ Bagmati P.Endemic
Synagelides kosi [101]Ramechhap/ Bagmati P.Endemic
Synagelides martensi [101]Synonyms: Synagelides dhaulagiricus ,Synagelides himalaicus, Synagelides jiricus, Synagelides thodungus & Synagelides wyszynskiiDolpa/ Karnali ProvinceEndemic
Synagelides nepalensis [24]Endemic
Synagelides nishikawai [25]Myagdi/ Gandaki P.Endemic
Synagelides oleksiaki [24]Synonym: Synagelides gorapanicusRamechhap/ Bagmati P.Endemic
Synagelides tukchensis [24]Mustang/ Gandaki P.Endemic
Synagelides ullerensis [24]Parbat/ Gandaki P.Endemic
Synagelides walesai [24]Lalitpur/ Bagmati P.Endemic
29.TelamoniaThorell,1887Telamonia dimidiata [66]Chitwan/ Bagmati P.Nepal, Bhutan, Malaysia
Telamonia festiva [66]Chitwan/ Bagmati P.Nepal, China, India
30.
ThyeneSimon,1885
Thyene bivittata [38]Kathmandu/ Bagmati P.Nepal, China, Pakistan
Thyene typica [38]Sankhuwasabha /Province 1Endemic
Thyene yuxiensis [38]
Tanahu/ Gandaki P.
Nepal and China
XXIV. FAMILY SCYTODIDAE Blackwall, 1864
1.
ScytodesLatreille, 1804
Scytodes mawphlongensis [10]
Lalitpur/ Bagmati Province
Nepal and India
XXV. FAMILY SELENOPIDAE Simon, 1897
1.
MakdiopsCrews and Harvey,2011
Makdiops montigena [102]
Chitwan/ Bagmati Province
Nepal and India
XXVI. FAMILY SPARASSIDAE Bertkau, 1872
1.BhutaniellaJäger, 2000Bhutaniella hillyardi [41]Sankhuwasabha/Province1Endemic
Bhutaniella rollardae [43]Pyuthan/ Lumbini P.Endemic
2.OliosWalckenaer,1837Olios rossetti [44]Kavre/ Bagmati ProvinceNepal, India, Pakistan
3.PseudopodaJäger, 2000Pseudopoda albolineata [82]Myagdi/ Gandaki P.Endemic
Pseudopoda alta [43]Kaski/ Gandaki ProvinceEndemic
Pseudopoda ausobskyi [43]Ilam/ Province 1Endemic
Pseudopoda brauni [43]Taplejung/ Province 1Endemic
Pseudopoda chauki [43]Terathum/ Province 1Endemic
Pseudopoda chulingensis [43]Gorkha/ Gandaki ProvinceEndemic
Pseudopoda cuneata [43]Myagdi/ Gandaki P.Endemic
Pseudopoda dama [43]Jhapa/ Province 1Endemic
Pseudopoda damana [43]Makwanpur/ Bagmati P.Endemic
Pseudopoda dhulensis [43]Baglung/ Gandaki P.Endemic
Pseudopoda diversipunctata [43]Kaski/ Gandaki ProvinceEndemic
Pseudopoda everesta [43]Solukhumbu/ Province 1Endemic
Pseudopoda grasshoffi [43]Sankhuwasabha /Province 1Endemic


Pseudopoda heteropodoides [43]Taplejung/ Province 1Endemic
Pseudopoda huberti [43]Pyuthan/ Lumbini P.Endemic
Pseudopoda hyatti [43]Myagdi/ Gandaki P.Endemic
Pseudopoda jirensis [43]Dolakha/ Bagmati P.Endemic
Pseudopoda kalinchoka [43]Dolakha/ Bagmati P.Endemic
Pseudopoda khimtensis [43]Ramechhap/ Bagmati P.Endemic
Pseudopoda latembola [43]Manang/ Gandaki P.Endemic
Pseudopoda marmoreal [43]Kaski/ Gandaki ProvinceEndemic
Pseudopoda martensi [43]Mustang/ Gandaki P.Endemic
Pseudopoda martinae [43]Rasuwa/ Bagmati P.Endemic
Pseudopoda monticola [43]Lalitpur/ Bagmati P.Endemic
Pseudopoda schawalleri [43]Panchthar/ Province 1Endemic
Pseudopoda sinopodoides [43]Kathmandu/ Bagmati P.Endemic
Pseudopoda tinjura [43]Tehrathum/ Province 1Endemic
Pseudopoda triapicata [43]Ilam/ Province 1Endemic
Pseudopoda trisuliensis [43]Rasuwa/ Bagmati P.Endemic
Pseudopoda varia [43]
Taplejung/ Province 1
Endemic
XXVII. FAMILY SYMPHYTOGNATHIDAE Hickman, 1931
1
IardinisSimon,1899
Iardinis martensi [7]
Dolakha/ Bagmati Province
Endemic
XXVIII. FAMILY TETRABLEMMIDAE O.P-Cambridge, 1873
1.BrignoliellaShear, 1978Brignoliella martensi [8]Lalitpur/ Bagmati ProvinceEndemic
2.
TetrablemmaO.P.-Cambridge, 1873
Tetrablemma phulchoki [14]

Endemic
XXIX. FAMILY TETRAGNATHIDAE Menge, 1866
1.LeucaugeWhite,1841Leucauge decorata [58]NepalNepal, Japan, Thailand, Bangladesh, China, India
2.
TetragnathaLatreille,1804
Tetragnatha bogotensis [103]Synonym: Tetragnatha boydi
Nepal
Nepal, Spain, Mexico to Paraguay
XXX. FAMILY THERAPHOSIDAE Thorell, 1870
1.
HaplocosmiaSchmidt & von Wirth, 1996
Haplocosmia nepalensis [104]
Kaski/ Gandaki Province
Endemic
XXXI. FAMILY THERIDIIDAE Sundevall, 1833
1.CarniellaThaler & SteinbergerCarniella nepalensis [105]Taplejung/ Province 1Endemic
2.
LactrodectusWalckenaer,1805
Lactrodectus elegans [65]
Gorkha/ Gandaki Province
Nepal, China, Japan , India, Myanmar
XXXII. FAMILY THOMISIDAE Sundevall, 1833
1.BassaniodesPocock, 1903Bassaniodes dolpoensis [15]∗transferred from genus XysticusDolpa/ Karnali provinceNepal and China
2.LysitelesSimon, 1895Lysiteles annapurnus [18]Kaski/ Gandaki ProvinceEndemic
Lysiteles himalayensis [18]Myagdi/ Gandaki ProvinceBhutan, Nepal
Lysiteles lepusculus [18]Mustang/ Gandaki P.Endemic
Lysiteles maius [18]Baitadi/ Gandaki P.Russia, Nepal to Japan
Lysiteles montivagus [18]Mustang/ Gandaki P.Endemic
Lysiteles niger [18]Makwanpur/ Bagmati P.Bhutan, Nepal
Lysiteles parvulus [18]Myagdi/ Gandaki ProvinceEndemic
Lysiteles saltus [18]Bhutan , Nepal, China
3.MonaesesThorell, 1869Monaeses aciculus [16]Taplejung/ Province 1Nepal to Japan, Philippines
4.PsammitisMenge, 1876Psammitis nepalhimalaicus [15]∗transferred from genus XysticusDolakha/ Bagmati ProvinceEndemic
Psammitis potamon [15]∗transferred from genus XysticusMyagdi/ Gandaki ProvinceEndemic
Psammitis simplicipalpatus [15]∗transferred from genus XysticusDolpa/ Karnali ProvinceNepal and Bhutan
5.RunciniaSimon, 1875Runcinia roonwali [58]NepalNepal and India
Runcinia insecta [58]previously published as Thomisus cherapunjeusNepalAsia, Africa, Australia
6.StiphropusGerstäcker, 1873Stiphropus soureni [17]Kavre/ Bagmati P.India, Nepal, Bhutan
7.
XysticusC.L. Koch, 1835
Xysticus alpinistus [15]Dolakha/ Bagmati P.Nepal, China
Xysticus cristatus [15]Mustang/ Gandaki P.Nepal, Kazakhstan, Iran
Xysticus croceus [2]NepalIndia, Nepal, Bhutan, China
Xysticus elephantus [15]Dolpa/ Karnali ProvinceNepal, China
Xysticus martensi [15]Endemic
Xysticus roonwali [106]Solukhumbu/ Province 1Nepal, India
Xysticus cf sikkimus [15]
Mustang/ Gandaki P.
Nepal, China, India
XXXIII. FAMILY TITANOECIDAE Lehtinen, 1967
1.
AnuvindaLehtinen, 1967
Anuvinda milloti [107]transferred from genus Amaurobius
Chitwan/ Bagmati P.
Endemic
XXXIV. FAMILY ZODARIIDAE Thorell, 1881
1.MallinellaStrand, 1906Mallinella erratica [19]∗transferred from genus StorenaIlam/ Province 1Endemic
Mallinella martensi [19]∗transferred from genus StorenaMustang/ Gandaki ProvinceEndemic
Mallinella nepalensis [19]∗transferred from genus StorenaRasuwa/ Bagmati ProvinceEndemic
Mallinella uncinata [19]∗transferred from genus StorenaKaski/ Gandaki ProvinceEndemic
2.SuffasiaJocqué, 1991Suffasia kanchenjunga [31]Ilam/ Province 1Endemic
Suffasia martensi [31]Ilam/ Province 1Endemic
Suffasia tumegaster [108]Lalitpur/ Bagmati P.Endemic
3.TropizodiumJocque & Churchill, 2005Tropizodium bengalensis [58]transferred from genus LuticaNepalNepal and India
  8 in total

1.  Thirty-five new species of the spider genus Pimoa (Araneae, Pimoidae) from Pan-Himalaya.

Authors:  Hao Xu; Xiaoqing Zhang; Zhiyuan Yao; Abid Ali; Shuqiang Li
Journal:  Zookeys       Date:  2021-04-08       Impact factor: 1.546

2.  Four new species of the spider genus Leclercera Deeleman-Reinhold (Araneae, Ochyroceratidae) from Nepal.

Authors:  Mingjie Xu; Fengyuan Li; Shuqiang Li
Journal:  Zootaxa       Date:  2018-08-23       Impact factor: 1.091

3.  The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental Region.

Authors:  Sarah C Crews; Mark S Harvey
Journal:  Zookeys       Date:  2011-05-19       Impact factor: 1.546

4.  The spider genus Olios Walckenaer, 1837 (Araneae: Sparassidae)-Part 1: species groups, diagnoses, identification keys, distribution maps and revision of the argelasius-, coenobitus- and auricomis-groups.

Authors:  Peter JÄger
Journal:  Zootaxa       Date:  2020-10-22       Impact factor: 1.091

5.  Global patterns of guild composition and functional diversity of spiders.

Authors:  Pedro Cardoso; Stano Pekár; Rudy Jocqué; Jonathan A Coddington
Journal:  PLoS One       Date:  2011-06-29       Impact factor: 3.240

6.  On the spider genus Amaurobius (Araneae, Amaurobiidae) in India and Nepal.

Authors:  Yuri M Marusik; Francesco Ballarin; Mikhail M Omelko
Journal:  Zookeys       Date:  2011-01-31       Impact factor: 1.546

7.  Thirty-one new species of the spider genus Leclercera from Southeast Asia (Araneae, Psilodercidae).

Authors:  Wan-Jin Chang; Shuqiang Li
Journal:  Zookeys       Date:  2020-02-19       Impact factor: 1.546

  8 in total

北京卡尤迪生物科技股份有限公司 © 2022-2023.