Duleepa Pathiraja1, June Wee2, Kijong Cho3, In-Geol Choi4. 1. Department of Biotechnology, College of Life Sciences and Biotechnology, Korea University, Seoul, 02841, Korea. 2. BK21 FOUR R&E Center for Environmental Science and Ecological Engineering, Korea University, Seoul, 02841, Korea. 3. Department of Environmental Science and Ecological Engineering, College of Life Sciences and Biotechnology, Korea University, Seoul, 02841, Korea. kjcho@korea.ac.kr. 4. Department of Biotechnology, College of Life Sciences and Biotechnology, Korea University, Seoul, 02841, Korea. igchoi@korea.ac.kr.