Wenkun Zhou1, Xiaoyue Zhang2. 1. State Key Laboratory of Plant Physiology and Biochemistry, College of Biological Sciences, China Agricultural University, Beijing, 100193, China. 2. State Key Laboratory of Plant Genomics, National Center for Plant Gene Research (Beijing), Institute of Genetics and Developmental Biology, Innovation Academy of Seed Design, the Chinese Academy of Sciences, Beijing, 100101, China.