David J McKenzie1, Benjamin Geffroy1, Anthony P Farrell2,3. 1. MARBEC, Univ Montpellier, CNRS, IFREMER, IRD, Montpellier, France. 2. Department of Zoology, University of British Columbia, Vancouver, British Columbia, Canada. 3. Faculty of Land and Food Systems, University of British Columbia, Vancouver, British Columbia, Canada.