Wenliang Zhou1, Meng Wang2, Mingpan Huang2, Fuwen Wei3,2,4. 1. Center for Evolution and Conservation Biology, Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou 511458, China. 2. Key Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China. 3. Center for Evolution and Conservation Biology, Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou 511458, China. weifw@ioz.ac.cn. 4. Endangered Species Scientific Commission of the People's Republic of China, Beijing 100101, China.