Jian Wang1, James F Parham2, Haitao Shi3. 1. College of Life Science and Technology, Honghe University, Mengzi 661199, China. 2. Department of Geological Sciences, California State University, Fullerton, CA 92834, USA. 3. Ministry of Education Key Laboratory for Ecology of Tropical Islands, College of Life Sciences, Hainan Normal University, Haikou 571158, China. haitao-shi@263.net.