Andrew D Letten1, Masato Yamamichi2,3. 1. School of Biological Sciences, The University of Queensland, Brisbane, QLD 4072, Australia; a.letten@uq.edu.au. 2. School of Biological Sciences, The University of Queensland, Brisbane, QLD 4072, Australia. 3. Department of International Health and Medical Anthropology, Institute of Tropical Medicine, Nagasaki University, Nagasaki 852-8523, Japan.