First checklist of the chrysidid wasps (Hymenoptera, Chrysididae) of Mongolia, with description of new species.

An annotated checklist of the Chrysididae from Mongolia is provided. A revision of the bibliographical data is provied, since most of the collecting localities published for "Mongolia" refer to places currently located in China. The known Mongolian cuckoo wasp fauna counts 90 species in 18 genera and two subfamilies. Four genera and 57 species are recorded for the first time, including two species here described as new for science: Cleptes mongolicus Rosa, Halada & Agnoli, sp. nov. (Dornod) and Spinolia spinosa Rosa & Halada, sp. nov. (Bayankhongor). Paolo Rosa, Maxim Yu. Proshchalykin, Marek Halada, Ulykpan Aibek.

Introduction

Mongolia is a large landlocked country in eastern Central Asia, covering 1,564,100 km². Politically, Mongolia is divided into 21 provinces named “aimags” with the capital Ulaanbaatar (Fig. 1). It is bordered by Russia to the north and China to the south, east, and west. Geographically and climatologically, it is an area of contrasts and extremes, between cold mountainous regions up to 4,000 m a.s.l. to the north and west and one of the largest deserts of the world in the south, the Gobi Desert. Most of the country is located on high plateaus, covered by steppes and extensive forested areas. It has an extreme continental climate with long, cold winters and short hot summers, during which most of its annual precipitation falls (Lavrenko 1979; Dathe and Proshchalykin 2016).

Figure 1.

Administrative map of Mongolia (from Dathe and Proshchalykin (2016) and Proshchalykin (2017) modified). Aimags: 1 Bayan-Ulgii 2 Uvs 3 Khovd 4 Zavkhan 5 Govi-Altai 6 Khuvsgul 7 Arkhangai 8 Bayankhongor 9 Bulgan 10 Orkhon 11 Uvurkhangai 12 Umnugovi 13 Selenge 14 Darkhan-Uul 15 Tuv 16 Ulaanbaatar 17 Dundgovi 18 Khentii 19 Govi-Sümber 20 Dornogovi 21 Dornod 22 Sukhbaatar.

Mongolian cuckoo wasps are scarcely known and a few occasional records are found in the literature (Rosa 2017a). Only one article (Móczár 1967) deals with Mongolian material collected by Dr Z. Kaszab during his entomological excursions in this country (1963–1968). Other scattered findings have been published (du Buysson 1901; Semenov-Tian-Shanskij 1912, 1932, 1967; Semenov-Tian-Shanskij and Nikol’skaya 1954; Linsenmaier 1997a; Rosa et al. 2017a, b), while most of the remaining bibliographical data recorded for “Mongolia” actually refer to localities currently included in China (Inner Mongolia, Xinjiang, Gansu) (du Buysson 1893; Radoszkowski 1877, 1891; Mocsáry 1890; Dalla Torre 1892; Bischoff 1913; Hammer 1936; Tsuneki 1947, 1953a; Linsenmaier 1959, 1968; Semenov-Tian-Shanskij 1967; Kimsey and Bohart 1991; Rosa et al. 2014, 2015). Approximately 30 species were properly recorded from Mongolia so far (Rosa 2017a) and we here add 57 new records for this country, mostly based on the materials collected by Czech entomologists (M. Halada, J. Halada, J. Straka, and M. Kadlecová) in 2003–2007 and mainly housed in the private collections of MH (České Budějovice, Czech Republic) and PR (Bernareggio, Italy). Other new records were found during the examination of the collection housed at the Zoological Institute in St. Petersburg (Russia, ZIN) and based on the material collected during the expeditions of V. Roborovskij and P. Kozlov in 1895 and P. Kozlov in 1926. Finally, a few specimens were examined from the material collected in Mongolia by Soviet-Mongolian expeditions in 1967–1982. Soviet-Mongolian expedition were conducted from 1967 to 1983 and led to the collection of extensive entomological material, which became the basis for the publication of numerous articles and books (including Insects of Mongolia in eleven volumes), devoted to the study of various insects families (Proshchalykin and Kuhlmann 2015), although the were never examined by anyone. Large part of the cuckoo wasps collected during these entomological expeditions is still unprepared and unidentified.

Unpublished distributional records from Mongolia were recently published in the volume on Russian (Rosa et al. 2019), for a better understanding of the distribution of the Asian species, but exact localities were omitted because they were not of interest for that publication. We here report the precise data of species recorded for the first time in Rosa et al. 2019, which are mostly based on material housed in the Linsenmaier collection (Luzern, Switzerland).

In the present paper, based on a comprehensive study of specimens (including primary types) deposited in various collections, we report additional records of 72 species, with two species described as new and 55 species recorded from Mongolia for the first time, resulting in a total number of 90 cuckoo wasps species known from this country (Table 1).

Table 1.

Records of Mongolian cuckoo wasp species by aimags.

No.	Species	Aimags
1.	Chrysis aestiva Dahlbom, 1854	7
2.	Chrysis angustula Schenck, 1856	7, 15
3.	Chrysis asahinai Tsuneki, 1950	8, 9, 12, 15, 20, 22
4.	Chrysis belokobylskiji Rosa, 2019	4, 12, 15
5.	Chrysis brevitarsis Thomson, 1870	9
6.	Chrysis castigata Linsenmaier, 1959	13, 15
7.	Chrysis chinensis Mocsáry, 1912	7, 13, 15
8.	Chrysis consanguinea Mocsáry, 1889	4, 7, 9, 13, 15, 16, 18, 21, 22
9.	Chrysis dauriana Linsenmaier, 1959	4, 7–9, 13, 18
10.	Chrysis equestris Dahlbom, 1854	7, 13
11.	Chrysis fulgida Linnaeus, 1761	7, 13, 15
12.	Chrysis ignita (Linnaeus, 1758)	9
13.	Chrysis illecebrosa Semenov, 1967	12
14.	Chrysis illigeri Wesmael, 1839	13, 15
15.	Chrysis ismaeli Semenov, 1967	12, 20, 21
16.	Chrysis jaxartis Semenov, 1910	12, 13, 15, 18, 21
17.	Chrysis leptomandibularis Niehuis, 2000	15
18.	Chrysis mane Semenov, 1912	15
19.	Chrysis matutina Semenov, 1967	7
20.	Chrysis mediata Linsenmaier, 1951	15
21.	Chrysis mocsaryi Radoszkowski, 1889	3
22.	Chrysis mysticalis Linsenmaier, 1959	4, 7, 9, 15, 20
23.	Chrysis nox Semenov, 1954	5, 15
24.	Chrysis pavesii Rosa, 2017	5, 15
25.	Chrysis priapus Rosa, 2018	5
26.	Chrysis pseudobrevitarsis Linsenmaier, 1951	7, 15
27.	Chrysis pupilla Semenov, 1967	12
28.	Chrysis rutilans Olivier, 1791	15
29.	Chrysis schencki Linsenmaier, 1968	7, 9
30.	Chrysis sibirica Rosa, 2017	7
31.	Chrysis solida Haupt, 1957	21
32.	Chrysis splendidula unica Radoszkowski, 1891	7
33.	Chrysis subcoriacea Linsenmaier, 1959	7
34.	Chrysis viridula Linnaeus, 1761	15
35.	Chrysura dichroa (Dahlbom, 1854)	4
36.	Chrysura ignifrons (Brullé, 1833)	4
37.	Cleptes dauriensis Móczár, 1997	3, 8, 11
38.	Cleptes mongolicus Rosa, Halada, & Agnoli, sp. nov.	21
39.	Colpopyga nesterovi Rosa, 2017	21
40.	Elampus albipennis (Mocsáry, 1889)	7, 20
41.	Elampus coloratus Rosa, 2017	22
42.	Elampus montanus (Mocsáry, 1890)	20
43.	Elampus panzeri (Fabricius, 1804)	4, 7
44.	Elampus sanzii Gogorza, 1887	15
45.	Elampus spinifemoris (Móczár, 1967)	11
46.	Euchroeus mongolicus Tsuneki, 1947	5, 11, 12
47.	Euchroeus orientis Semenov, 1910	22
48.	Hedychridium ardens (Coquebert, 1801)	4, 7, 8, 11, 13, 16, 18, 21, 22
49.	Hedychridium asianum Linsenmaier, 1997	7–9, 16
50.	Hedychridium belokobylskiji Rosa, 2017	15
51.	Hedychridium cupreum (Dahlbom, 1845)	4, 5, 8, 11, 12, 15, 20
52.	Hedychridium gabriellae Rosa, 2017	8, 15, 20
53.	Hedychridium longigena Rosa, 2017	8, 9, 13, 15, 18, 20, 21
54.	Hedychridium propodeale Rosa, 2017	5
55.	Hedychridium roseum (Rossi, 1790)	7, 20–22
56.	Hedychrum chalybaeum Dahlbom, 1854	5, 8, 13, 15, 16, 21, 22
57.	Hedychrum gerstaeckeri Chevrier, 1869	13, 15, 18
58.	Hedychrum lama du Buysson, 1891	3
59.	Hedychrum longicolle Abeille de Perrin, 1877	9, 12, 15, 21, 22
60.	Hedychrum nobile (Scopoli, 1763)	4, 7, 13, 15
61.	Hedychrum rutilans ermak Semenov, 1967	7, 13, 15, 21, 22
62.	Holopyga generosa asiatica Trautmann, 1926	13
63.	Holopyga kaszabi Móczár, 1967	11, 12, 20
64.	Holopyga minuma Linsenmaier, 1959	21, 22
65.	Omalus aeneus (Fabricius, 1787)	15, 16
66.	Omalus berezovskii (Semenov, 1932)	16
67.	Omalus margianus (Semenov, 1932)	7–9, 15, 22
68.	Omalus miramae (Semenov, 1932)	8, 20, 22
69.	Omalus stella (Semenov, 1932)	7, 11, 15
70.	Parnopes glasunowi Semenov, 1901	3
71.	Parnopes popovii Eversmann, 1858	7, 9, 12, 15, 20–22
72.	Pentachrysis amoena (Eversmann, 1858)	without locality
73.	Philoctetes bogdanovii (Radoszkowski, 1877)	7
74.	Philoctetes cynthiae Rosa, 2017	8, 11, 16, 22
75.	Philoctetes diakonovi (Semenov, 1932)	20
76.	Philoctetes lyubae Rosa, 2017	20
77.	Philoctetes mongolicus (du Buysson, 1901)	7, 8, 11, 15, 16, 18, 22
78.	Philoctetes shokalskii (Semenov, 1932)	8, 11, 12, 15, 16, 18–22
79.	Pseudochrysis gengiskhan Rosa, 2017	8, 9, 13, 15, 21, 22
80.	Pseudochrysis neglecta (Shuckard, 1837)	15
81.	Pseudomalus auratus nigridorsus (Tsuneki, 1953)	4, 9, 15, 18
82.	Pseudomalus corensis (Uchida, 1927)	9, 13, 15, 16, 18, 21
83.	Pseudomalus punctatus (Uchida, 1927)	9, 15, 18, 21
84.	Pseudomalus pusillus (Fabricius, 1804)	8, 9, 11–13, 15, 18, 21
85.	Spinolia spinosa Rosa & Halada, sp. nov.	8
86.	Spinolia unicolor (Dahlbom, 1831)	5
87.	Stilbum calens (Fabricius, 1781)	7, 9, 11, 15, 20
88.	Trichrysis cyanea (Linnaeus, 1758)	8, 13, 15
89.	Trichrysis pellucida (du Buysson, 1887)	without locality
90.	Trichrysis secernenda (Mocsáry, 1912)	13

Comment. Aimag designation as in Fig. 1.

Materials and methods

Terminology follows Lanes et al. (2020), Hymenoptera Anatomy Ontology (HAO 2020), and partly Kimsey and Bohart (1991). Abbreviations used in the descriptions are as follows:

flagellomeres 1, 2, 3, etc., respectively;

l/w length/width;

anterior ocellus diameter;

malar space, the shortest distance between base of mandible and lower margin of compound eye;

the shortest distance between posterior ocellus and compound eye;

P pedicel;

puncture diameter;

the shortest distance between posterior ocelli;

T1–T5 metasomal terga numbered consecutively, starting with 1 at the second abdominal segment.

Pictures of the types were taken with Nikon D700 connected to the microscope Togal SCZ and stacked with the software Combine ZP.

The checklist follows the genera subdivision proposed by Kimsey and Bohart (1991), with few exceptions for some genera (e.g., Latreille, 1809, Semenov, 1891 and Semenov, 1954). The species are listed alphabetically. We have used the following abbreviations for collectors: – J. Halada; – J. Straka; – M. Halada; – M. Kadlecová. An asterisk (*) marks the new records.

Types and other specimens are deposited in the following Institutions and private collections:

Entomology Institute, Hokkaido University (Japan);

Hungarian Natural History Museum, Zoological Department, Budapest (Hungary);

Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Kraków (Poland);

Linnean Society, London (England);

Museo National de Ciencias Naturales, Madrid (Spain);

Museum für Naturkunde, Berlin (Germany);

Museum d’Histoire Naturelle, Geneva (Switzerland);

National Museum of Natural History, Paris (France);

Museo di Storia Naturale, Genova (Italy);

British Museum of Natural History, London (UK);

Museum of Natural History, Vienna (Austria);

National Institute of Agro-Environmetal Science, Tsukuba (Japan);

Natur Museum, Luzern (Switzerland);

Osaka Museum of Natural History, Osaka (Japan);

Zoological Institute, Russian Academy of Sciences, St. Petersburg (Russia);

Zoological Museum of Moscow Lomonosov State University (Russia);

Zoologiska Museet, Lund Zoological Museum, University of Lund (Sweden);

G.L. Agnoli collection (Bologna, Italy);

M. Halada collection (České Budějovice, Czech Republic);

P. Rosa collection (Bernareggio, Italy);

U. Koschwitz collection (Eppenbraun, Germany).

Results

Taxa from Mongolia

Subfamily

Genus Latreille, 1802

Latreille, 1802: 316. Type species: Linnaeus, 1761 [= (Linnaeus, 1761)], by monotypy.

Móczár, 1997

0FA19F1D-FEBD-5567-A083-9689B8EEB63C

Móczár, 1997: 36. Holotype ♀: Russia: Dauria, leg. F. Sahlb., “

:

Material examined.

Mongolia: Khovd, 1 ♂, Bodongin-Gol River, 12 km SW Altai, 22.VII.1970, leg. M. Kozlov (ZIN); Uvurkhangai, 1 ♀, 12 km E of Arvaykheer, , 1800 m, 3.VII.2004, leg. JH (GLAC); Bayankhongor, 1 ♂, 16 km SW of Bayankhongor, , 2165 m, 10.VII.2004, leg. JH (GLAC).

Distribution.

Mongolia (Bayankhongor, Khovd, Uvurkhangai); Russia (Zabaikalskii Terr.) (Rosa 2017a).

Rosa, Halada & Agnoli sp. nov.

DAB361FF-A06A-5BAC-A41B-DC0DFA0A4D9D

http://zoobank.org/73389B93-F683-41CC-84AC-3E16ED9B3000

Figures 2 , 3

Figure 2.

sp. nov., female, holotype A habitus, dorsal view B head, frontal view C head and mesosoma, dorsal view D head and mesosoma, lateral view E metasoma, postero-lateral view F metasoma, dorso-lateral view. Scale bars: 1.0 mm.

Figure 3.

sp. nov., male, paratype A habitus, dorsal view B head, frontal view C head and mesosoma, dorsal view D mesosoma, lateral view E metasoma, postero-lateral view F metasoma, dorsal view. Scale bars: 1.0 mm.

Type material.

: ♀, Mongolia: Dornod, 100 km W of Choibalsan, 820 m, 23.VII.2007, leg. M. Halada (ZIN). : 1 ♂, same collecting locality and date (GLAC); 1 ♂, 20 km W of Choibalsan, , 800 m, 24.VII.2007, leg. M. Halada (PRC).

Diagnosis.

sp. nov. belongs to the species group, based on the pronotum without posterior pit row and without longitudinal median sulcus or posterior median keel. It is closely related only to Móczár, 2000 from Tajikistan, for its general habitus and colouration. The latter belongs to the group (Móczár 2000), for the modified pronotal structure, without posterior transversal groove, but with a posteromedian longitudinal keel. Besides the unmodified pronotum, the female of sp. nov. can be easily separated from the female of by: a) pubescence whitish, shorter on metasoma (max 2.5 MOD) (vs. blackish, longer on metasoma, up to 3 MOD); b) punctation on metasoma with polished T1, shallow and sparse tiny punctures on T2, double punctures on T3 (vs. scattered punctate on T1, densely and evenly punctate on T2 and T3); c) colouration: head entirely black; propodeum entirely blue; T3 and T4 laterally blue; pedicel and F1 yellow; femora apically, tibiae and tarsi yellow (vs. head blue; propodeum black with median blue spot; T3 and T4 fully black; pedicel and flagellum dark brown). The male of is currently unknown.

Description.

Female. Holotype (Fig. 2A–F). Body length 4.6 mm. Forewing length 2.7 mm. POL = 2.2 MOD; OOL = 2.7 MOD. MS = 2.0 MOD. P:F1:F2:F3 = 1.0:1.0:0.7:0.7. F1 1.5 × as long as wide, F2 1.1 × as long as wide. Head in frontal view 1.2 × as broad as long between lower edge of clypeus and vertex. Face and vertex with small, even, and sparse punctures (1–4 PD) (Fig. 2B). Clypeal lower margin simple, unmodified, 2 MOD width, without acute teeth at corners; lateral edges subparallel. Frontal sulcus broad and deep in the first part, from anterior ocellus to mid of face, faint in the second half, from mid-face to the clypeal margin (Fig. 2B). Mandibles tridentate. Ocellar triangle isosceles, without post-ocellar sulcus. Postero-lateral pits close to posterior ocelli deep and elongate. Pedicel as long as F1. Malar spaces elongate (2.0 MOD). Pronotum unmodified; pronotal neck finely striated transversally; posterior margin of pronotum simple, without transverse row of pits or median keel. Pronotum with small punctures similar to those on vertex. Mesoscutum and mesoscutellum scarcely punctate, with tiny and scattered punctures (Fig. 2C), largely impunctate; notauli and parapsidal lines deep and complete. Mesopleuron with small, deep punctures; transversely aligned medially; with short, deep scrobal sulcus on posterior half (Fig. 2D). Metascutellum noticeably reduced by large metanotal trough and by deep and large anteromedian suture. Metapleuron transversely striate. Metapostnotum (dorsal surface of metapectal-propodeal complex) short, irregularly reticulate, with large foveae along posterior margin, before the propodeal declivity. Propodeal posterior projections short, stout, and divergent. Wing veins and cells unmodified. All metasomal terga with impunctate, brownish stripe along posterior margin (Fig. 2F); T1 mostly impunctate, with a few, sparse, tiny punctures; T2 with even, sparse, small punctures (3–5 PD), posteromedially polished; T3 with dense, irregular and double punctation; scattered to polished toward the apical margin; T4 with large, scattered punctures. Head black, with violet reflections medially on clypeus; scapus dorsally violet, ventrally brownish without metallic reflections; P light brown and F1 yellow; other flagellomeres dark brown to blackish. Mandible dark brown, medially yellowish. Pronotal neck medially black; pronotum, mesonotum, mesopleuron, metanotum (excluding black anterior suture and axillary trough), metapleuron metallic red with purple reflections dorsally; propodeum dorsally blue, propodeal declivity black; body ventrally black. Metasoma entirely black; apical margin of each tergum with brownish stripe; laterally on T3 with feeble green reflections; laterally on T4 with extended blue reflections (Fig. 2E). Tegulae brown. Legs with tibiae and tarsi yellowish; coxae red to golden; profemur anteriorly metallic red excluding distal joint; metafemur posteriorly metallic; other parts brown.

Male. Paratypes. Body length 4.0–4.2 mm. POL = 1.6 MOD; OOL = 1.0 MOD. MS = 1.9 MOD. P:F1:F2:F3 = 1.0:1.4:0.9:0.9. F1 3.5 × as long as wide (width taken at distal apex), F2 1.5×. Head in frontal view 1.3 × as broad as long between lower edge of clypeus and vertex. Face and vertex with small, even, and denser punctures (1–2 PD) compared to female (Fig. 3B). Frontal sulcus narrow and visible in the first part, from anterior ocellus to brow, faint in the second half, from mid-face to the clypeal margin (Fig. 3B). Lower face medially with punctures more spaced 4–5 PD. Ocellar triangle, post-ocellar sulcus, and posterolateral pits similar to female. F1 1.5 × as long as P. Punctation overall similar to that of female; metascutellum larger, with narrow anteromedian mesoscutellar-metascutal suture; metapleuron polished. Other characters as in female. T1 with denser (2–5 PD), tiny punctures; T2 with even, denser (1–3 PD), small punctures (3–5 PD), posteromedially sparser to polished; T3 with dense, irregular and double punctation; scattered to polished toward the apical margin; T4 with similar punctures; T5 almost polished, with scattered punctures. Species sexually dimorphic with head and mesosoma bright green, including ventral side; propodeum blue. Mandible metallic green from base to half length. Scapus green, pedicel and flagellum black. Metasoma entirely black, with terga apically brownish and laterally with feeble blue reflections on T3 and T4 (Fig. 3E). Tegulae brown. Coxae and femora medially green; trochanters brown, femora distally and tarsi yellowish.

Etymology.

The specific epithet is named after the country of origin.

Mongolia (Dornod).

Tribe

Genus Linnaeus, 1761

Linnaeus, 1761: 414. Type species: Linnaeus, 1758 [= (Linnaeus, 1758)], by subsequent designation of Latreille 1810: 437.

Lichtenstein, 1876: 27. Type species: Dahlbom, 1854, by subsequent designation of Ashmead 1902: 226. Synonymized by Linsenmaier 1959: 91.

Dahlbom, 1854

AEE4D3F6-6EF6-571E-83D9-3949A5A4B9E4

Dahlbom, 1854: 286. Holotype ♀; Greece: Rhodes (Berlin ?) (

Mongolia: Arkhangai, 1 ♂, 90 km NE of Tsetserleg, , 27.VII.2005, leg. JH (MHC).

*Mongolia (Arkhangai); Asiatic-European, from Caucasus, Turkey, Greece, Iran, Palestine, European part of Russia to Mongolia (Rosa et al. 2019, present record).

Remarks.

This is the most eastern record for .

Schenck, 1856

223D1189-8F3B-5019-81C2-120F40534B84

Schenck, 1856: 28. Lectotype ♀ (designated by

Mongolia: Arkhangai, 5 ♀♀, 1 ♂, Chuluut Gol River, , 23.VII.2005, leg. JH (MHC); 4 ♀♀, 1 ♂, 70 km NE of Tsetserleg, 25.VII.2005, leg. JH (MHC); Tuv, 1 ♀, 2 ♂♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC).

*Mongolia (Arkhangai, Tuv); Asiatic-European, from western Europe to China and Russia (Rosa et al. 2019).

Tsuneki, 1950

66561CA1-3312-586D-A518-10BDD2E4C95C

Tsuneki, 1950: 80. Holotype ♀; China, Manchuria, 22.VIII.1938, leg. S. Asahina (

:

Mongolia: Bayankhongor, 12 ♂♂, 130 km S of Bayankhongor, , 1240 m, 6.VII.2004, leg. JH, MK (MHC, PRC); 1 ♀, ibid, Orog Nuur, 6–7.VII.2004, on Saxaul, leg. JS (PRC); Bulgan, 13 ♀♀, 4 ♂♂, Mongol Els Nat. Res., dunes, , 31.VII.2005, leg. JH (MHC); Sukhbaatar, 1 ♂, 100 km SSW of Baruun-Urt, 1100 m, 30.VII.2007, leg. MH (MHC); Tuv, 2 ♀♀, 2 ♂♂, 75 km W of Ulaanbaatar, dunes, 2.VIII.2005, leg. JH (MHC); 39 ♀♀, 37 ♂♂, 70 km W of Ulaanbaatar, 1070 m, dunes, 16.VIII.2007, leg. JH, MH (MHC); Umnugovi, 1 ♂, Gobi, 100 km SW of Dalanzadgad, Bayanzag, on Saxaul, 1–2.VII.2003, leg. JH (MHC).

Mongolia (*Bayankhongor, *Bulgan, Dornogovi, *Sukhbaatar, *Tuv, *Umnugovi); China (Liaoning) (Rosa et al. 2014).

Rosa, 2019

3891D06F-DD86-5B4F-AEF5-4D96F7222DFB

Rosa, 2019: 2. Holotype ♀; Kyrgyzstan: Naryn River near Karakolka (

Mongolia: Umnugovi, 1 ♀, Nogon-kub, N. Gobi, 1.VIII.1926, P. Kozlov (ZIN); Tuv, 1 ♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC); Zavkhan, 1 ♀, 40 km SW of Uliastay, dunes, 18.VII.2005, leg. JH (MHC).

Mongolia (Tuv, Umnugovi, Zavkhan); China (Qinghai), Kyrgyzstan, Tajikistan (Rosa 2019).

Thomson, 1870

7562EB77-12BF-5469-A482-AB7ED095ADD6

Thomson, 1870: 107. Holotype ♀; Sweden: Nerike [= Närke] (Lund) (examined) (

Mongolia: Bulgan, 1 ♀, 137 km NE of Aravaykheer, , 1250 m, 26.VII.2004, leg. JH (MHC).

*Mongolia (Bulgan); Asiatic-European, from western Europe to Russia (Rosa et al. 2019).

Linsenmaier, 1959

CF45C6EF-C88A-5FD3-802A-04D45CF4DBEA

Linsenmaier, 1951: 82. Holotype ♀; Uzbekistan: Ferghana (Budapest) (examined) (

Linsenmaier, 1959: 155. Replacement name for

Mongolia: Selenge, 2 ♂♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Tuv, 1 ♂, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MH (MHC).

*Mongolia (Selenge, Tuv); Kazakhstan, Kyrgyzstan, Turkmenistan, Uzbekistan, Russia (Eastern Siberia) (Rosa et al. 2019).

Mocsáry, 1912

81563555-1A1A-5F3B-93BA-469182A7B1C4

Mocsáry, 1912: 589. Holotype ♀; China: Shanghai (

:

Mongolia: Arkhangai, 24 ♂♂, Chuluut Gol River, , 23.VII.2005, leg. JH (MHC); 2 ♀♀, 4 ♂♂, 90 km NE of Tsetserleg, , 24.VII.2004, leg. JH (MHC); 2 ♀♀, ibid, 27.VII.2005, leg. JH (MHC); Tuv, 1 ♀, 1 ♂, Ulaanbaatar Bog Duul, 11.VII.1983, leg. Karl Bleyl (NMLS); 7 ♀♀, 28 ♂♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC); 6 ♂♂, Khangayn Mts, 5 km N of Khunt, 20.VII.2005, leg. JH (MHC); Selenge, 11 ♀♀, 19 ♂♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC).

Mongolia (*Arkhangai, *Selenge, *Tuv); Asiatic-European, from western Europe (Switzerland) to China (Helongjiang, Shanghai) (Linsenmaier 1959).

This species was previously reported from Mongolia (Rosa et al. 2019) without exact locality.

Mocsáry, 1889

BDDA689D-AC3F-5D71-BF0C-2E5790DA4889

Mocsáry, 1889: 299. Syntypes ♀♀; Italy: Sicily; Algeria (

Mongolia: Arkhangai, 1 ♂, Chuluut Gol River, , 23.VII.2005, leg. JH (MHC); 3 ♀♀, 90 km NE of Tsetserleg, , 27.VII.2005, leg. JH (MHC); Bulgan, 1 ♀, 137 km NE of Aravaykheer, , 1250 m, 2.VII.2004, leg. JH (MHC); Dornod, 1 ♀, 100 km W of Choilbalsan, 820 m, 23.VII.2007, leg. JH (MHC); Khentii, 4 ♂♂, 100 km NE of Ondorkhaan, Kerulen River, 970 m, 22.VII.2007, leg. MH (MHC); Selenge, 1 ♀, 2 ♂♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Sukhbaatar, 2 ♂♂, 100 km SSW of Baruun-Urt, 1100 m, 30.VII.2007, leg. MH (MHC); Tuv, 7 ♂♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC); 1 ♂, Khangaun Mts, 5 km N of Khunt, 20.VII.2005, leg. JH (MHC); 1 ♀, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. JH (MHC); Ulaanbaatar, 1 ♀, 7 km E of Ulaanbaatar, Gachuurt, , 31.VII.2002, leg. JS (MHC); Zavkhan, 1 ♂, 40 km SW of Uliastay, dunes, 18.VII.2005, leg. JH (MHC).

*Mongolia (Arkhangai, Bulgan, Dornod, Khentii, Selenge, Sukhbaatar, Tuv, Ulaanbaatar, Zavkhan); Palaearctic, from southern Europe and northern Africa to Eastern Siberia ang Mongolia (Rosa et al. 2019, present records).

Linsenmaier, 1959

137B485D-37FC-5FA2-BCB4-38B6933A8E8B

Linsenmaier, 1959: 112. Holotype ♀; Russia: Dauria (

Semenov-Tian-Shanskij, 1967: 178, nec Mocsáry, 1914. Holotype ♀; Russia [not Mongolia]; Transbaikalia: Ingoda River (St. Petersburg) (examined).

Bohart in Kimsey and Bohart 1991: 440. Replacement name for

Mongolia: Arkhangai, 1 ♀, 90 km NE of Tsetserleg, , 27.VII.2005, leg. JH (MHC); Bayankhongor, 2 ♂♂, 163 km S of Bayankhongor, , 2165 m, 10.VII.2004, leg. JH (MHC); Bulgan, 1 ♂, 170 km W of Ulaanbaatar, dunes, 1070 m, 16.VIII.2007, leg. MH (MHC); Khentii, 1 ♀, 100 km NE of Ondorkhaan, Kerulen River, 970 m, 22.VII.2007, leg. MH (MHC); Selenge, 4 ♀♀, 2 ♂♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Tuv, 2 ♀♀, 4 ♂♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC); 2 ♂♂, ibid, 12.VII.2003, leg. JH (MHC); 10 ♀♀, 6 ♂♂, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MH (MHC, PRC); Zavkhan, 2 ♂♂, 40 km SW of Uliastay, dunes, 18.VII.2005, leg. JH (MHC).

Mongolia (*Arkhangai, *Bayankhongor, *Bulgan, *Khentii, *Selenge, *Zavkhan); Russia (Eastern Siberia) (Rosa et al. 2017a).

This species was previously reported from Mongolia (Rosa et al. 2019) without exact locality.

Dahlbom, 1854

6DBC0BFA-F658-5F25-AC09-CA485AA6C786

Dahlbom, 1854: 307. Holotype ♀; locality unknown [most likely Sweden] (Stockholm) (examined) (

Mongolia: Arkhangai, 1 ♀, 2 ♂♂, 90 km NE of Tsetserleg, , 24.VII.2004, leg. JH (MHC); 1 ♀, 1 ♂, ibid, leg. MK (MHC); 1 ♀, 1 ♂, 70 km NE Tsetserleg, 25.VII.2005, leg. JH (MHC); Selenge, 1 ♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC).

*Mongolia (Arkhangai, Selenge); Asiatic-European, from western Europe to Russia (Rosa et al. 2019).

Linnaeus, 1761

01844BC8-DB38-5BA0-B72A-5A25C8E21806

Linnaeus, 1761: 415. Lectotype ♀ (designated by

Mongolia: (Form A): Arkhangai, 2 ♀♀, 1 ♂, 70 km NE of Tsetserleg, 25.VII.2005, leg. JH (MHC); Selenge, 1 ♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Tuv, 5 ♀♀, 3 ♂♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC); (Form B): Arkhangai, 2 ♀♀, 1 ♂, 70 km NE of Tsetserleg, 25.VII.2005, leg. JH (MHC); 1 ♀, 1 ♂, Chuluut Gol River, , 23.VII.2005, leg. JH (MHC); Tuv, 1 ♀, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC).

*Mongolia (Arkhangai, Selenge, Tuv); Asiatic-European, from Europe to eastern Siberia, Russian Far East and North-East China (Manchuria) (Rosa et al. 2014, 2019).

Two distinct colour forms (Fig. 4) are recorded from Mongolia, Siberia and Primorsky Territory (Russia), and Heilongjiang (China). Form A is matching with the typical European (Fig. 4A, C). Form B is chromatic different without the typical blue colouration on male and female metasoma and with non-metallic black areas on head vertex and mesosoma (Fig. 4B, D). Male T1 golden-greenish, with or without a narrow transversal green or bluish stripe or patch; T2 red, with or without a basal, narrow black stripe; female T1 golden-greenish, with green to bluish colour on T1 frontal declivity to petiolar insertion. This colour variation has also been observed in specimens from Russia (Siberia and Primorsky Territory) and China (Heilongjiang). The Chinese form was mentioned by Linsenmaier (1968) as Linsenmaier, 1968, which is anyway an unnecessary replacement name for Mocsáry, 1912 nec Mocsáry, 1892 (actually male and female of the same taxon). Other evident different morphological characteristics are not recognizable. However, these two forms may represent two sister species, genetically separate, but difficult to identify on the basis of morphological characteristics, as in other known cases of of the group (Paukkunen et al. 2015; Orlovskytė et al. 2016).

Figure 4.

Linnaeus, habitus dorsal view A form A, ♀ B form B, ♀ C form A, ♂ D form B, ♂. Scale bars: 1.0 mm.

(Linnaeus, 1758)

CA3F82A1-F484-5363-83F7-3340C4A7BD14

Linnaeus, 1758: 571. Lectotype ♀ (designated by Richards 1935); Europe (

:

None examined.

The identification of by Buyanjargal and Abasheev (2015) is doubtful and very likely represent another species of the group or even a member of another species group (e.g., group). In fact, the host association with , as observed by the two authors, is unusual. is known as a possible host for members of the group ( and (sub ) Pauli et al. 2019, supplementary file 4). For example, Linsenmaier was erroneously identified as by several authors, including Trautmann (identification label pinned with the type of ).

Mongolia (Bulgan) [doubtful]; West-Palaearctic: from West Europe to central Asia (Linsenmaier 1997b).

Semenov, 1967

A728A8EB-FE41-5160-8B37-087E90107832

Semenov-Tian-Shanskij, 1967: 166. Holotype ♂; Bugas near Khami, SE from Tian Shan [China, Xinjiang] (

Mongolia: Umnugovi, 1 ♂, Deemgin-gobi, 25 km SSO of Khajlastyn-Khuduka, 20.VI.1971, leg. M. Kozlov (ZIN).

*Mongolia (Umnugovi); China (Xinjiang) (Rosa et al. 2014).

Wesmael, 1839

4532A2FC-4222-5E81-9A47-5C0C2D2690C0

Wesmael, 1839: 176. Syntypes ♂♀; Belgium (Bruxelles,

Mongolia: Selenge, 1 ♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Tuv, 1 ♀, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MH (MHC).

*Mongolia (Selenge, Tuv); Asiatic-European, from western Europe to Mongolia (present record).

Semenov, 1967

439A7EA6-B583-5B71-9ACA-E1D1E3A57B6F

Semenov-Tian-Shanskij, 1967: 124. Holotype ♀; Kazakhstan: Balamurun, Karatau Mountain ridge foothills, leg. V. Kozhantschikov (

Mongolia: Dornod, 1 ♂, 100 km W of Choilbalsan, 820 m, 23.VII.2007, leg. MK (MHC); Dornogovi, 5 ♂♂, 65 km SE of Chatan-Bulag, 1020 m, 2.VIII.2007, leg. MH (PRC/MHC); Umnugovi, 1 ♀, Gobi, Dalanzadgad, 24–26.VI.2003, leg. JH (MHC); 12 ♀♀, 70 km S of Saynshand, 1100 m, 6.VIII.2007, leg. MH (PRC/MHC).

*Mongolia (Dornod, Dornogovi, Umnugovi); Kazakhstan (Rosa 2018).

Notes.

As supposed by Rosa (2018), living specimens are red and change to greenish after preparation.

Semenov, 1910

53D00E54-48D9-579F-9FED-E8D99843191A

Semenov-Tian-Shansky, 1910: 222. Lectotype ♂ (designated by

Mongolia: Dornod, 6 ♀♀, 100 km W of Choilbalsan, 820 m, 23.VII.2007, leg. MH (MHC); 9 ♀♀, 2 ♂♂, 20 km W of Choilbalsan, 800 m, 24.VII.2007, leg. MH (MHC); Khentii, 11 ♀♀, 4 ♂♂, 100 km NE of Ondorkhaan, Kerulen River, 970 m, 22.VII.2007, leg. MH (MHC); Selenge, 3 ♀♀, 1 ♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Tuv, 5 ♀♀, 1 ♂, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MH (MHC); 1 ♀, same date and locality, and collector (P. Tyrner priv. coll.); Umnugovi, 1 ♂, Gobi, Dalanzadgad, 25.VI.2003, leg. JH (MHC).

*Mongolia (Dornod, Khentii, Selenge, Tuv, Umnugovi); Asiatic-European, from Greece, Iran, and Turkey to Central Asia (Rosa et al. 2019).

Niehuis, 2000

3C8BE02B-C06D-575C-9780-9B1CF6CF652B

Niehuis, 2000: 192. Holotype ♀; Germany: Rheinland-Pfalz, Monsheim (Frankfurt) (

Mongolia: Tuv, 3 ♀♀, 2 ♂♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH, det. J. van der Smissen and MH (MHC).

*Mongolia (Tuv); Asiatic-European from Europe to Russia (Rosa et al. 2019).

Semenov, 1912

87D6DEE1-93D6-557C-80F8-762EB2D7D7E4

Semenov-Tian-Shanskij, 1912: 192. Lectotype ♂ (designated by Bohart in Kimsey and Bohart 1991: 436); China: Alashan (192 (descr.), depository:

: Kimsey and Bohart 1991: 436 (China [not Mongolia]: Gansu, Quingai, cat.,

Mongolia: Tuv, 1 ♂, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MK (PRC).

*Mongolia (Tuv); China (Gansu, Qinghai, Inner Mongolia) (Rosa et al. 2014).

Semenov, 1967

BE57E98A-32E1-5A22-8A53-6BCCE5D6A040

Semenov-Tian-Shanskij, 1967: 179. Holotype ♀; China: Gansu (

Mongolia: Arkhangai, 1 ♂, 90 km NE of Tsetserleg, , 27.VII.2005, leg. JH (MHC).

*Mongolia (Arkhangai); China (Gansu) (Rosa et al. 2014).

Linsenmaier, 1951

7E117F5F-8980-504D-89F1-8AC4547BE8EB

Linsenmaier, 1951: 76. Lectotype ♀ (designated by

Mongolia: Tuv, 2 ♀♀, Ulaanbaatar Bog Duul, 11.VII.1983, leg. Karl Bleyl, det. Linsenmaier 1992 (NMLS).

*Mongolia (Tuv); Palaearctic region excluding Japan (Linsenmaier 1997b).

Radoszkowski, 1889

6A0D935B-FC48-57B3-A16D-9E29DBD2CD34

:

Mongolia: Holotype ♀, golden rounded label, Kansu Kobden-Owatu 12/VIII [handwritten] Mocsáry [handwritten by Radoszkowski] // Rad. (tres interep.) [?] [handwritten by Mocsáry], label with right flagellum and metasoma, Mus. Pan Krakow [hadwritten by Dylewska].

Mongolia (Khovd) (Radoszkowski 1889).

Linsenmaier, 1959

076C949E-E9F3-5BFF-89EE-AC90C3C3EA13

Linsenmaier, 1959: 165. Holotype ♀; Spain: Zamora (Luzern) (examined) (

Mongolia: Arkhangai, 1 ♀, 1 ♂, 90 km NE of Tsetserleg, , 24.VII.2004, leg. MK (MHC); Bulgan, 2 ♂♂, Mongol Els Nat. Res., dunes, , 31.VII.2005, leg. JH (MHC); Dornogovi, 1 ♀, 28 km of SE Chatan-Bulag, 3.VIII.2007, leg. MHMHC); Tuv, 1 ♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC); 2 ♀♀, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MH (MHC); Zavkhan, 1 ♀, 1 ♂, 40 km SW of Uliastay, dunes, 18.VII.2005, leg. JH (MHC).

*Mongolia (Arkhangai, Bulgan, Dornogovi, Tuv, Zavkhan); from southern Europe to eastern Siberia (Rosa et al. 2017b).

Semenov, 1954

949297CE-690B-5D5A-B643-5D23C246FD4D

Semenov in

: Kimsey and Bohart 1991: 444 (cat., Mongolia: Yihe Bogdo, Peter the Great Range,

Mongolia: Govi-Altai, 4 ♀♀, 1 ♂, Ikhe-Bogdo, Gobi Altai, 30.VI–12.VII.1926, leg. P. Kozlov // Paratypes (ZIN); 4 ♀♀, idem, 15–17.VII.1926, Paratypes (ZIN); 1 ♀, North slope of Ikhe-Bogdo, 30.VI–12.VII.1926, leg. P. Kozlov, Paratypes (ZIN); 1 ♀, Ihe-Bogdo, Gob. Altai, 15–17.VII.1926, leg. P. Kozlov [in Cyrillic], det. M. Nikol’skaya (NMLS); Tuv, 1 ♀, Ulaanbaatar Bog Duul, 11.VII.1983, leg. Karl Bleyl, det. Linsenmaier 1990 (NMLS).

Mongolia (Govi-Altai, Tuv); Tajikistan (Rosa et al. 2017a).

Rosa, 2017

61885804-0C34-59CA-9F6A-79EA9450938B

Rosa in

Mongolia: Govi-Altai, 1 ♂, 10 km SSE of Ich-Oba-Ula, 18.VII.1970, leg. E. Narchuk (ZIN).

*Mongolia (Govi-Altai, Tuv); Russia (western Siberia) (Rosa et al. 2017c).

Rosa, 2018

F6C71551-A6FE-58EA-9B54-9C5E032C3913

Rosa, 2018: 281. Holotype ♂; Mongolia: Govi-Altai Prov., 8 km SE of Argalant-Ula (

Mongolia: Govi-Altai, 1 ♂, 8 km SE of Argalant-Ula, 20.VI.1980, leg. G. Medvedev (ZIN).

Mongolia (Govi-Altai) (Rosa 2018).

Linsenmaier, 1951

237AC968-D491-5D8E-BC76-281C06048A5D

Linsenmaier, 1951: 79. Lectotype ♀ (designated by

:

:

Mongolia: Arkhangai, 1 ♂, 70 km NE of Tsetserleg, 25.VII.2005, leg. JH (MHC); Tuv, 1 ♀, Tereltz, 8.VII.1983, leg. Karl Bleyl, det. Linsenmaier 1992 (NMLS); 2 ♀♀, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MH (MHC).

Mongolia (Arkhangai, Tuv); Asiatic-European, from western Europe to Mongolia (Linsenmaier 1997a).

Semenov, 1967

7E381F1A-3960-520B-8FEE-EF644453A548

Semenov-Tian-Shanskij, 1967: 174. Holotype ♀; Uzbekistan: Termez (

Mongolia: Umnugovi, 1 ♀, “Yuzhno-Gobiyskiy Ajmag, sajr Undyn-Gol, 25 km S of Khan-Bogdo, 7.VII.1971”, leg. M. Kozlov (ZIN).

*Mongolia (Umnugovi); Uzbekistan (Semenov-Tian-Shanskij 1967).

Olivier, 1791

E5B29A70-A3E3-5AFC-B1B3-123163EDDF94

Olivier, 1791: 676. Type unknown; France: Angoumois (depository unknown) (

Mongolia: Tuv, 1 ♀, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MH (MHC).

*Mongolia (Tuv); Palaearctic, from western Europe and North Africa to China and Japan (Linsenmaier 1997b).

Linsenmaier, 1968

7304B10D-C499-5D86-A27A-D9D906C9FCCC

Linsenmaier, 1959: 156, nom. praeocc., nec Mocsáry, 1912. Holotype ♀; Switzerland: Graubünden (Luzern) (examined) (

Linsenmaier, 1968: 99. Replacement name for

Mongolia: Arkhangai, 2 ♀♀, Chuluut Gol River, , 23.VII.2005, leg. JH, det. J. Van der Smissen (MHC); Bulgan, 2 ♀♀, Mongol Els Nat. Res., dunes, , 31.VII.2005, leg. JH, det. J. Van der Smissen (MHC).

*Mongolia (Arkhangai, Bulgan); Asiatic-European, from western Europe to Central Asia, Siberia and Japan (Rosa et al. 2019).

Rosa, 2017

C208046B-73F3-508C-AC60-9EE23C33F069

Rosa in

Mongolia: Arkhangai, 1 ♀, Chuluut Gol River, , 23.VII.2005, leg. JH (MHC).

*Mongolia (Arkhangai); Russia (Eastern Siberia) (Rosa et al. 2017c).

Haupt, 1957

B106AAD1-4734-5029-99D9-B1B84C677B6E

Haupt, 1957: 115. Lectotype ♀ (designated by

Linsenmaier, 1959:

None examined.

Mongolia (Dornod); Asiatic-European, from western Europe to Japan (Linsenmaier 1997b).

Radoszkowski, 1891

770ED8E5-784A-5F85-AE43-D999B483568D

Radoszkowski, 1891: 189. Syntypes ♂, ♀; Turkmenistan: Ashgabad (

Mongolia: Arkhangai, 1 ♂, 90 km NE of Tsetserleg, 27.VII.2005, leg. JH (MHC).

*Mongolia (Arkhangai); Turkmenistan (Radoszkowski 1891).

Linsenmaier, 1959

256A09E7-C5E4-5E29-A0E0-4878B2FCB50A

Linsenmaier, 1959: 160. Holotype ♀; Finland: Kyrkslätt [= Kirkkonummi] (Luzern) (examined) (

Mongolia: Arkhangai, 1 ♂, 70 km NE of Tsetserleg, 25.VII.2005, leg. JH, det. J. Van der Smissen (MHC).

*Mongolia (Arkhangai); Asiatic-European, from western Europe to Central Asia, Russia and Japan (Rosa et al. 2019).

Linnaeus, 1761

261C71FD-11B5-51B4-96FA-284AFA77C85C

Linnaeus, 1761: 415. Type unknown; Sweden (unknown) (

Mongolia: Tuv, 1 ♀, 100 km E of Ulaanbaatar, 20 km NE of Tereltz, Tuul River, 15–21.VII.2003, leg. JH (PRC).

*Mongolia (Tuv); Asiatic-European, from western Europe to Central Asia, Russia, and Japan (Rosa et al. 2019).

Genus Dahlbom, 1845

Dahlbom, 1845: 6. Type species: Fabricius, 1804, by subsequent designation of Bodenstein 1939: 125.

(Dahlbom, 1854)

299D26E3-082D-5C99-9399-2C4FF34C8CBC

Dahlbom, 1854: 146. Lectotype ♀ (designated by Rosa and Xu 2015: 17); Hungary: Budapest (MSNT) (

Mongolia: Zavkhan, 1 ♀, 40 km SW of Uliastay, dunes, 18.VII.2005, leg. JH (MHC).

*Mongolia (Zavkhan); Asiatic-European, from western Europe to Central Asia and western Siberia (Rosa et al. 2019).

Brullé, 1833

F682CC3D-ADAB-51C8-A32D-DFD7C2531284

Brullé, 1833: 375. Holotype ♂ [not ♀]; Greece: Peloponnese (Paris) (examined) (

Mongolia: Zavkhan, 1 ♂, 40 km SW of Uliastay, dunes, 18.VII.2005, leg. JH (MHC).

*Mongolia (Zavkhan); Palaeartcic, from southern Europe and northern Africa to Middle East and Central Asia (Rosa et al. 2019).

Genus Latreille, 1809

Latreille, 1809: 49. Type species: Fabricius, 1787 [= (Fabricius, 1787)], by monotypy.

Tsuneki, 1947

58F1AEE0-8E40-53F1-9E4E-4301DC131A36

Tsuneki, 1947: 54. Holotype ♀; China: Inner Mongolia: Apaka (

:

Semenov, 1967: 189 (cat., Mongolia: 1 ♀, Uburchangaj aimag: Changaj Mt., 8 km W of Somon Chajrchandulaan, 2000 m, Exp. Dr. Z. Kaszab, 1964, nr. 217, 28.VI.1964; 1 ♀, Southgobi aimag: 60 km E of Somon Bulgan, 1120 m, Exp. Dr. Z. Kaszab 1964, nr. 262, 4.VII.1964).

: Kimsey and Bohart 1991: 296 (cat., China [not Mongolia]: Apaka).

:

Mongolia: Govi-Altai, 25 ♀♀, 1 ♂, 70 km E of Altay city, Guulin, 14.VII.2005, leg. JH (MHC, PRC).

Mongolia (*Govi-Altai, Umnugovi, Uvurkhangai); China (Inner Mongolia, Shanxi) (Rosa et al. 2014).

Semenov, 1910

CBC93850-32D7-5A45-B075-7A5AC6976BD3

Semenov-Tian-Shansky, 1910: 214. Lectotype ♂, designated by Kimsey in Kimsey & Bohart 1991: 296; China: Bugas near Khami, SE of Tian-Shan [China, Xinjiang], 3–5.IX.1895, leg. V. Roborovskij & P. Kozlov (

:

:

None examined.

Mongolia (Sukhbaatar); China (Xinjiang) (Rosa et al. 2014).

Genus Lichtenstein, 1876

Lichtenstein, 1876: 227. Type species: Eversmann 1858 [= (Eversmann, 1858)], by subsequent designation of Ashmead 1902: 226

(Eversmann, 1858)

72011E11-19C8-5705-849B-1F2D3F1481EC

Eversmann, 1858: 562. Holotype ♀; Russian SFSR: ‘campis transuralensibus’ (

: Kimsey and Bohart 1991: 521 (Mongolia, without specific locality);

None examined.

Mongolia (without locality); Asiatic-European, from eastern Europe to Mongolia (Kimsey and Bohart 1991).

Genus Semenov, 1891

Semenov, 1891: 444. Type species: Dahlbom, 1845: 6 [= (Dahlbom, 1845)], by subsequent designation of Semenov 1892: 485.

Linsenmaier, 1951: 65 (as subgenus of Latreille, 1809). Type species: Dahlbom, 1854: 149, by original designation. Synonymized by Rosa et al. 2017b

Rosa, 2017

73DDA6E3-505B-5632-89FE-F2339B3E73E8

Rosa in

Mongolia: Bayankhongor, 1 ♀, 1 ♂, 129 km NW of Bayankhongor, , 2590 m, 16.VII.2004, leg. JH (MHC); Bulgan, 4 ♀♀, Mongolia, 137 km NE of Aravaykheer, , 1250 m, 26.VII.2004, leg. JH (PRC/ZIN); 1 ♂, Mongol Els Nat. Res., dunes, , 31.VII.2005, leg. JH (MHC); Dornod, 2 ♂♂, 100 km W of Choibalsan, 820 m, 23.VII.2007, leg. MH (MHC); 3 ♂♂, 20 km W of Choibalsan, , 800 m, 24.VII.2007, leg. MH (MHC); 2 ♂♂, 50 km SW of Choibalsan, 960 m, 25.VII.2007, leg. JH (MHC); Selenge, 2 ♂♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII. 2003, leg. JH (MHC); Sukhbaatar, 1 ♂, 200 km SSE of Baruun-Urt, Moltsoy Els, 1250 m, 27.VII.2007, Allotype, leg. MK (ZIN); 3 ♂♂, ibid, 27.VII.2007, leg. MH (MHC); 2 ♀♀, 7 ♂♂, 100 km SSW of Baruun-Urt, 1100 m, 30.VII.2007, leg. MH (MHC); Tuv, 4 ♂♂, Khangayn Mts, 5 km N of Khunt, 20.VII.2005, leg. JH (MHC); 4 ♂♂, 75 km W of Ulaanbaatar, dunes, 2.VIII.2005, leg. JH (MHC); 13 ♀♀, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. JH (MHC); 8 ♀♀, ibid, leg. MH (MHC).

Mongolia (*Bayankhongor, *Dornod, *Selenge, Sukhbaatar, *Tuv, Bulgan); Russia (Siberia) (Rosa et al. 2017c).

(Shuckard, 1837)

62F88D4F-437D-55AB-B8EB-DA74F2DB614F

Shuckard, 1837: 169. Lectotype ♀ (designated by

Mongolia: Tuv, 1 ♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC).

*Mongolia (Tuv); Holarctic: from west Europe to Turkey, Siberia, Manchuria and Russian Far East (Rosa et al. 2019); North America (Bohart and Kimsey 1982).

Genus Dahlbom, 1854

Dahlbom, 1854: 363. Type species: Dahlbom, 1854 [= (Förster, 1853)], by monotypy.

Rosa & Halada sp. nov.

688E8405-0A11-500A-ABD2-9A33A2377945

http://zoobank.org/A105F4B1-87F4-4005-B1A0-09844A7247B0

Figures 5A, D , 6A, D

Figure 5.

species, females A sp. nov., dorsal view B, dorsal view C, dorsal view D sp. nov., lateral view: arrows pointing at pronotal and propleural spines. Scale bars: 1.0 mm.

Figure 6.

species, females, head in frontal view (A–C), and metasoma in dorsal view (D–F). A sp. nov. BC. D sp. nov. EF. Scale bars: 1.0 mm.

: ♀, Mongolia: Bayankhongor, Edringiyn-Nuru Ridge, 100 km SSW of Bayan-Under, 5.IX.1970, leg. V. Zaitzev (ZIN).

sp. nov. is closely related to Central Asian species of the group, which includes (Mocsáry, 1890), (Semenov, 1901), (Bingham, 1903) and other small species so far considered synonyms of (Kimsey and Bohart 1991). sp. nov. female can be easily separated from all these species by: lateral pronotal area and propleuron ventrally V-shaped carinate, displaying two teeth in lateral view (Fig. 5D) (vs. unmodified in other species); mesopleuron with large and deep scrobal sulcus subtended by large projecting subrectangular carina (Fig. 5D) (vs. U-shaped carina); sparse, deep and large punctures on mesosoma (Fig. 6D), and sparse and deep punctures on metasoma (vs. punctation with dense, shallow and tiny punctures on mesosoma, denser and shallower on metasoma); antennae yellowish, distinctly elongate (Fig. 5A) (vs. black to dark brown, with short to very short flagellomeres); head, in frontal view, transversely subrectangular (Fig. 6A) and not triangular (Fig. 6B); with bulging eyes, similarly to . It is additionally separated from by punctation, elongate and yellowish antennae and bronze body colour (entirely blue body in , with shortened, blackish flagellomeres).

Female. Body length 6.0 mm. Fore wing length 3.8 mm. OOL = 2.3 MOD; POL = 1.9 MOD; MS = 0.7 MOD; relative length of P:F1:F2:F3 = 1.0:1.4:1.0:0.8; subantennal space: 1.4 MOD. Vertex with deep and contiguous punctures, as large as 0.25 MOD; vertex moderately depressed and impunctate in front of anterior ocellus and impunctate laterad of posterior ocelli; median anterior depression developed to upper scapal basin; TFC faint; frons continuous, without two flattened or concave, striate areas; scapal basin almost flat, laterally densely micro-punctate, medially with contiguous punctures forming transverse winkles (Fig. 6A); lower part of scapal basin medially impunctate and sulcate; apex of clypeus discoloured, W-shaped and bent under, medially the folded part measures 0.6 MOD. Malar space very short, distinctly less than 1 MOD. Antennae elongate, with flagellomeres as long as 1.5 × their width. Mouth parts elongate (as long as 0.8 × head length) and evidently protruding from oral fossa. Pronotal groove barely visible; anterolateral corner of the pronotum projected to form an acute humeral angle (Fig. 5A); lateral pronotal area ventrally V-shaped carinate forming an acute tooth (Fig. 5D); propleuron ventrally carinate in a large V-shaped tooth (Fig. 5D). Mesosoma punctation dorsally with large, spaced punctures; interspaces medially polished, laterally micro-punctate; notauli incomplete, visible and deep only basally towards the transscutal fissure; parapsidal furrows fully visible; mesopleuron with a large subrectangular area subtended the mesepimeron + mesepisternum; posterior propodeal projections narrow, acute and downward directed. Wing venation unmodified, with long Rs bending slightly away from costal margin, leaving marginal cell broadly open. Punctation on T1 with tiny, sparse punctures (separated by 1–4 PD) (Fig. 6D), laterally micro-punctate on interspaces; T2 with larger and deeper punctures, anterodorsally denser (0.1–2 PD), laterally micro-punctate on interspaces; T3 with coarse to contiguous small punctures; T3 pit row barely sunken, with small, round pits, equally spaced; posterior pit row area almost polished, with a few, sparse, tiny punctures; T3 with two lateral angles and fully bordered by hyaline margin. Metasomal invaginated T5, T6, and S5 with several dorsal and lateral lobes. S2 black spots oval, transversally placed and separated 0.5 MOD each other. Body coppery-bronze, darker to black on median area of mesoscutum; ventrally golden to copper; tegulae golden to non-metallic yellowish on outer margin; tarsi dark brown. Mandible brown, lighter medially. Scape and pedicel coppery, antennomeres yellowish-orange, darker on distal segments. Legs pale coloured, with slight metallic reflections, with non-metallic proximal and distal joints; tarsi yellowish. Forewings hyaline, slightly amber, with light brown veins. . Whitish, short and sparse setae on head and mesosoma (up to 1.5 MOD long); face with short whitish setae (less than 1.0 MOD); metasoma with short (less than 1. MOD) whitish, sparse setae on T3 and ventrally on S2 and S3 and femora.

Male. Unknown.

The specific epithet (feminine) is derived from the Latin adjective spinosus (thorny) for the long and acute teeth ventrally displayed on pronotum and propleuron and clearly visible in lateral view (Fig. 5D).

Mongolia (Bayankhongor).

(Dahlbom, 1831)

EEFB9EE8-9503-51AA-B34A-B12B7D9AA051

Dahlbom, 1831: 32. Syntypes ♂♂; Sweden: Scania: Lomma and Käflinge [= Kävlinge] (

: Kimsey and Bohart 1991: 552 (cat., Mongolia, without locality).

Mongolia: Govi-Altai, 1 ♂, 70 km E of Altay city, Guulin, 14.VII.2005, leg. JH (MHC).

Mongolia (*Govi-Altai); Asiatic-European: from eastern Europe to Mongolia (Linsenmaier 1959, Kimsey and Bohart 1991).

Genus Spinola, 1806

Spinola, 1806: 9. Type species: Fabricius, 1781, by subsequent desigation of Latreille 1810: 437.

(Fabricius, 1781)

D018A140-B734-5F89-8B9A-0990943D0C66

Fabricius, 1781: 455. Holotype ♀; Russia: Siberia (

Linsenmaier, 1959:

Mongolia: Arkhangai, 9 ♀♀, 25 km NE of Tsetserleg, , 23.VII.2004, leg. JH (MHC); Bulgan, 1 ♀, Mongol Els Nat. Res., dunes, , 31.VII.2005, leg. JH (MHC); Dornogovi, 1 ♂, 28 km SE of Chatan-Bulag, steppe, 3.VIII.2007, leg. MK (PRC); Tuv, 2 ♀♀, 1 ♂, Khangayn Mts, 5 km N of Khunt, 20.VII.2005, leg. JH (MHC); 1 ♂, 75 km W of Ulaanbaatar, dunes, 2.VIII.2005, leg. JH (MHC); Uvurkhangai, 3 ♀♀, 12 km E of Aravaykheer, , 1800 m, 3.VII.2004, leg. JH (MHC).

*Mongolia (Arkhangai, Bulgan, Dornogovi, Tuv, Uvurkhangai); widely distributed in the Palaearctic Region (Tsuneki 1948, Linsenmaier 1959), Russia (Siberia), China (Liaoning, Beijing, Inner Mongolia, Shanxi) (Rosa et al. 2019). Linsenmaier (1997b) mentioned Mongolia in the distribution range of , yet the specimen examined by the Swiss author was collected in China, Inner Mongolia: 1 ♂, Hutjertu Gol [currently Khujirt Gol River, near Bailingmiaozhen monastery, N of Baotou, Inner Mongolia, China] 1927, Sven Hedins Exp. Ctr. Asien Dr. Hummel, det. Linsenmaier 1963 (NMLS).

Lichtenstein, 1876: 27. Type species: Linnaeus, 1758 [= (Linnaeus, 1758)], by monotypy.

(Linnaeus, 1758)

3664516F-0F76-52A0-B871-62051D404D5F

Linnaeus, 1758: 572. Lectotype ♂ (designated by

Mongolia: Bayankhongor, 1 ♀, 16 km SW of Bayankhongor, , 2165 m, 10.VII.2004, leg. JH (MHC); Selenge, 1 ♀, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Tuv, 1 ♀, 100 km E of Ulaanbaatar, 20 km NE Tereltz, Tuul River, 15–21.VII.2003, leg. JH (MHC); 2 ♀♀, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MH (MHC).

*Mongolia (Bayankhongor, Selenge, Tuv); Palaearctic, from western Europe and northern Africa to Central Asia, China and Japan (Rosa et al. 2019).

(du Buysson, 1887)

1C358ADE-4B45-5CE2-B464-CA9C792790D5

du Buysson, 1887: 183. Lectotype ♀ (designated by

Mocsáry, 1889: 323. Unnecessary replacement name for

Mocsáry, 1914: 24. Lectotype ♀ (designated by Bohart in Kimsey and Bohart 1991: 571); Mongolia (HMNH).

Mongolia: 1 ♀, Mocs. typ. det. Mocsáry, red label, L. Buyss. Linsenmaier det. 59, Lectotype Mocs. ♀ RM Bohart, id nr. 135554 HNHM Hym. coll. Paralectotypes: 4 ♀♀, Mongolia, Mocs. typ. det. Mocsáry, red label, L. Buyss. Linsenmaier det. 59, Paralectotype Mocs. ♀ RM Bohart, id nr. 135555–135558 HNHM Hym. coll.

Mongolia (without locality); East-Palaearctic: Russia (Far East), China (Liaoning, Inner Mongolia, Hebei, Beijing, Hunan) (Rosa et al. 2016, 2019).

(Mocsáry, 1912)

846820A8-4737-54AA-A4D8-2A90DB6C28E1

Mocsáry, 1912: 376. Lectotype ♂ (designated by Bohart in

Mongolia: Selenge, 1 ♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC).

*Mongolia (Selenge); Afghanistan, Uzbekistan, China (Xinjiang, Ningxia) (Rosa et al. 2016).

Tribe

Genus Semenov, 1954

Semenov, 1954: 137. Type species: Eversmann, 1858 [= (Eversmann, 1858)], by original designation.

Rosa, 2017

2C119BA7-224A-5337-90D9-A3A7D6D7DF01

Rosa, 2017b: 301. Holotype ♀; Kazakhstan: Aktobe Prov., Mugodzhary Mt., Emba River valley, 17.vi.1985, leg. M. Nesterov (

Mongolia: Dornod, 2 ♀♀, 1 ♂, 20 km W of Choilbalsan, 800 m, , 24.VII.2007, leg. MH (MHC).

*Mongolia (Dornod); Kazakhstan (Rosa 2017b).

Spinola, 1806: 10. Type species: Fabricius, 1804 [= (Fabricius, 1804)], by subsequent designation of Latreille 1810: 437.

Agassiz, 1846: 136. Unjustified emendation of Spinola, 1806 (part.).

Förster, 1853: 351. Type species: Förster, 1853 [= (Lepeletier, 1806)], by subsequent designation of Ashmead 1902: 228.

(Mocsáry, 1889)

049AE56C-772D-5084-9B17-B3684BFCC398

Mocsáry, 1889: 80. Lectotype ♂ (designated by

Mongolia: Arkhangai, 93 ♀♀, 30 ♂♂, Chuluut Gol River, , 23.VII.2005, leg. JH (MHC); Dornogovi, 1 ♂, Orgon, 11.VII.2005, leg. JH (MHC).

*Mongolia (Arkhangai, Dornogovi); Asiatic-European, from eastern Europe, Saudi Arabia, UAE to Central Asia and eastern Siberia (Rosa et al. 2019).

Rosa, 2017

15084A1B-CE65-5126-9378-90CD870E62C2

Rosa in

Mongolia: Sukhbaatar, 1 ♂, Lun-Ula, 30 km WSW of Dariganga, 1.VII.1971, leg. I. Kerzhner (ZIN).

Mongolia (Sukhbaatar); Russia (Tyva Rep.) (Rosa et al. 2017d).

(Mocsáry, 1890)

6486FD88-10D0-55F2-8AB1-693AB14437AC

Mocsáry, 1890: 49. Holotype ♂; Turkey: Buyuk Agri Dagi (Mount Ararat) (

Mongolia: Dornogovi, 6 ♂♂, Orgon, 11.VII.2005, leg. JH (MHC).

*Mongolia (Dornogovi); Turkey (Mocsáry 1890) and Central Asia (unpubl. data).

(Fabricius, 1804)

502428A1-21F3-5429-85F3-203C18346F6D

Panzer, 1798: fig. 51, pl. 11. Type unknown; Germany: Nürnberg (depository unknown), nom. praeocc., nec Fabricius, 1794.

Fabricius, 1804: 172. Replacement name for

Mongolia: Arkhangai, 1 ♂, 25 km NE of Tsetserleg, , 23.VII.2004, leg MK (MHC); Zavkhan, 1 ♀, 2 ♂♂, 40 km SW of Uliastay, dunes, 18.VII.2005, leg. JH (MHC).

*Mongolia (Arkhangai, Zavkhan); Asiatic-European, from western Europe to eastern Siberia, Russian Far East, and China (Heilongjiang) (Rosa et al. 2019).

Gogorza, 1887

7CD44450-8295-5AB2-8411-2D5AD5BAD52C

Gogorza, 1887: 33. Holotype ♂; Spain: Madrid (

:

None examined.

Mongolia (Tuv); Asiatic-European, from Iberian Peninsula to Mongolia and eastern Siberia (Rosa et al. 2019).

(Móczár, 1967)

CB1F0770-9B73-5930-8278-23A5197DFD1F

Móczár, 1967: 185. Holotype ♀; Mongolia: Uvurkhangai aimag: Arc Bogd ul, ca. 20 km S of von Somon Chovd, 1760 m, Exp. Dr. Z. Kaszab, 1964, 22.VI.1964 (

Mongolia: Uvurkhangai, 1 ♀, Uburchangaj aimag, Arc Bogd ul, cca 20 km S of von somon Chovd, 1760 m Exp. Dr. Z. Kaszab, 1964, Nr. 170, 22.VI.1964, sp. nov. ♀ det. Móczár 965, ♀ Pz. F. Linsenmaier det. 1964, Holotype ♀ L. Móczár 1966, Hym. Typ. No. 87 Mus. Budapest, id nr. 134892 HNHM Hym. coll. (HNHM).

Mongolia (Uvurkhangai) (Móczár 1967).

Abeille de Perrin, 1878: 3. Type species: Lepeletier, 1806 [= (Coquebert, 1801)], by subsequent designation of Ashmead 1902: 227.

(Coquebert, 1801)

B1965D70-88B2-5844-A04D-9F09C697D48D

Coquebert, 1801: 59. Holotype ♀; France: Bordeaux (

:

Mongolia: Arkhangai, 1 ♀, 90 km NE of Tsetserleg, , 24.VII.2004, leg. JH (MHC); Bayankhongor, 9 ♀♀, 8 ♂♂, 16 km SW of Bayankhongor, , 2165 m, 10.VII.2004, leg. JH (MHC); Dornod, 2 ♀♀, 3 ♂♂, 100 km W of Choilbalsan, 820 m, 23.VII.2007, leg. MH (MHC); 5 ♀♀, 2 ♂♂, 20 km W of Choilbalsan, 800 m, , 24.VII.2007, leg. MH (MHC); Khentii, 1 ♀, 100 km NE of Ondorkhaan, Kerulen River, 970 m, 22.VII.2007, leg. MH (MHC); Selenge, 2 ♀♀, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Sukhbaatar, 1 ♀, 1 ♂, 100 km SSW of Baruun-Urt, 1100 m, 30.VII.2007, leg. MH (MHC); Ulaanbaatar, 7 ♀♀, Ulaanbaatar, Tuul River valley, 12.VII.2003, leg. JH (MHC); Zavkhan, 2 ♂♂, 40 km SW of Uliastay, dunes, 18.VII.2005, leg. JH (MHC).

Mongolia (*Arkhangai, *Bayankhongor, *Dornod, *Khentii, *Selenge, *Sukhbaatar, *Ulaanbaatar, Uvurkhangai, *Zavkhan); Asiatic-European, from Europe and Middle East to Russia (Far East) (Kimsey and Bohart 1991; Kurzenko and Lelej 2007).

Linsenmaier, 1997

D2CECE62-6B04-5D59-A6ED-53CE40D63EDD

Linsenmaier, 1997a: 254. Holotype ♂, Mongolia: Ulan Bator, 1900 m (

Mongolia: Arkhangai, 1 ♂, 90 km NE of Tsetserleg, , 24.VII.2004, leg. JH (MHC); Bayankhongor, 1 ♂, 16 km SW of Bayankhongor, , 2165 m, 10.VII.2004, leg. JH (MHC); Bulgan, 3 ♂♂, 137 km NE of Aravaykheer, , 1250 m, 2.VII.2004, leg. JS (MHC); 2 ♀♀, 2 ♂♂, 143 km NE of Aravaykheer, , 1300 m, 26.VII.2004, leg. MH (MHC); Ulaanbaatar, 1 ♂, Ulaanbaatar, 16.VII.1989, 1900 m, leg. Peter Salk, det. Linsenmaier, 1997 (NMLS).

Mongolia (*Arkhangai, *Bayankhongor, *Bulgan, Ulaanbaatar); China (Gansu) (Rosa et al. 2014).

was described as a subspecies of . As recently pointed out by Paukkunen et al. (2014), is a synonym of and sensu Linsenmaier (1959) is .

Rosa, 2017

74DD55EE-4F99-5D9C-A1CC-71EACCDE496D

Rosa in

Mongolia: Tuv, 1 ♀, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MK (PRC).

*Mongolia (Tuv); Russia (Eastern Siberia) (Rosa et al. 2019).

(Dahlbom, 1845)

5CBA7E09-D20F-5506-8563-6C0F4E0E3283

Dahlbom, 1845: 3. Lectotype ♀ (designated by

Mongolia: Bayankhongor, 2 ♀♀, 16 km SW of Bayankhongor, , 2165 m, 10.VII.2004, leg. JH (MHC); Dornogovi, 1 ♂, 65 km SE of Chatan-Bulag, 1020 m, 2.VIII.2007, leg. MH (MHC); Govi-Altai, 4 ♀♀, 70 km E of Altay City, Guulin, 14.VII.2005, leg. JH (MHC); Tuv, 2 ♀♀, 1 ♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC); 1 ♀, Khangaun Mts, 5 km N of Khunt, 20.VII.2005, leg. JH (MHC); Umnugovi, 3 ♀♀, Gobi, 100 km SW of Dalanzadgad, Bayanzag, 1–2.VII.2003, leg. JH (MHC); Uvurkhangai, 1 ♂, 12 km E of Aravaykheer, , 1800 m, 3.VII.2004, leg. JH (MHC); Zavkhan, 2 ♀♀, 1 ♂, 40 km SW of Uliastay, dunes, 18.VII.2005, leg. JH (MHC).

*Mongolia (Bayankhongor, Dornogovi, Govi-Altai, Tuv, Umnugovi, Uvurkhangai, Zavkhan); Asiatic-European, from north-western Europe to Mongolia and China (Rosa et al. 2014).

Specimens from Mongolia display an unusual red colouration.

Rosa, 2017

339A9801-E63D-58CF-A7C4-C614B90B745D

Rosa in

Mongolia: Bayankhongor, 18 ♂♂, 22 ♀♀, 75 km S of Bayankhongor, , 1330 m, 8.VII.2004, leg. JH, JS (MHC); Dornogovi, 5 ♀♀, 5 ♂♂, 65 km SE of Chatan-Bulag, 1020 m, 2.VIII.2007, leg. MH (MHC); Tuv, 1 ♂, 70 km W of Ulaanbaatar, 1070 m, dunes, 16.VIII.2007, leg. MH (MHC).

*Mongolia (Bayankhongor, Dornogovi, Tuv); Russia (Eastern Siberia) (Rosa et al. 2019).

Rosa, 2017

C4414CEE-E8DE-5093-9652-CBFAABC69B44

Rosa in

Mongolia: Bayankhongor, 1 ♀, 1 ♂, 56 km NW of Bayankhongor, , 2200 m, 12.VII.2004, leg. JS (MHC); Bulgan, 2 ♀♀, 137 km NE of Aravaykheer, , 1250 m, 2.VII.2004, leg. MK (MHC); 3 ♀♀, ibid, 26.VII.2004, JH (MHC); Dornod, 1 ♂, 100 km W of Choilbalsan, 820 m, 23.VII.2007, leg. MH (MHC); Dornogovi, 1 ♂, 65 km SE of Chatan-Bulag, 1020 m, 2.VIII.2007, leg. MH (MHC); Khentii, 2 ♀♀, 2 ♂♂, 100 km NE of Ondorkhaan, Kerulen River, 970 m, 22.VII.2007, leg. MH (MHC); Selenge, 1 ♀, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Tuv, 10 ♀♀, 9 ♂♂, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MK (PRC); 4 ♂♂, 70 km W of Ulaanbaatar, 1070 m, dunes, 16.VIII.2007, leg. MH (MHC).

*Mongolia (Bayankhongor, Bulgan, Dornod, Dornogovi, Khentii, Selenge, Tuv); Russia (Eastern Siberia) (Rosa et al. 2019).

Rosa, 2017

18B1546C-4CE5-5BCB-A5B3-1885747B0B11

Rosa in

Mongolia: Govi-Altai, 1 ♀, Mongolia W, 70 km E of Altay city, Guulin, 14.VII.2005, leg. JH (MHC).

*Mongolia (Govi-Altai); Russia (Eastern Siberia) (Rosa et al. 2019).

(Rossi, 1790)

83478E37-22EE-54CF-86C3-395A84137EBA

Rossi, 1790: 75. Syntypes; Italy (Berlin?).

Mongolia: Arkhangai, 1 ♂, 90 km NE of Tsetserleg, , 24.7.2004, leg. JH (MHC); Dornod, 21 ♀♀, 4 ♂♂, 100 km W of Choilbalsan, 820 m, 23.VII.2007, leg. MH (MHC); 19 ♀♀, 1 ♂, 20 km W of Choilbalsan, 800 m, , 24.7.2007, leg. MH (MHC); Dornogovi, 4 ♂♂, 2 km SE of Khuvsgol, 5.VIII.2007, leg. MH (MHC); Sukhbaatar, 1 ♀, 1 ♂, 100 km SSW of Baruun-Urt, 1100 m, 30.VII.2007, leg. MH (MHC); 4 ♂♂.

*Mongolia (Arkhangai, Dornod, Dornogovi, Sukhbaatar); Asiatic-European, from western Europe to Russia (Far East) (Rosa et al. 2019).

The record from Korea (Tsuneki 1953b and Korean checklists) must be referred to Linsenmaier, 1959. In fact, Linsenmaier (1959) described as the Korean specimens collected and identified by Tsuneki (1953) as .

Latreille, 1802: 317. Type species: Fabricius, 1775 [= (Scopoli, 1763)], by monotypy.

Dahlbom, 1854

E2991175-823B-5B88-B08F-BFEC4A9FBC84

Dahlbom, 1854: 64. Syntypes ♂♂; Europe: ‘Europa media et meridionali’, Russia, Prussia, Silesia (MfN,

Mongolia: Bayankhongor, 1 ♂, 2 km S of Bayankhongor, , 1800 m, 10.VII.2004, leg. JH (PRC); 20 ♀♀, 2 ♂♂, 56 km NW of Bayankhongor, , 2200 m, 12.VII.2004, leg. JH (PRC); Dornod, 16 ♀♀, 9 ♂♂, 100 km W of Choilbalsan, 820 m, 23.VII.2007, leg. MH (MHC); 16 ♀♀, 6 ♂♂, 20 km W of Choilbalsan, 800 m, 24.VII.2007, leg. MH (MHC); 1 ♂, 15 km W of Choibalsan, Kerulen River, 770 m, 24.VII.2007, leg. MK (PRC); 1 ♂, 50 km SW of Choibalsan, 960 m, 25.VII.2007, leg. JH (MHC); 1 ♀, 50 km SW of Choibalsan, 960 m, 25.VII.2007, leg. MH (MHC); Govi-Altai, 2 ♂♂, 70 km E of Altay city, Guulin, 14.VII.2005, leg. JH (MHC); Selenge, 1 ♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (PRC); Sukhbaatar, 6 ♀♀, 2 ♂♂, 100 km SSW of Baruun-Urt, 1100 m, 30.VII.2007, leg. MH (MHC); Tuv, 1 ♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC); Ulaanbaatar, 1 ♂, Ulaanbaatar, Tuul River valley, 12.VII.2003, leg. JH (PRC); 1 ♀, 7 km E of Ulaanbaatar, Gachuurt, , 31.VII.2002, 1310 m, leg. JS (PRC).

Mongolia (Bayankhongor, *Dornod, *Govi-Altai, *Selenge, Sukhbaatar, Tuv, *Ulaanbaatar); widely distributed in the Palaearctic Region (Linsenmaier 1959; Kurzenko and Lelej 2007), China (Heilongjiang, Inner Mongolia, Gansu) (Rosa et al. 2014).

Chevrier, 1869

4FA957BD-3FFE-5603-8C54-EAD079F88868

Chevrier, 1869: 47. Syntypes ♀♀, ♂♂, [not holotype]; Switzerland (Geneva) (examined).

Mongolia: Khentii, 3 ♀♀, 100 km NE of Ondorkhaan, Kerulen River, 970 m, 22.VII.2007, leg. MH (MHC); Selenge, 1 ♀, 5 ♂♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Tuv, 4 ♀♀, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MH (MHC).

*Mongolia (Khentii, Selenge, Tuv); Palaearctic and Oriental region, from western Europe to Russian Far East, Japan, China and Taiwan (Rosa et al. 2014, 2019).

du Buysson, 1891

98F28AEC-29A3-5A91-9FF7-2EAC92978EB9

du Buysson, 1891: 31. Lectotype ♂ (designated by Kimsey in Kimsey and Bohart 1991: 215); Mongolia: Kansu-Kobden Owatu (

Mongolia: Khovd, 1 ♂, Mongolie O. Radoszkowsky, Kansu-Kobden Owatu, 12.8, Mongolie Coll. R. du Buysson 1900, Type, Museum Paris.

Mongolia (Khovd) (du Buysson 1891).

Abeille de Perrin, 1877

B975A0F8-264A-5644-A238-43EF3DD6BBC8

Abeille de Perrin, 1877: 65. Lectotype ♀ (designated by

Mongolia: Bulgan, 1 ♂, 170 km W of Ulaanbaatar, dunes, 1070 m, 16.VIII.2007, leg. MH (MHC); Dornod, 1 ♂, 50 km SW of Choibalsan, 960 m, 25.VII.2007, leg. JH (MHC); Sukhbaatar, 1 ♂, 200 km SSE of Baruun-Urt, Moltsoy Els, 1250 m, 27.VII.2007, leg. MH (MHC); 2 ♂♂, 6 ♀♀, 100 km SSW of Baruun-Urt, 1100 m, 30.VII.2007, leg. MH (MHC); Tuv, 1 ♀, 75 km W Ulaanbaatar, dunes, 2.VIII.2005, leg. JH (MHC); Umnugovi, 1 ♀, Gobi Gurvansaikhan National Park, , 10.VII.2005, leg. JH (MHC).

*Mongolia (Bulgan, Dornod, Sukhbaatar, Tuv, Umnugovi); Palaearctic, from southern Europe and northern Africa, to western Asia, Siberia, and China (Rosa et al. 2019).

(Scopoli, 1763)

EA24A491-3936-5BB7-8D89-3F4109B386E0

Scopoli, 1763: 297. Holotype ♀; Italy [not Austria] (lost).

Mongolia: Arkhangai, 1 ♀, 25 km NE of Tsetserleg, , 23.VII.2004, leg. JH (MHC); 4 ♂♂, 90 km NE of Tsetserleg, , 24.VII.2004, leg. JH (MHC); 4 ♀♀, 9 ♂♂, ibid, 27.VII.2005, leg. JH (MHC); Selenge, 2 ♀♀, 12 ♂♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Tuv, 2 ♀♀, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MH (MHC); Zavkhan, 3 ♂♂, 40 km SW of Uliastay, dunes, 18.VII.2005, leg. JH (MHC).

*Mongolia (Arkhangai, Selenge, Tuv, Zavkhan); Asiatic-European, from western Europe to Siberia (Rosa et al. 2019).

Semenov, 1967

82CFF819-5B80-53B1-BFA2-87CAF48DD9E1

Semenov-Tian-Shanskij, 1967: 142. Holotype ♂; Russia: Siberia, Shira Lake [Khakass Rep.], 24.VII.1897, Yu. Wagner (

Mongolia: Arkhangai, 1 ♀, 90 km NE of Tsetserleg, , 24.VII.2004, leg. JH (MHC); 1 ♀, 2 ♂♂, 90 km NE of Tsetserleg, , 27.VII.2005, leg. JH (MHC); Dornod, 2 ♂♂, 20 km W of Choibalsan, , 800 m, 24.VII.2007, leg. MH (MHC); Selenge, 1 ♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Tuv, 1 ♀, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MK (PRC); 7 ♀♀, 3 ♂♂, ibid, 8–13.VIII.2007, leg. MH (MHC).

Mongolia (*Arkhangai, *Dornod, *Selenge, Sukhbaatar, *Tuv); Russia (Siberia, Far East) (Rosa et al. 2019).

Dahlbom, 1845: 4. Type species: Dahlbom, 1845, by subsequent designation of Ashmead 1902: 227.

Trautmann, 1926

7A268874-78C1-5ED1-8276-6C68833AB6C7

Trautmann, 1926: 5. Holotype ♀; Turkey: İzmir prov.: Smyrna (

Mongolia: Selenge, 3 ♀♀, 5 ♂♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC).

*Mongolia (Selenge); Asiatic-European, from southern Europe to China (Rosa et al. 2019).

Móczár, 1967

ED915242-2305-599A-92B6-74F7EDDFDADD

Móczár, 1967: 187. Holotype ♂; Mongolia: Ostgobi aimag 40 km NW of Chara-Eireg 1150 m Exp. Dr. Z. Kaszab, 1963 (

Mongolia: Dornogovi, 1 ♂, Ostgobi aimag 40 km NW of Chara-Eireg, 1150 m Exp. Dr. Z. Kaszab, 1963, Nr. 62, 30.VI.1963, Holotype ♂ n. sp. det. Móczár 1966, Hym. Typ. No. 89, id nr. 134927 HNHM Hym. coll. (HNHM); 1 ♀, Ostgobi aimag 40 km NW of Chara-Eireg 1150 m Exp. Dr. Z. Kaszab, 1963, Nr. 62, 30.VI.1963, Allotype ♀ n. sp. det. Móczár 1966, Hym. Typ. No. 96, id nr. 134933 HNHM Hym. Coll. (HNHM). 1 ♂: Ostgobi aimag 40 km NW of Chara-Eireg 1150 m Exp. Dr. Z. Kaszab, 1963, Nr. 62, 30.VI.1963, sp. n. ? , ♂ Allotype Dhlb. diversicolor Lins. Linsenmaier 1966, Paratype ♂ n. sp. det. Móczár 1966, Hym. Typ. No. 90 (HNHM); 1 ♂, Mongolia, Ostgobi aimag 40 km NW of Chara-Eireg 1150 m Exp. Dr. Z. Kaszab, 1963, Nr. 62, 30.VI.1963, Paratype ♂ n. sp. det. Móczár 1966, Hym. Typ. No. 91, id nr. 134929 HNHM Hym. coll. (HNHM); 1 ♂: Mongolia, Ostgobi aimag 40 km NW of Chara-Eireg 1150 m Exp. Dr. Z. Kaszab, 1963, Nr. 62, 30.VI.1963, Paratype ♂ n. sp. det. Móczár 1966, Hym. Typ. No. 93, id nr. 134930 HNHM Hym. coll. (HNHM); 1 ♂, Mongolia, Ostgobi aimag 40 km NW of Chara-Eireg 1150 m Exp. Dr. Z. Kaszab, 1963, Nr. 62, 30.VI.1963, Paratype ♂ n. sp. det. Móczár 1966, Hym. Typ. No. 94, id nr. 134931 HNHM Hym. coll. (HNHM); 1 ♀: Mongolia, Ostgobi aimag 40 km NW of Chara-Eireg 1150 m Exp. Dr. Z. Kaszab, 1963, Nr. 62, 30.VI.63, ? det. L. Móczár, Type Dhlb. diversicolor Lins. Linsenmaier 1966, Paratype ♀ n. sp. det. Móczár 1966, Hym. Typ. No. 95 / id nr. 134932 HNHM Hym. coll. (HNHM); 1 ♂, Mongolia, Catgobi aimag 40 Km NW of Chara-Eireg 1150 m, Exp.Dr. Z. Kaszab, 1963, Nr.62, 30.VI.1963, Paratype (NMLS); 1 ♀, Mongolia, Catgobi aimag 40 Km NW of Chara-Eireg, 1150 m, Exp. Dr. Z. Kaszab, 1963 / Nr.62, 30.VI.1963, Paratype (NMLS); Umnugovi, 1 ♂, Gobi, Dalanzadgad, 25.6.2003, leg. JH (MHC); Uvurkhangai, 1 ♂, 139 km SW of Aravaykheer, , 1430 m, 4.VII.2004, leg. JS (MHC).

Mongolia (Dornogovi, *Umnugovi, *Uvurkhangai) (Móczár 1967).

Linsenmaier, 1959

B223D605-87A7-598A-B882-8FD677537D97

Linsenmaier, 1959a: 31. Holotype ♀; Turkey: Niğde prov.: Niğde (

Mongolia: Dornod, 22 ♀♀, 14 ♂♂, 100 km W of Choilbalsan, 820 m, 23.VII.2007, leg. MH (MHC); 25 ♀♀, 6 ♂♂, 20 km W of Choilbalsan, 800 m, 24.VII.2007, leg. MH (MHC); Sukhbaatar, 1 ♂, 200 km SSE of Baruun-Urt, Moltsoy Els, 1250 m, 27.VII.2007, leg. MH (MHC).

*Mongolia (Dornod, Sukhbaatar); Asiatic-European, from central Europe to eastern Siberia (Rosa et al. 2019).

Genus Panzer, 1801

Panzer, 1801: 13. Type species: Fabricius, 1787, by monotypy.

(Fabricius, 1787)

F4EAF101-00E8-5CC4-B51B-64BD794EECD0

Fabricius, 1787: 284. Holotype ♀; Germany: Hala Saxonum [= Halle] (Copenhagen) (examined).

Mongolia: Tuv, 1 ♀, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC); Ulaanbaatar, 1 ♀, Ulaanbaatar, Tuul River valley, 12.VII.2003, leg. JH (MHC).

*Mongolia (Tuv, Ulaanbaatar); Holarctic and Oriental: from Europe and North Africa to Japan and Taiwan (Wei et al. 2014). Probably accidentally introduced to North America (Kimsey and Bohart 1991).

(Semenov, 1932)

47569CF4-012C-5877-831B-8F8249F440C4

Semenov-Tian-Shanskij, 1932: 12. Holotype ♀; China: Sichuan (

Mongolia: Ulaanbaatar, 1 ♂, Ulaanbaatar, Tuul River valley, 12.VII.2003, leg. JH (MHC).

*Mongolia (Ulaanbaatar); East-Palaearctic: Russia (Eastern Siberia, Far East), China (Ningxia, Sichuan) (Rosa et al. 2019).

(Semenov, 1932)

E31AB170-F5C6-5FBD-BDAF-87400BD505B7

Semenov-Tian-Shanskij, 1932: 15. Lectotype ♀ (designated by

Mongolia: Arkhangai, 1 ♀, 90 km NE of Tsetserleg, , 27.VII.2005, leg. JH (MHC); Bayankhongor, 1 ♀, 75 km S of Bayankhongor, , 1330 m, 8.VII.2004, leg. JS (MHC); Bulgan, 2 ♂♂, 137 km NE of Aravaykheer, , 1250 m, 2.VII.2004, leg. JH (MHC); 1 ♀, 1 ♂, 143 km NE of Aravaykheer, , 1300 m, 26.VII.2004, leg. MH (MHC); 2 ♂♂, Mongol Els Nat. Res., dunes, , 31.VII.2005, leg. JH (MHC); Sukhbaatar, 2 ♀♀, 100 km SSW of Baruun-Urt, 1100 m, 30.VII.2007, leg. MH (MHC); Tuv, 1 ♂, 80 km W of Ulaanbaatar, 1230 m, dunes, 17.VIII.2007, leg. MH (MHC).

*Mongolia (Arkhangai, Bayankhongor, Bulgan, Sukhbaatar, Tuv); Central Asia (Kimsey and Bohart 1991).

(Semenov, 1932)

F908B839-05A2-5BC7-B4CD-7AFEDCB1C90D

Semenov-Tian-Shanskij, 1932: 13. Lectotype ♀ (designated by

Mongolia: Bayankhongor, 1 ♂, 75 km S of Bayankhongor, , 1330 m, 8.VII.2004, leg. JH (MHC); Dornogovi, 2 ♀♀, Orgon, 11.VII.2005, leg. JH (MHC); Sukhbaatar, 2 ♀♀, 200 km SSE of Baruun-Urt, Moltsoy Els, 1250 m, 27.VII.2007, leg. MH (MHC).

*Mongolia (Bayankhongor, Dornogovi, Sukhbaatar); Central Asia (Kimsey and Bohart 1991).

(Semenov, 1932)

4305C1CA-26A7-5417-B7CB-603E941ABDC8

Semenov-Tian-Shanskij and Nikol’skaya, 1954: 93. Lectotype ♀ (designated by

Mongolia: Arkhangai, 1 ♀, 90 km NE of Tsetserleg, , 24.VII.2004, leg. JH (MHC); Tuv, 3 ♂♂, 75 km W of Ulaanbaatar, dunes, 2.VIII.2005, leg. JH (MHC); Uvurkhangai, 1 ♀, 159 km of SW Aravaykheer, , 1250 m, 5.VII.2004, leg. JH (MHC).

*Mongolia (Arkhangai, Tuv, Uvurkhangai); Central Asia (Kimsey and Bohart 1991).

Genus Abeille de Perrin, 1879

Abeille de Perrin, 1879: 27. Type species: Abeille de Perrin, 1879 [= (Klug, 1835)], by subsequent designation of Ashmead 1902: 228.

Agassiz, 1846: 136. Unjustified emendation of Spinola, 1806 (part.).

(Radoszkowski, 1877)

BDDA00C1-24C2-52AF-A99B-A0246391A210

Radoszkowski, 1877: 5. Holotype ♂; Uzbekistan: Zarafshan (

Mongolia: Arkhangai, 1 ♂, 90 km NE of Tsetserleg, , 27.7.2005, leg. JH (MHC).

*Mongolia (Arkhangai); Asiatic-European, from southern Europe, western Asia, Iran, and Turkey to Mongolia (Rosa et al. 2013; present record).

Rosa, 2017

FECC811F-AE39-54B2-A96D-6DEC9D18E368

Rosa in

Mongolia: Bayankhongor, 1 ♀, 1 ♂, 75 km S of Bayankhongor, , 1330 m, 8.VII.2004, leg. JH (MHC); 1 ♀, 56 km NW of Bayankhongor, , 2200 m, 12.VII.2004, leg. JS (MHC); 3 ♀♀, 7 ♂♂, 86 km NW of Bayankhongor, , 2070 m, 14.VII.2004, leg. JH, MK (MHC); Sukhbaatar, 1 ♀, SE Mongolia, 200 km SSE of Baruun-Urt, Moltsoy Els, 1250 m, 27.VII.2007, leg. MK, paratype (PRC); 1 ♂, ibid, leg. MH (MHC); Ulaanbaatar, 2 ♂♂, Ulaanbaatar, Tuul River valley, 12.VII.2003, leg. JH (MHC); Uvurkhangai, 4 ♂♂, 12 km E of Aravaykheer, , 1800 m, 3.VII.2004, leg. JH (MHC);

Mongolia (*Bayankhongor, Sukhbaatar, *Ulaanbaatar, *Uvurkhangai); Russia (Tyva Rep.) (Rosa et al. 2017c).

(Semenov, 1932)

9A776651-90CF-5AA0-B0D9-9A731FE3E0AE

Semenov-Tian-Shanskij, 1932: 34. Holotype ♀; Kazakhstan: “Turkestan septentr.: Bajgakum (

Mongolia: Dornogovi, 1 ♀, 1 ♂, Orgon, 11.VII.2005, leg. JH (MHC).

*Mongolia (Dornogovi); Central Asia (Kimsey and Bohart 1991).

Rosa, 2017

94CBFEEC-2798-533A-A6C0-8004628A77F3

Rosa in

Mongolia: Dornogovi, 1 ♂, Atayn Mts, Gichigniv Nuruu, 10 km SW of Sain-Shand, 12.VII.2005, leg. JH (MHC).

*Mongolia (Dornogovi); Russia (East Siberia) (Rosa et al. 2017c).

(du Buysson, 1901)

2E72708C-2D43-546D-B26E-0E43FB88D320

du Buysson, 1901: 98. Lectotype ♂ (designated by

(!) var.

:

:

: Kimsey and Bohart 1991: 256 (cat., North Mongolia).

:

Mongolia: Arkhangai, 1 ♂, 25 km NE of Tsetserleg, , 23.VII.2004, leg. JH (MHC); Bayankhongor, 2 ♂♂, 16 km SW of Bayankhongor, , 2165 m, 10.VII.2004, leg. JH (MHC); 1 ♂, 56 km NW of Bayankhongor, , 2200 m, 12.VII.2004, leg. JH (MHC); 1 ♂, 86 km NW of Bayankhongor, , 2070 m, 14.VII.2004, leg. JH (MHC); Khentii, 2 ♀♀, 100 km NE of Ondorkhaan, Kerulen River, 970 m, 22.VII.2007, leg. MH (MHC); Selenge, 1 ♀, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Sukhbaatar, 1 ♀, SE Mongolia, 200 km SSE of Baruun-Urt, Moltsoy Els, 1250 m, 27.VII.2007, MK (PRC); Tuv, 1 ♀, 50 km N of Ulaanbataar, E of Mandal, 1180 m, 8–13.VII.2007, leg. MK (PRC); 1 ♀, 1 ♂, ibid, leg. MH (MHC); Uvurkhangai, 1 ♂, N. Mongolei Leder 92, Buyss. var. nov. R. du Buysson det. 1901 ♂, Lectotype v. Buysson det. L. Móczár (NHMW). Paralectotypes: 1 ♂ N. Mongolei Leder 92, Buyss. var. nov. R. du Buysson det. 1901 ♂, Paralectotype v. Buysson det. L. Móczár (NHMW); 1 ♀, N. Mongolei Leder 92, Buyss. var. nov. R. du Buysson det. 1901 ♀, Mocs. det. L. Móczár (NHMW).

Mongolia (*Arkhangai, *Bayankhongor, *Khentii, *Sukhbaatar, *Tuv, Ulaanbaatar, Uvurkhangai); widely distributed from Mongolia to Central Asia and southern Russia to Volga (Trautmann 1927), China (Shanxi) (Rosa et al. 2014).

(Semenov, 1932)

122EF46C-0557-5897-B18D-6EBF6EB68780

Semenov-Tian-Shanskij, 1932: 24. Lectotype ♀ (designated by

: Móczár, 1967: 186 (cat., Mongolia: 1 ♂, Ostgobi aimag; 40 km NW of Chara-Eireg, 1150 m, Exp. Dr. Z. Kaszab, 1963, nr. 62, 30.VI.1963; 1 ♂, Ostgobi aimag: 20 km SO of Čojren, 1200 m, Exp. Dr. Z. Kaszab, 1963, nr. 70, 1.VII.1963; 1 ♂, Sukhbaatar aimag: 44 km SSW of Baruun urt, 1050 m, Exp. Dr. Z. Kaszab, 1965, nr. 353, 3.VIII.1965; 1 ♀, Chentej aimag: 10 km W of Somon Delgerchaan, 1250 m, Exp. Dr. Z. Kaszab, 1965, nr. 476, 24.VIII.1965, allotype).

: Kimsey and Bohart 1991: 257 (cat., Mongolia: Urga).

Mongolia: Bayankhongor, 5 ♀♀, 11 ♂♂, 86 km NW of Bayankhongor, , 2070 m, 14.VII. 2004, leg. JS, MK, JH (MHC); Dornod, 1 ♀, 50 km SW of Choilbalsan, 960 m, 25.VII.2007, leg. JH (MHC); Govi-Altai, 1 ♀, 1 ♂, 70 km of Altay city, Guulin, 14.VII.2005, leg. JH (MHC); Govi-Sümber, 2 ♂♂, 20 km SE of Choyr, 1480 m, 7.VIII.2007, leg. MK (PRC); 1 ♀, 1 ♂, ibid, leg. MH (MHC); Sukhbaatar, 4 ♂♂, 200 km SSE of Baruun-Urt, Moltsoy Els, 1250 m, 27.VII.2007, leg. MK (PRC); 1 ♂, 210 km SSE of Baruun-Urt, 29.VII.2007, steppe, leg. MK (PRC); 12 ♂♂, 100 km SSW of Baruun-Urt, 1100 m, 30.VII.2007, leg. MH (MHC); Tuv, 1 ♀, Teregtin, 1350 m, Exp. Dr. Z. Kaszab, 1963, Nr.73, 2.VII.1963, det. Linsenmaier 1966 (NMLS); Ulaanbaatar, ♀ [not ♂], env. Urga [Ulaanbaatar], 1–4.VI.1909, leg. P. Kozlov, Ellamp. shokalskii m. [mihi] Typ. ♂. A. Semenov-Tian-Shansky det. V.19, Lectotype Sem. design. LS Kimsey (ZIN); 1 ♀, same data, paralectotype (ZIN); Umnugovi, 1 ♀, Gobi Gurvansaikhan National Park, 40 km W of Dalanzadgad, 2000 m, 28–30.VI. 2003, leg. JH (MHC); Uvurkhangai, 9 ♂♂, 12 km E of Aravaykheer, , 1800 m, 3.VII.2004, leg. JH (MHC).

Mongolia (*Bayankhongor, *Dornod, Dornogovi, *Govi-Sümber, Khentii, Sukhbaatar,*Tuv, Ulaanbaatar, *Umnugovi, Uvurkhangai) (Kimsey 1986).

Genus Ashmead, 1902

Ashmead, 1902: 229. Type species: Aaron, 1885 [= (Haldeman, 1844)], by original designation.

(Tsuneki, 1953)

935EF421-D5F4-5670-81C1-D29CF280862A

Tsuneki, 1953a: 54. Syntypes ♂, ♀; Japan, Korea, Manchuria (

Mongolia: Bulgan, 1 ♀, 137 km NE of Aravaykheer, , 1250 m, 2.VII.2004, leg. JS (MHC); 1 ♀, Mongol Els Nat. Res., dunes, , 31.VII.2005, leg. JH (MHC); Khentii, 1 ♀, 100 km NE of Ondorkhaan, Kerulen River, 970 m, 22.VII.2007, leg. MH (MHC); Tuv, 2 ♀♀, 18 ♂♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC); Zavkhan, 2 ♀♀, 40 km SW of Uliastay, dunes, 18.VII.2005, leg. JH (MHC).

*Mongolia (Bulgan, Khentii, Tuv, Zavkhan); Russia (Eastern Siberia, Far East), China, Korea, Japan (Rosa et al. 2019).

(Uchida, 1927)

6B40105F-768D-5161-A13A-2F46F6123712

Uchida, 1927: 153. Holotype ♂; Korea: Seiryori (descr.) (

du Buysson, 1908:

Mongolia: Bulgan, 1 ♀, Mongol Els Nat. Res., dunes, , 31.VII.2005, leg. JH (MHC); Dornod, 1 ♀, 100 km W of Choilbalsan, 820 m, 23.VII.2007, leg. MH (MHC); 2 ♂♂, 20 km W of Choilbalsan, 800 m, , 24.VII.2007, leg. MH (MHC/PRC); Khentii, 1 ♀, 1 ♂, 100 km NE of Ondorkhaan, Kerulen River, 970 m, 22.VII.2007, leg. MH (MHC); Selenge, 7 ♀♀, 5 ♂♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC/PRC); Tuv, 7 ♀♀, 3 ♂♂, 50 km N of Ulaanbaatar, E of Mandal, 1180 m, 8–13.VIII.2007, leg. MH (MHC, PRC); Ulaanbaatar, 3 ♀♀, Ulaanbaatar, Tuul River valley, 12.VII.2003, leg. JH (MHC).

Mongolia (*Bulgan, *Dornod, *Khentii, *Selenge, Tuv, *Ulaanbaatar); Russia (Eastern Siberia, Far East); Japan (Hokkaido) (Rosa et al. 2019).

The specimen illustrated in the volume of Russian cuckoo wasps (Rosa et al. 2019: fig. 18) is apparently misidentified and currently belonging to an unidentified species. Examination of type material by Uchida is needed for further studies.

(Uchida, 1927)

3E3FC438-A781-5016-9809-7985AFE51E92

Uchida, 1927: 152. Syntypes ♂, ♀; Japan: Hokkaido and Honshu (

: Móczár, 1967: 185 (cat., Mongolia: 1 ♀, 1 ♂, Čojbalsan [= Dornod] aimag: Chamardavaa ul, 80 km SO of Somon Chalchingol, 600 m, Exp. Dr. Z. Kaszab, 1965, nr. 401, 13.VIII.1963).

Mongolia: Bulgan, 1 ♀, Mongol Els Nat. Res., dunes, , 31.VII.2005, leg. JH (MHC); Khentii, 14 ♀♀, 100 km NE of Ondorkhaan, Kerulen River, 970 m, 22.VII.2007, leg. JH, MH (MHC); Tuv, 18 ♂♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC).

Mongolia (*Bulgan, Dornod, *Khentii, *Tuv); Russia (Eastern Siberia, Far East); Korea, Japan (Rosa et al. 2019).

(Fabricius, 1804)

6260FC01-21A3-55BE-B156-45A6B3395518

Fabricius, 1804: 176. Lectotype ♀ (designated by

: Móczár, 1967: 195 (cat., Mongolia: 1 ♀, Čojbalsan [= Dornod] aimag: 50 km SO of Čojbalsan [= Choibalsan], 700 m, Exp. Dr. Z. Kaszab, 1965, nr. 421, 16.VIII.1965; 1 ♂, Čojbalsan [= Dornod] aimag: 44 km NW of Čojbalsan [= Choibalsan], 750 m, Exp. Dr. Z. Kaszab, 1965, nr. 425, 17.VIII.1965).

Mongolia: Bayankhongor, 2 ♀♀, 16 km SW of Bayankhongor, , 2165 m, 10.VII.2004, leg. JS (MHC); Bulgan, 2 ♀♀, 137 km NE of Aravaykheer, , 1250 m, 2.VII.2004, leg. JS (MHC); Dornod, 4 ♀♀, 5 ♂♂, 100 km W of Choilbalsan, 820 m, 23.VII.2007, leg. MH (MHC); 4 ♀♀, 20 km W of Choilbalsan, 800 m, 24.VII.2007, leg. MH (MHC); 2 ♂♂, 50 km SW of Choilbalsan, 960 m, 25.VII.2007, leg. JH (MHC); Khentii, 3 ♀♀, 5 ♂♂, 100 km NE of Ondorkhaan, Kerulen River, 970 m, 22.VII.2007, leg. MH (MHC); Selenge, 2 ♂♂, 90 km N of Ulaanbaatar, Segnez River, 1450 m, 6–8.VII.2003, leg. JH (MHC); Tuv, 2 ♂♂, 50 km E of Ulaanbaatar, Tuul River, 22.VI.2003, leg. JH (MHC); Umnugovi, 3 ♀♀, 12 ♂♂, Gobi Gurvansaikhan National Park, 40 km W of Dalanzadgad, 2000 m, 28–30.VI.2003, leg. JH (MHC); Uvurkhangai, 4 ♀♀, 13 ♂♂, 12 km E of Aravaykheer, , 1800 m, 3.VII.2004, leg. JH (MHC).

Mongolia (*Bayankhongor, *Bulgan, Dornod, *Khentii, *Selenge, *Tuv, *Umnugovi, *Uvurkhangai); Palaearctic, from western Europe and northern Africa to Russian Far East (Kurzenko and Lelej 2007).

Genus Latreille, 1797

Latreille, 1797: 126. Type species: Fabricius, 1775 [= (Pallas, 1771)], by monotypy.

Semenov, 1901

F1D126B9-5C46-5E88-8C82-1E798FC161C3

Semenow, 1901: 25. Holotype ♂; Tajikistan: “Turkestan occid.[entalis]: Jagnob: Rovat, 12.VII.1892, leg. D. Glasunow” (

Mongolia: Khovd, 1 ♂, ur. Elkhon, 20 km SE of Altai, Bodoncha, 27.V.1970, leg. E. Narchuk (ZIN).

*Mongolia (Khovd); Central Asia, Russia (south of European part) (Rosa et al. 2019).

Eversmann, 1858

5F9B12C8-F505-5FB1-A538-5E512D7429EC

Eversmann, 1858: 567. Holotype ♀; Russia: Siberia “campis orientalibus” (

Mongolia: Arkhangai, 1 ♂, 25 km NE of Tsetserleg, , 23.VII.2004, leg. JH (MHC); 1 ♂, 90 km NE of Tsetserleg, , 24.VII.2004, leg. JH (MHC); 1 ♀, ibid, 27.VII.2005, leg. JH (MHC); Bulgan, 1 ♂, 143 km NE of Arvaykheer, , 26.VII.2004, 1300 m, sandy dunes, JS (PRC); 4 ♀♀, 3 ♂♂, Mongol Els Nat. Res., , dunes, 1320 m, 31.VII.2005, leg. JH (PRC); Dornod, 2 ♂♂, 50 km SW of Choibalsan, 960 m, 25.VII.2007, leg. JH (MHC); Dornogovi, 1 ♀, 2 ♂♂, 28 km SE of Chatan-Bulag, 3.VIII.2007, leg. MH (MHC); Sukhbaatar, 4 ♀♀, 4 ♂♂, 200 km SSE of Baruun-Urt, Moltsoy Els, 1250 m, 27.VII.2007, leg. MH (MHC); 1 ♂, 3 ♀♀, ibid, 27.VII.2007, leg. JH (MHC); 2 ♂♂, 100 km SSW of Baruun-Urt, 1100 m, 30.VII.2007, leg. MH (MHC); Tuv, 1 ♀, 75 km W of Ulaanbaatar, dunes, 2.VIII.2005, leg. JH (MHC); Umnugovi, 1 ♀, Gobi, Dalanzadgad, 25.VI.2003, leg. JH (MHC).

Mongolia (*Arkhangai, *Bulgan, *Dornod, *Dornogovi, *Sukhbaatar, *Tuv, *Umnugovi); China (Heilongjiang, Shanghai, Shandong), Korea, Russia (Rosa et al. 2014).

Species to be excluded from Mongolian fauna

The following 19 taxa were described or listed for Mongolia by Radoszkowski (1887, 1891), Mocsáry (1890), Dalla Torre (1892), du Buysson (1893), Bischoff (1913), Linsenmaier (1959), and Kimsey and Bohart (1991), yet the type localities are situated in Inner Mongolia (China) or Central Asian countries. These species are expected in Mongolia due to the close vicinity of the collecting localities, with the only exception of (du Buysson, 1909), which belongs to the Oriental fauna.

(!) Radoszkowski, 1887: 45. Holotype ♀; Mongolia [= China]: Qinghai: Zaïdam (ISEA-PAS) (examined).

: Mocsáry 1889: 80. Justified emendation of Radoszkowski, 1887. Dalla Torre 1892: 14 (cat., Mongolia [= China]).

: Dalla Torre 1892: 14 (cat., Mongolia [= China]); Kimsey and Bohart 1991: 168 (cat., Mongolia [= China]: Zaidam).

: Bischoff 1913: 6 (cat., Mongolia [= China]).

: Linsenmaier 1959: 16 (key), 24 (tax., descr., Mongolia [= China]).

Mocsáry, 1890: 49. Holotype ♀; Mongolia [= China, Inner Mongolia]: Mongolia meridionalis (Ta-Wan) (ISEA-PAS) (examined).

: Dalla Torre 1892: 18 (cat., Mongolia [= Inner Mongolia]).

: Kimsey and Bohart 1991: 171 (cat., Mongolia [= Inner Mongolia]: Ta-Wan).

: Bischoff 1913: 7 (cat., Mongolia [= Inner Mongolia]).

: Linsenmaier 1959: 16 (key), 24 (descr., Mongolia [= Inner Mongolia]).

Tsuneki, 1947: 47. Holotype ♀; China: Inner Mongolia: Apaka (NIAS).

: Kimsey and Bohart 1991: 188 (cat., Mongolia [= China, Inner Mongolia]: Apaka).

Mocsáry in Radoszkowski 1889: 10, nec Brullé, 1846. Lectotype ♀ (designated by French, in Bohart and French 1986: 341); China “Ta-schian-sy” (HNHM) (examined).

Mocsáry, 1889: 157. Replacement name for Radoszkowski, 1889, nec Brullé, 1846.

Tsuneki, 1947: 54. Syntypes ♀♀, ♂♂; China: Inner Mongolia: Apaka (NIAS).

: Linsenmaier 1959: 39 (descr., key, Mongolia [= China, Inner Mongolia]); Kimsey and Bohart 1991: 220. Mongolia (cat., without locality, related to the record of from Inner Mongolia by Tsuneki 1947).

du Buysson, 1893: 246. Syntypes ♀♀; Mongolia [= China, Inner Mongolia], Kansu-Jelisyn-Kuse (ISEA-PAS); Persia (MNHN) (examined). Bischoff 1913: 8 (cat., Mongolia [= China, Inner Mongolia])

: Kimsey and Bohart 1991: 248. Incorrect subsequent spelling.

: Rosa et al. 2015: 77.

: Farhad et al. 2018: 199. Lectotype designation: ♂; China: Kansu Jelisyn Kuse (ISEA-PAS).

Semenov in Semenov-Tian-Shanskji and Nikol’skaya, 1954: 93. Holotype ♂; China: Sandzhu [Xinjiang], Gushan Gobi (depository: ZIN) (examined). Rosa et al. 2017a: 80 (cat., type series), 221 (plate 223).

: Kimsey and Bohart 1991: 270 (cat., Mongolia [= China]: Sachow Gobi [= Oasis Sachzhou, Gashunskoe Gobi [= Dunhuang, Gansu]).

Semenov-Tian-Shanskij, 1967: 160. Holotype ♀; China: Alashan, Maladzhin (ZIN) (examined).

: Kimsey and Bohart 1991: 379 (cat., Mongolia [= China, Inner Mongolia]: Alashan, Maladzhin).

Mocsáry, 1912: 586. Holotype ♀; Kazakhstan: Altai: Semipalatinsk (HNHM) (examined).

: Kimsey and Bohart 1991: 381 (cat., Mongolia [= Kazakhstan]: Altai Mts).

du Buysson, 1909: 210. Holotype ♀; Viet Nam: Tonkin (MNHN).

: Kimsey and Bohart 1991: 412 (cat., Mongolia for the erroneous synonymy of Mocsáry, 1912).

Mocsáry, 1912: 406. Lectotype ♂ (designated by Bohart in Bohart and French 1986: 341); Kazakhstan: Semipalatinsk (HMNH) (examined).

: Kimsey and Bohart 1991: 412 (cat., Mongolia [= Kazakhstan]: Altai Mts).

Radoszkowski, 1891: 186. Syntypes ♀♀; Mongolia [= China]: Kansu, Jelissyn-Kuce (ISEA-PAS, MfN) (examined).

: Bischoff 1913: 54 (cat., Mongolia [= China]: Totau (locality not found)).

: Kimsey and Bohart 1991: 34 (cat., Mongolia [= China]: Kansu).

Radoszkowski, 1887: 47. Holotype ♂ [not ♀]; China: Xinjiang, Keria-Daria (ISEA-PAS) (examined).

: Mocsáry 1889: 516 (tax., descr., Mongolia [= China]); Bischoff 1913: 54 (cat., Mongolia [= China]).

: Dalla Torre 1892: 73 (cat., Mongolia [= China]); Kimsey and Bohart 1991: 427 (cat., Mongolia [= China]: Keria Daria).

Semenov-Tian-Shanskij, 1967: 160. Holotype ♂; China: Alashan, Uzosto canyon, 14.5.1908, leg. P. Kozlov (ZIN) (examined).

: Kimsey and Bohart 1991: 429 (cat., Mongolia [= China, Inner Mongolia]: Alashan, Uzosto Canyon).

Semenov-Tian-Shanskij, 1967: 178, nec Mocsáry, 1914. Holotype ♀; Russia: Transbaikalia: Ingoda River (ZIN) (examined).

Bohart in Kimsey and Bohart 1991: 440. Replacement name for Semenov-Tian-Shanskij, 1967, nec Mocsáry, 1914 (cat., Mongolia: Transbaikalia: Ingoda river).

Radoszkowski, 1891: 186. Holotype ♂; Mongolia [= China]: Tufyn (ISEA-PAS) (examined).

: Bischoff 1913: 57 (cat., Mongolia [= China]).

: Kimsey and Bohart 1991: 450 (cat., Mongolia [= China]: Tufyn).

Radoszkowski, 1887: 46. Holotype ♂; Mongolia [= China]: Zaïdam (ISEA-PAS) (examined).

: Mocsáry 1889: 504 (tax., descr., Mongolia [= China]); Bischoff 1913: 57 (cat., Mongolia [= China]).

: Dalla Torre 1892: 86 (cat., Mongolia [= China]); Kimsey and Bohart 1991: 452 (cat., Mongolia [= China]: Zaidam, Keria Mts).

Mocsáry in Radoszkowski, 1887: 48. Holotype ♂; Mongolia [= China]: Zaïdam (ISEA-PAS) (examined). Mocsáry 1889: 516 (cat., descr., Mongolia [= China]); Bischoff 1913: 59 (cat., Mongolia [= China]).

: Dalla Torre 1892: 97 (cat., Mongolia [= China]); Kimsey and Bohart 1991: 464 (cat., Mongolia: Zaidam).

Semenov-Tian-Shanskij, 1912: 190. Lectotype ♀ (designated by Rosa et al. 2017a: 45); Kyrgyzstan: Peter the Great Range, Gardan-Kaftar Pass (ZIN) (examined).

: Kimsey and Bohart 1991: 486 (cat., Mongolia [= Kyrgyzstan]).

Conclusions

Approximately 1500 chrysidid specimens were examined for the compilation of this first checklist of the Mongolian . Fifty-seven resulted newly recorded, but still a large number of specimens are laying unidentified in museum and private collections. Nineteen species were excluded from the fauna of Mongolia, because collecting localities are currently included in China territories; however, these species are expected for Mongolia. Based on the available data, distributional records for 90 Mongolian species are listed, representing 18 genera grouped in two subfamilies. In terms of species richness, are represented only by two species so far, and the subfamily is the most speciose (88 species, 98%). Among , is the most speciose tribe (47 species, 53.4%), followed by (39 species, 44.3%), and finally (2 species, 2.2%).

Currently eight species (9% of known taxa) are provisionally considered endemic: Rosa, Halada & Agnoli, sp. nov., Radoszkowski, 1889, Rosa, 2018, Rosa & Halada, sp. nov., (Móczár, 1967), du Buysson, 1891, Móczár, 1967, and (Semenov, 1932).

From a chorological point of view, one species has a Holarctic distribution (), ten have Palaearctic distributions, one has a Holarctic and Oriental distribution (), one a Palaearctic and Oriental distribution (), 28 species have an Asiatic-European distribution, 21 have an East Palaearctic distribution, and 19 have a Central Asian distribution.

Another result of the present study is a better knowledge of the distributional limits of some species, and Mongolia represents the easternmost record for seven species: , , , , , and .

The most widespread Mongolian species is the endemic recorded in ten aimags. and were recorded in nine aimags. This is not surprising because resulted in being one of the most common species from Western to Eastern Siberia also (Rosa et al. 2017c, d), whereas is one of the most common Euro-Siberian species ranging from Central Europe to China (Rosa et al. 2014). was recorded in eight aimags, whereas , and were recorded in seven7 aimags; they are Asiatic-European species, sometimes locally abundant. and were recorded in seven aimags, yet they are East-Palaearctic species.

Although most of the Mongolian aimags are under-represented in the existing data due to inadequacy of surveys, based on the currently available data we can state that the highest number of recorded species was collected in Tuv (42 species), Arkhangai (27 species), and Selenge and Dornod (20 species) aimags (Table 2). The family has not yet been documented from Bayan-Ulgii, Darkhan-Uul, Dundgovi, Khuvsgul, Orkhon, and Uvs although it is probable that this cosmopolitan family is present in these aimags and it is only a matter of time before the fauna is sampled and recorded.

Table 2.

Species diversity of aimags in terms of area size, number of specimens and number of collecting sites.

Aimags	Area, km2	No. of species	No. of coll. sites	No. of specimens
Arkhangai	55 314	27	4	237
Bayankhongor	115 978	18	10	142
Bulgan	48 733	18	4	61
Dornod	123 597	19	4	228
Dornogovi	109 472	16	4	42
Govi-Altai	141 448	8	5	49
Govi-Sümber	5 542	1	1	4
Khentii	80 325	12	1	55
Khovd	76 061	4	4	4
Selenge	41 153	20	1	100
Sukhbaatar	82 286	17	4	79
Tuv	74 042	42	10	350
Ulaanbaatar	4 704	10	4	20
Umnugovi	165 381	13	7	34
Uvurkhangai	62 895	12	5	42
Zavkhan	82 457	11	1	21

Comment. are not known in Bayan-Ulgii, Darkhan-Uul, Dundgovi, Khuvsgul, Orkhon, and Uvs.

Overall, the Mongolian fauna is still too poorly known for a complete analysis of species richness and composition. The faunal richness of Mongolia is doubtless much higher than we currently know, in comparison with the chrysidid fauna of the adjacent countries and considering the geographic position of Mongolian aimags and their different biotopes. For example, at least another 75 species recorded for Siberia (Rosa et al. 2017d) are expected for Mongolia, as well as other five genera known from bordering and central Asian countries (: Bischoff, 1913, Semenov, 1892; : Semenov, 1932, Semenov, 1954; : Bischoff, 1910). Copious undescribed members of the genus Linsenmaier, 1968 were collected in Mongolia. The descriptions of Mongolian , and a revision of this genus, are in preparation. Only a single record of an undescribed was previously known in literature for Central Asia (Turkmenistan) (Linsenmaier 1994).