Jinbiao Liu1,2, Changhai Chen3, Xiangdong Li1,2. 1. Jiangsu Co-innovation Center for Prevention and Control of Important Animal Infectious Diseases and Zoonoses, College of Veterinary Medicine, Yangzhou University, Yangzhou, China. 2. Joint International Research Laboratory of Agriculture and Agri-Product Safety, the Ministry of Education of China, Yangzhou University, Yangzhou, China. 3. Jiangsu Provincial Center for Animal Disease Control and Prevention, Nanjing, China.