Literature DB >> 32626487

List of non-EU Scolytinae of coniferous hosts.

Claude Bragard, Katharina Dehnen-Schmutz, Francesco Di Serio, Paolo Gonthier, Marie-Agnès Jacques, Josep Anton Jaques Miret, Annemarie Fejer Justesen, Alan MacLeod, Christer Sven Magnusson, Juan A Navas-Cortes, Stephen Parnell, Roel Potting, Philippe Lucien Reignault, Hans-Hermann Thulke, Wopke van der Werf, Antonio Vicent Civera, Jonathan Yuen, Lucia Zappalà, Jean-Claude Grégoire, Franz Streissl, Virág Kertész, Panagiotis Milonas.

Abstract

Following a request from the European Commission, the EFSA Panel on Plant Health prepared a list of non-EU Scolytinae spp. (Coleoptera: Curculionidae) affecting coniferous hosts. A literature review and search of databases, conducted up to January 2019, identified 804 Scolytinae species and subspecies of coniferous hosts. These Scolytinae were assigned to two categories (a) 705 non-EU species and subspecies, known to occur only outside the EU or having only limited presence in the EU, and (b) 99 species and subspecies with substantial presence in the EU (i.e. they are only reported so far from the EU or known to occur or be widespread in some Member States or reported in more than three EU MS). Scolytinae of category (b) will be excluded from further categorisation efforts. The main knowledge gaps and uncertainties of this listing concern (i) the status of species that are present in only a few MS and (ii) the status of the species that are present only at boundaries of the EU territory. The non-EU Scolytinae will be categorised by the Panel in a separate opinion.

Entities: Chemical Disease Gene Species

Keywords: European Union; bark beetles; conifers; non‐EU; pest risk; plant health; plant pest; quarantine

Year: 2020 PMID： 32626487 PMCID： PMC7008835 DOI： 10.2903/j.efsa.2020.5933

Source DB: PubMed Journal: EFSA J ISSN： 1831-4732

Introduction

Background and Terms of Reference as provided by the requestor

Background

Council Directive 2000/29/EC1 on protective measures against the introduction into the Community of organisms harmful to plants or plant products and against their spread within the Community establishes the present European Union plant health regime. The Directive lays down the phytosanitary provisions and the control checks to be carried out at the place of origin on plants and plant products destined for the Union or to be moved within the Union. In the Directive's 2000/29/EC annexes, the list of harmful organisms (pests) whose introduction into or spread within the Union is prohibited, is detailed together with specific requirements for import or internal movement. Following the evaluation of the plant health regime, the new basic plant health law, Regulation (EU) 2016/20312 on protective measures against pests of plants, was adopted on 26 October 2016 and will apply from 14 December 2019 onwards, repealing Directive 2000/29/EC. In line with the principles of the above mentioned legislation and the follow‐up work of the secondary legislation for the listing of EU regulated pests, EFSA is requested to provide pest categorizations of the harmful organisms included in the annexes of Directive 2000/29/EC, in the cases where recent pest risk assessment/pest categorisation is not available.

Terms of Reference

EFSA is requested, pursuant to Article 22(5.b) and Article 29(1) of Regulation (EC) No 178/2002,3 to provide scientific opinion in the field of plant health. EFSA is requested to prepare and deliver a pest categorisation (step 1 analysis) for each of the regulated pests included in the appendices of the annex to this mandate. The methodology and template of pest categorisation have already been developed in past mandates for the organisms listed in Annex II Part A Section II of Directive 2000/29/EC. The same methodology and outcome is expected for this work as well. The list of the harmful organisms included in the annex to this mandate comprises 133 harmful organisms or groups. A pest categorisation is expected for these 133 pests or groups and the delivery of the work would be stepwise at regular intervals through the year as detailed below. First priority covers the harmful organisms included in Appendix 1, comprising pests from Annex II Part A Section I and Annex II Part B of Directive 2000/29/EC. The delivery of all pest categorisations for the pests included in Appendix 1 is June 2018. The second priority is the pests included in Appendix 2, comprising the group of Cicadellidae (non‐EU) known to be vector of Pierce's disease (caused by Xylella fastidiosa), the group of Tephritidae (non‐EU), the group of potato viruses and virus‐like organisms, the group of viruses and virus‐like organisms of Cydonia Mill., Fragaria L., Malus Mill., Prunus L., Pyrus L., Ribes L., Rubus L. and Vitis L. and the group of Margarodes (non‐EU species). The delivery of all pest categorisations for the pests included in Appendix 2 is end 2019. The pests included in Appendix 3 cover pests of Annex I part A section I and all pests categorisations should be delivered by end 2020. For the above mentioned groups, each covering a large number of pests, the pest categorisation will be performed for the group and not the individual harmful organisms listed under “such as” notation in the Annexes of the Directive 2000/29/EC. The criteria to be taken particularly under consideration for these cases, is the analysis of host pest combination, investigation of pathways, the damages occurring and the relevant impact. Finally, as indicated in the text above, all references to ‘non‐European’ should be avoided and replaced by ‘non‐EU’ and refer to all territories with exception of the Union territories as defined in Article 1 point 3 of Regulation (EU) 2016/2031.

Terms of Reference: Appendix 1

List of harmful organisms for which pest categorisation is requested. The list below follows the annexes of Directive 2000/29/EC.

Terms of Reference: Appendix 2

List of harmful organisms for which pest categorisation is requested per group. The list below follows the categorisation included in the annexes of Directive 2000/29/EC.

Terms of Reference: Appendix 3

List of harmful organisms for which pest categorisation is requested. The list below follows the annexes of Directive 2000/29/EC.

Interpretation of the Terms of Reference

This scientific opinion presents the list of non‐EU Scolytinae spp. on coniferous host plants. They are listed as Scolytidae4 (non‐EU) in the Appendices to the Terms of Reference (ToR) to be subject to pest categorisation to determine whether they fulfil the criteria of quarantine pests or those of regulated non‐quarantine pests for the area of the EU excluding Ceuta, Melilla and the outermost regions of Member States (MS) referred to in Article 355(1) of the Treaty on the Functioning of the European Union (TFEU), other than Madeira and the Azores. As a first step toward this goal, the Panel prepared a list of Scolytinae species on coniferous host plants. The list is based on information collected from various literature sources and databases up to January 2019. In the process, two groups of Scolytinae were distinguished: non‐EU Scolytinae of coniferous hosts and Scolytinae with significant presence in the EU (known to occur in several MS, frequently reported in the EU, widespread in some MS) or so far reported only from the EU. A non‐EU Scolytinae is defined by its geographical distribution outside of the EU territory. As such, Scolytinae not reported from the EU and occurring only outside of the EU territory are considered as non‐EU Scolytinae. Furthermore, Scolytinae occurring outside the EU and having only a limited presence in the EU (reported from only a few MS, with restricted distribution) are also considered as non‐EU. The status of each species occupying 1, 2 or 3 MSs was then further examined. The panel identified three different groups: (i) species endemic only in the EU, (ii) species known for a long time to be established in MSs at the border of the EU, obviously at the margin of their distribution area outside of the EU, and (iii) species apparently distributed at random within the EU, suggesting recent and distinct introductions. The first two groups were excluded from further categorisation while the species in the last group were included as ‘non‐EU species’. This opinion provides the methodology and results for this classification which prepares but does not preclude the actual pest categorisation linked to the present mandate. This means that the Panel will then perform a pest categorisation for the non‐EU Scolytinae of coniferous hosts.

Data and methodologies

The preparatory work (literature review and data collection) for the current opinion was outsourced to the University of Padova (Department of Agronomy, Food, Natural Resources, Animals and Environment). The raw data can be downloaded under supporting documents (Annex A, Full list non‐EU Scolytinae and Annex B, Short list non‐EU Scolytinae). A comprehensive worldwide list of Scolytinae species living on conifers was generated by screening the databases of the European and Mediterranean Plant Protection Organisation (EPPO) and the Centre for Agriculture and Biosciences International (CABI), as well as available comprehensive and updated species lists and catalogues specific literature concerning all Scolytinae species in the world (Wood and Bright, 1992; Bright and Skidmore, 1997; Bright and Skidmore, 2002; Bright, 2014), or occurring in geographical areas hosting native conifers, such as North, Central and South America (Wood, 1982, 2007), Europe and North Africa (Pfeffer 1995, Knízek, 2011; Alonso‐Zarazaga et al., 2017), Australia (http://www.padil.gov.au), New Zealand (Brockerhoff et al., 2003) and Asia (Stark, 1952, Knízek, 2011; Alonso‐Zarazaga et al., 2017). Information on host(s) and distribution of Scolytinae in the first instance were retrieved from the EPPO Global Database (EPPO, 2019), the Centre for Agriculture and Biosciences International (CABI) and relevant publications. From the above‐mentioned sources, information was collected about species taxonomy (tribe); geographic distribution at continental level, considering the following macroregions: North America, Central America, South America, Europe, Asia, Oceania, North Africa, sub‐Saharan Africa). For all the species reported in the EU the geographic distribution was provided both at EU‐member states and at non‐EU European country levels in order to understand their distribution in Europe; feeding habits (herbiphagy, spermatophagy, mycophagy, myelophagy, phloeophagy, xylomycetophagy, xylophagy); list of host trees (at genus level) considering the most common conifer genera that are hosts of bark beetles: Abies spp., Cupressus spp., Larix spp., Picea spp., Pinus spp., Taxus spp., Juniperus spp., Cedrus spp., Chamaecyparis spp., Pseudotsuga spp., Tsuga spp., Thuja spp.; other possible hosts will be included in the category ‘other conifers’; potential for introduction: dispersal by trade reflected by the total number of interceptions of a given non‐EU Scolytinae species or genus recorded in the past years in the international ports and airports of North America, Europe, Australia and New Zealand. This information was obtained in the databases of USDA APHIS – Agriculture Quarantine Inspection, Australian Quarantine and Inspection Service, New Zealand Ministry of Primary Industries, Canadian Food Inspection Agency and Canadian Border Service Agency, EUROPHYT, and in specific literature and reviews such as, for instance, Brockerhoff et al. (2006) or Haack and Rabaglia (2013). Natural dispersal capacities by flight are rarely described and therefore were not investigated when building this list. Further references and information were obtained from experts and from citations within primary references. Data collected from National Plant Protection Organisations of the MS were also considered. The collected information was used to fill an extraction table (Annex A) that presents all raw data obtained. Only the non‐EU Scolytinae on conifers will be subject of further categorisation efforts in the frame of the present mandate. The Scolytinae excluded from this group are therefore referred to in the present opinion as Scolytinae excluded from further categorisation in the frame of the present mandate.

Listing of non‐EU Scolytinae

All Scolytinae species and subspecies identified in the literature and database searches have been listed and subsequently organised into the two groups specified below.

Scolytinae considered as non‐EU

The Scolytinae considered as non‐EU (Appendix A) belong to two subcategories: Scolytinae not known to be present in the EU, Scolytinae known to be present outside the EU and with only limited presence (i.e. reported in up to three MS, known to have restricted distribution) in the EU.

Scolytinae excluded from further categorisation in the frame of the present mandate

The Scolytinae species excluded from further categorisation in the frame of the present mandate are listed in Appendix B. They belong to two subcategories: Scolytinae species reported only from the EU, Scolytinae species known to be present outside the EU, but with a substantial presence also in the EU (known to occur in several MS, frequently reported in the EU, widespread in some MSs)

Uncertainties

When a species is present in only a few MS, its status (widely or locally established; transient) is often unclear from the literature. It is sometimes unclear whether species that are only present at the boundaries of the EU as well as in contiguous non‐EU countries are in the process of moving into the EU or have been established in these EU MS for a long time. Limited efforts on surveillance and therefore incomplete distribution data are an additional uncertainty that might be included. Non‐English (local) literature reports may have been overlooked.

Conclusions

The Panel was requested by the European Commission to produce a pest categorisation of 133 harmful organisms or groups listed in annexes of Directive 2000/29/EC. One of the groups for which the categorisation will be prepared is non‐EU Scolytinae spp. As a first step, a systematic approach identified 804 species and subspecies of Scolytinae species feeding on conifers. Among these Scolytinae species and subspecies, based on information on distribution and prevalence both inside and outside the EU, the Panel identified 705 non‐EU species and subspecies, known to occur only outside the EU or having only a limited presence in the EU (Appendix A). These Scolytinae will be further categorised in a separate opinion. The remaining 99 Scolytinae species and subspecies which have a substantial presence in the EU or are so far reported from the EU only (Appendix B), will not be categorised within the current mandate. However, the Commission may, at any time, request EFSA to categorise some or all the Scolytinae excluded from the present exercise. The main knowledge gaps and uncertainties of this listing concern (i) the status of species that are present in only a few MS, and (ii) and the status of the species present only at the boundaries of the EU.

Abbreviations

Centre for Agriculture and Bioscience International European and Mediterranean Plant Protection Organization Food and Agriculture Organization International Plant Protection Convention Member State National Center for Biotechnology Information EFSA Panel on Plant Health Treaty on the Functioning of the European Union Terms of Reference The appendix lists the Scolytinae species considered as non‐EU, their geographic occurrence in the EU (at MS level) and outside the EU (at continent level, or at country level for those species that are present also in the EU), the associated uncertainties or comments, as well as the main references from which the information was extracted. Presence in France and Spain is doubtful FR NPPO: the alleged presence in France is due to an error in labelling In the sense of phytosanitary terms, in line with Article 1 point 3 of Regulation (EU) 2016/2031, the Canary Islands are considered as Third Countries though they are part of Spain. Cognato A.I., 2000. Phylogenetic reveals new genus of Ipini bark beetle (Scolytidae). Annals of the Entomological Society of America 93:(362‐366.) and the species Ips (=Orthotomicus) orientalis Wood & Yin, 1986 was moved under Pseudips orientalis (Wood & Yin, 1986). The recent catalog of the palaearctic species of Alonso‐Zarazaga et al. (2017) adopts this new classification. The appendix lists the Scolytinae species present in the EU, their geographic occurrence in the EU MS and in non‐EU European countries, as well as outside the EU (at continent level), the associated uncertainties or comments, and the main references from which the information was extracted. FI NPPO: occurred in Finland before 1960. Endemic EU species SE NPPO: first observed in Sweden already around 1920. Listed as vulnerable on Swedish red‐list. Should be on EU list. FI NPPO: the species has not been reported in Finland, though it may occur SE NPPO: first observed in Sweden before 1960. Red‐listed in Sweden. FI NPPO: the species does not occur in Finland. The species was found in Poland before 1960 and Karpinski (1931) described it from Poland FI NPPO: occurred in Finland before 1960. Vulnerable in Finland according to the Finnish red‐list. SE NPPO: established in Sweden with old records e.g. from 1937. Red‐listed in Sweden. Should be on the EU list. FI NPPO: occurred in Finland before 1960. SE NPPO: established in Sweden with old records e.g. from 1944. Should be on the EU list. BE NPPO: the species is present in Belgium. SI NPPO: the species is present in Slovenia The annex contains detailed information on occurrence and the hosts of different non‐EU Scolytinae species. The annex contains detailed information of those non‐EU Scolytinae species for which information on their biology and impact was available. Full list non‐EU Scolytinae Click here for additional data file. Short list non EU Scolytinae Click here for additional data file.

Annex IIAI
(a) Insects, mites and nematodes, at all stages of their development
Aleurocantus spp.	Numonia pyrivorella (Matsumura)
Anthonomus bisignifer (Schenkling)	Oligonychus perditus Pritchard and Baker
Anthonomus signatus (Say)	Pissodes spp. (non‐EU)
Aschistonyx eppoi Inouye	Scirtothrips aurantii Faure
Carposina niponensis Walsingham	Scirtothrips citri (Moultex)
Enarmonia packardi (Zeller)	Scolytidae spp. (non‐EU)
Enarmonia prunivora Walsh	Scrobipalpopsis solanivora Povolny
Grapholita inopinata Heinrich	Tachypterellus quadrigibbus Say
Hishomonus phycitis	Toxoptera citricida Kirk.
Leucaspis japonica Ckll.	Unaspis citri Comstock
Listronotus bonariensis (Kuschel)
(b) Bacteria
Citrus variegated chlorosis	Xanthomonas campestris pv. oryzae (Ishiyama) Dye and pv. oryzicola (Fang. et al.) Dye
Erwinia stewartii (Smith) Dye
(c) Fungi
Alternaria alternata (Fr.) Keissler (non‐EU pathogenic isolates)	Elsinoe spp. Bitanc. and Jenk. Mendes
Anisogramma anomala (Peck) E. Müller	Fusarium oxysporum f. sp. albedinis (Kilian and Maire) Gordon
Apiosporina morbosa (Schwein.) v. Arx	Guignardia piricola (Nosa) Yamamoto
Ceratocystis virescens (Davidson) Moreau	Puccinia pittieriana Hennings
Cercoseptoria pini‐densiflorae (Hori and Nambu) Deighton	Stegophora ulmea (Schweinitz: Fries) Sydow & Sydow
Cercospora angolensis Carv. and Mendes	Venturia nashicola Tanaka and Yamamoto
(d) Virus and virus‐like organisms
Beet curly top virus (non‐EU isolates)	Little cherry pathogen (non‐ EU isolates)
Black raspberry latent virus	Naturally spreading psorosis
Blight and blight‐like	Palm lethal yellowing mycoplasm
Cadang‐Cadang viroid	Satsuma dwarf virus
Citrus tristeza virus (non‐EU isolates)	Tatter leaf virus
Leprosis	Witches’ broom (MLO)
Annex IIB
(a) Insect mites and nematodes, at all stages of their development
Anthonomus grandis (Boh.)	Ips cembrae Heer
Cephalcia lariciphila (Klug)	Ips duplicatus Sahlberg
Dendroctonus micans Kugelan	Ips sexdentatus Börner
Gilphinia hercyniae (Hartig)	Ips typographus Heer
Gonipterus scutellatus Gyll.	Sternochetus mangiferae Fabricius
Ips amitinus Eichhof
(b) Bacteria
Curtobacterium flaccumfaciens pv. flaccumfaciens (Hedges) Collins and Jones
(c) Fungi
Glomerella gossypii Edgerton	Hypoxylon mammatum (Wahl.) J. Miller
Gremmeniella abietina (Lag.) Morelet

Annex IAI
(a) Insects, mites and nematodes, at all stages of their development
Group of Cicadellidae (non‐EU) known to be vector of Pierce's disease (caused by Xylella fastidiosa), such as:
1) Carneocephala fulgida Nottingham	3) Graphocephala atropunctata (Signoret)
2) Draeculacephala minerva Ball
Group of Tephritidae (non‐EU) such as:
1) Anastrepha fraterculus (Wiedemann)	12) Pardalaspis cyanescens Bezzi
2) Anastrepha ludens (Loew)	13) Pardalaspis quinaria Bezzi
3) Anastrepha obliqua Macquart	14) Pterandrus rosa (Karsch)
4) Anastrepha suspensa (Loew)	15) Rhacochlaena japonica Ito
5) Dacus ciliatus Loew	16) Rhagoletis completa Cresson
6) Dacus curcurbitae Coquillet	17) Rhagoletis fausta (Osten‐Sacken)
7) Dacus dorsalis Hendel	18) Rhagoletis indifferens Curran
8) Dacus tryoni (Froggatt)	19) Rhagoletis mendax Curran
9) Dacus tsuneonis Miyake	20) Rhagoletis pomonella Walsh
10) Dacus zonatus Saund.	21) Rhagoletis suavis (Loew)
11) Epochra canadensis (Loew)
(c) Viruses and virus‐like organisms
Group of potato viruses and virus‐like organisms such as:
1) Andean potato latent virus	5) Potato virus T
2) Andean potato mottle virus	6) non‐EU isolates of potato viruses A, M, S, V, X and Y (including Yo, Yn and Yc) and Potato leafroll virus
3) Arracacha virus B, oca strain
4) Potato black ringspot virus
Group of viruses and virus‐like organisms of Cydonia Mill., Fragaria L., Malus Mill., Prunus L., Pyrus L., Ribes L., Rubus L. and Vitis L., such as:
1) Blueberry leaf mottle virus	8) Peach yellows mycoplasm
2) Cherry rasp leaf virus (American)	9) Plum line pattern virus (American)
3) Peach mosaic virus (American)	10) Raspberry leaf curl virus (American)
4) Peach phony rickettsia	11) Strawberry witches’ broom mycoplasma
5) Peach rosette mosaic virus	12) Non‐EU viruses and virus‐like organisms of Cydonia Mill., Fragaria L., Malus Mill., Prunus L., Pyrus L., Ribes L., Rubus L. and Vitis L.
6) Peach rosette mycoplasm
7) Peach X‐disease mycoplasm
Annex IIAI
(a) Insects, mites and nematodes, at all stages of their development
Group of Margarodes (non‐EU species) such as:
1) Margarodes vitis (Phillipi)	3) Margarodes prieskaensis Jakubski
2) Margarodes vredendalensis de Klerk

Annex IAI
(a) Insects, mites and nematodes, at all stages of their development
Acleris spp. (non‐EU)	Longidorus diadecturus Eveleigh and Allen
Amauromyza maculosa (Malloch)	Monochamus spp. (non‐EU)
Anomala orientalis Waterhouse	Myndus crudus Van Duzee
Arrhenodes minutus Drury	Nacobbus aberrans (Thorne) Thorne and Allen
Choristoneura spp. (non‐EU)	Naupactus leucoloma Boheman
Conotrachelus nenuphar (Herbst)	Premnotrypes spp. (non‐EU)
Dendrolimus sibiricus Tschetverikov	Pseudopityophthorus minutissimus (Zimmermann)
Diabrotica barberi Smith and Lawrence	Pseudopityophthorus pruinosus (Eichhoff)
Diabrotica undecimpunctata howardi Barber	Scaphoideus luteolus (Van Duzee)
Diabrotica undecimpunctata undecimpunctata Mannerheim	Spodoptera eridania (Cramer)
Diabrotica virgifera zeae Krysan & Smith	Spodoptera frugiperda (Smith)
Diaphorina citri Kuway	Spodoptera litura (Fabricus)
Heliothis zea (Boddie)	Thrips palmi Karny
Hirschmanniella spp., other than Hirschmanniella gracilis (de Man) Luc and Goodey	Xiphinema americanum Cobb sensu lato (non‐EU populations)
Liriomyza sativae Blanchard	Xiphinema californicum Lamberti and Bleve‐Zacheo
(b) Fungi
Ceratocystis fagacearum (Bretz) Hunt	Mycosphaerella larici‐leptolepis Ito et al.
Chrysomyxa arctostaphyli Dietel	Mycosphaerella populorum G. E. Thompson
Cronartium spp. (non‐EU)	Phoma andina Turkensteen
Endocronartium spp. (non‐EU)	Phyllosticta solitaria Ell. and Ev.
Guignardia laricina (Saw.) Yamamoto and Ito	Septoria lycopersici Speg. var. malagutii Ciccarone and Boerema
Gymnosporangium spp. (non‐EU)	Thecaphora solani Barrus
Inonotus weirii (Murril) Kotlaba and Pouzar	Trechispora brinkmannii (Bresad.) Rogers
Melampsora farlowii (Arthur) Davis
(c) Viruses and virus‐like organisms
Tobacco ringspot virus	Pepper mild tigré virus
Tomato ringspot virus	Squash leaf curl virus
Bean golden mosaic virus	Euphorbia mosaic virus
Cowpea mild mottle virus	Florida tomato virus
Lettuce infectious yellows virus
(d) Parasitic plants
Arceuthobium spp. (non‐EU)
Annex IAII
(a) Insects, mites and nematodes, at all stages of their development
Meloidogyne fallax Karssen	Rhizoecus hibisci Kawai and Takagi
Popillia japonica Newman
(b) Bacteria
Clavibacter michiganensis (Smith) Davis et al. ssp. sepedonicus (Spieckermann and Kotthoff) Davis et al.	Ralstonia solanacearum (Smith) Yabuuchi et al.
(c) Fungi
Melampsora medusae Thümen	Synchytrium endobioticum (Schilbersky) Percival
Annex I B
(a) Insects, mites and nematodes, at all stages of their development
Leptinotarsa decemlineata Say	Liriomyza bryoniae (Kaltenbach)
(b) Viruses and virus‐like organisms
Beet necrotic yellow vein virus

ID	Species	Presence in EU MS	Presence outside EU	Comments/Uncertainties	Reference
1.	Acanthotomicus curvidens		Sub‐Saharan Africa		Wood and Bright (1992)
2.	Ambrosiodmus eichhoffi		Sub‐Saharan Africa		Wood and Bright (1992)
3.	Ambrosiodmus funebris		South America		Atkinson (2018)
4.	Ambrosiodmus hagedorni		North America, Central America, South America		Atkinson (2018)
5.	Ambrosiodmus innominatus		Oceania		Wood and Bright (1992)
6.	Ambrosiodmus lecontei		North America, Central America		Atkinson (2018)
7.	Ambrosiodmus lewisi		North America, Asia		Atkinson (2018)
8.	Ambrosiodmus rusticus		North America, Central America, South America		Atkinson (2018)
9.	Araptus andinus		South America		Atkinson (2018)
10.	Araptus araucariae		South America		Atkinson (2018)
11.	Araptus impensus		South America		Atkinson (2018)
12.	Araptus sobrinus		Central America		Atkinson (2018)
13.	Cactopinus koebelei		North America		Atkinson (2018)
14.	Cactopinus pini		North America		Atkinson (2018)
15.	Carphobius arizonicus		North America		Atkinson (2018)
16.	Carphobius cupressi		Central America		Atkinson (2018)
17.	Carphobius pilifer		North America		Atkinson (2018)
18.	Carphoborus andersoni		North America		Atkinson (2018)
19.	Carphoborus bicornis		North America		Atkinson (2018)
20.	Carphoborus bifurcus		North America, Central America		Atkinson (2018)
21.	Carphoborus blaisdelli		North America		Atkinson (2018)
22.	Carphoborus bonnairei		Asia, North Africa		Alonso‐Zarazaga et al. (2017)
23.	Carphoborus brevisetosus		North America		Atkinson (2018)
24.	Carphoborus carri		North America		Atkinson (2018)
25.	Carphoborus convexifrons		North America		Atkinson (2018)
26.	Carphoborus costatus		Asia		Alonso‐Zarazaga et al. (2017)
27.	Carphoborus declivis		North America		Atkinson (2018)
28.	Carphoborus dunni		North America		Atkinson (2018)
29.	Carphoborus frontalis		North America		Atkinson (2018)
30.	Carphoborus henscheli	Cyprus	Georgia, Turkey, Israel, Syria		Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
31.	Carphoborus intermedius		North America		Atkinson (2018)
32.	Carphoborus jurinskii		Asia		Alonso‐Zarazaga et al. (2017)
33.	Carphoborus marani	Greece, Hungary	European Russia	Presence in Hungary only mentioned once and never confirmed later (Milos Knizek, pers. comm.)	Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
34.	Carphoborus mexicanus		North America		Atkinson (2018)
35.	Carphoborus perplexus		North America		Atkinson (2018)
36.	Carphoborus piceae		North America		Atkinson (2018)
37.	Carphoborus pinicolens		North America		Atkinson (2018)
38.	Carphoborus ponderosae		North America		Atkinson (2018)
39.	Carphoborus pseudotsugae		North America		Atkinson (2018)
40.	Carphoborus radiatae		North America		Atkinson (2018)
41.	Carphoborus sansoni		North America		Atkinson (2018)
42.	Carphoborus simplex		North America		Atkinson (2018)
43.	Carphoborus taireiensis		Asia		Alonso‐Zarazaga et al. (2017)
44.	Carphoborus vandykei		North America		Atkinson (2018)
45.	Carphoborus zhobi		Asia		Alonso‐Zarazaga et al. (2017)
46.	Chramesus spinosus		South America		Atkinson (2018)
47.	Cnestus mutilatus		North America, Asia, Oceania		Alonso‐Zarazaga et al. (2017) and Atkinson (2018)
48.	Coccotrypes advena		North America, Central America, Asia, Oceania		Wood and Bright (1992)
49.	Coccotrypes cinnamomi		Asia		Wood and Bright (1992)
50.	Coccotrypes fijianus		Oceania		Wood and Bright (1992)
51.	Coccotrypes leveri		Oceania		Wood and Bright (1992)
52.	Coccotrypes medius		Asia		Wood and Bright (1992)
53.	Coccotrypes norimasanus		Asia		Alonso‐Zarazaga et al. (2017)
54.	Coccotrypes nubilus		Asia, Sub‐Saharan Africa		Alonso‐Zarazaga et al. (2017)
55.	Coccotrypes recticollis		Asia		Wood and Bright (1992)
56.	Conophthorus apachecae		North America		Atkinson (2018)
57.	Conophthorus conicolens		North America		Atkinson (2018)
58.	Conophthorus coniperda		North America		Atkinson (2018)
59.	Conophthorus echinatae		North America		Atkinson (2018)
60.	Conophthorus edulis		North America		Atkinson (2018)
61.	Conophthorus mexicanus		North America, Central America		Atkinson (2018)
62.	Conophthorus michoacanae		North America		Atkinson (2018)
63.	Conophthorus monophyllae		North America		Atkinson (2018)
64.	Conophthorus ponderosae		North America		Atkinson (2018)
65.	Conophthorus radiatae		North America		Atkinson (2018)
66.	Conophthorus resinosae		North America		Atkinson (2018)
67.	Conophthorus teocotum		North America		Atkinson (2018)
68.	Conophthorus terminalis		North America		Atkinson (2018)
69.	Corthylus rufopilosus		South America		Atkinson (2018)
70.	Corthylus schaufussi		South America		Atkinson (2018)
71.	Corthylus simplex		Central America		Atkinson (2018)
72.	Cryphalus araucariae		Oceania		Wood and Bright (1992)
73.	Cryphalus brunneus		Oceania		Wood and Bright (1992)
74.	Cryphalus chamaecipariae		Asia		Alonso‐Zarazaga et al. (2017)
75.	Cryphalus chinlingensis		Asia		Alonso‐Zarazaga et al. (2017)
76.	Cryphalus cryptomeriae		Asia		Alonso‐Zarazaga et al. (2017)
77.	Cryphalus cylindrus		Oceania		Wood and Bright (1992)
78.	Cryphalus dissimilis		Asia		Alonso‐Zarazaga et al. (2017)
79.	Cryphalus diversicolor		Oceania		Wood and Bright (1992)
80.	Cryphalus fulvus		Asia		Alonso‐Zarazaga et al. (2017)
81.	Cryphalus jeholensis		Asia		Alonso‐Zarazaga et al. (2017)
82.	Cryphalus jezoensis		Asia		Alonso‐Zarazaga et al. (2017)
83.	Cryphalus kurenzovi		Asia		Alonso‐Zarazaga et al. (2017)
84.	Cryphalus laricis		Asia		Alonso‐Zarazaga et al. (2017)
85.	Cryphalus latus		Asia		Alonso‐Zarazaga et al. (2017)
86.	Cryphalus lepocrinus		Asia		Alonso‐Zarazaga et al. (2017)
87.	Cryphalus lipingensis		Asia		Alonso‐Zarazaga et al. (2017)
88.	Cryphalus longisetosus		Asia		Alonso‐Zarazaga et al. (2017)
89.	Cryphalus major		Asia		Alonso‐Zarazaga et al. (2017)
90.	Cryphalus malloti		Asia		Wood and Bright (1992)
91.	Cryphalus markangensis		Asia		Alonso‐Zarazaga et al. (2017)
92.	Cryphalus massonianus		Asia		Alonso‐Zarazaga et al. (2017)
93.	Cryphalus miyalopiceus		Asia		Alonso‐Zarazaga et al. (2017)
94.	Cryphalus montanus		Asia		Alonso‐Zarazaga et al. (2017)
95.	Cryphalus niponensis		Asia		Alonso‐Zarazaga et al. (2017)
96.	Cryphalus piceus		Asia		Alonso‐Zarazaga et al. (2017)
97.	Cryphalus pilosus		Asia		Alonso‐Zarazaga et al. (2017)
98.	Cryphalus pseudochinlingensis		Asia		Alonso‐Zarazaga et al. (2017)
99.	Cryphalus pseudotabulaeformis		Asia		Alonso‐Zarazaga et al. (2017)
100.	Cryphalus pubescens		North America		Atkinson (2018)
101.	Cryphalus redikorzevi		Asia		Alonso‐Zarazaga et al. (2017)
102.	Cryphalus rubentis		North America		Atkinson (2018)
103.	Cryphalus ruficollis		North America		Atkinson (2018)
104.	Cryphalus sawadai		Asia		Alonso‐Zarazaga et al. (2017)
105.	Cryphalus sichotensis		Asia		Alonso‐Zarazaga et al. (2017)
106.	Cryphalus sinoabietis		Asia		Alonso‐Zarazaga et al. (2017)
107.	Cryphalus strohmeyeri		Asia		Alonso‐Zarazaga et al. (2017)
108.	Cryphalus sylvicola		Oceania		Wood and Bright (1992)
109.	Cryphalus szechuanensis		Asia		Alonso‐Zarazaga et al. (2017)
110.	Cryphalus tabulaeformis		Asia		Alonso‐Zarazaga et al. (2017)
111.	Cryphalus taiwanus		Asia		Alonso‐Zarazaga et al. (2017)
112.	Cryphalus yamaguchii		Asia		Alonso‐Zarazaga et al. (2017)
113.	Cryptoxyleborus gracilior		Oceania		Wood and Bright (1992)
114.	Crypturgus alutaceus		North America, Central America		Atkinson (2018)
115.	Crypturgus beesoni		Asia		Alonso‐Zarazaga et al. (2017)
116.	Crypturgus borealis		North America		Atkinson (2018)
117.	Crypturgus cedri		North Africa		Alonso‐Zarazaga et al. (2017)
118.	Crypturgus concolor		North Africa		Alonso‐Zarazaga et al. (2017)
119.	Crypturgus dubius	France, Spain	Turkey, Iran	Presence in France and Spain is doubtful FR NPPO: the alleged presence in France is due to an error in labelling	Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
120.	Crypturgus tuberosus		Asia		Alonso‐Zarazaga et al. (2017)
121.	Cyrtogenius agathis		Asia		Wood and Bright (1992)
122.	Cyrtogenius brevior		Asia		Wood and Bright (1992)
123.	Cyrtogenius luteus	Italy	China, Japan, South Korea, South America	Introduced species	Atkinson (2018) and Gomez et al. (2012)
124.	Cyrtogenius perakensis		Asia		Alonso‐Zarazaga et al. (2017)
125.	Cyrtogenius philippinensis		Asia		Alonso‐Zarazaga et al. (2017)
126.	Dendroctonus adjunctus		North America, Central America		Atkinson (2018)
127.	Dendroctonus approximatus		North America, Central America		Atkinson (2018)
128.	Dendroctonus armandi		Asia		Alonso‐Zarazaga et al. (2017)
129.	Dendroctonus brevicomis		North America		Atkinson (2018)
130.	Dendroctonus frontalis		North America, Central America		Atkinson (2018)
131.	Dendroctonus jeffreyi		North America		Atkinson (2018)
132.	Dendroctonus mesoamericanus		North America, Central America		Atkinson (2018)
133.	Dendroctonus mexicanus		North America, Central America		Atkinson (2018)
134.	Dendroctonus murrayanae		North America		Atkinson (2018)
135.	Dendroctonus parallelocollis		North America, Central America		Atkinson (2018)
136.	Dendroctonus ponderosae		North America		Atkinson (2018)
137.	Dendroctonus pseudotsugae barragani		North America		Atkinson (2018)
138.	Dendroctonus pseudotsugae pseudotsugae		North America		Atkinson (2018)
139.	Dendroctonus punctatus		North America		Atkinson (2018)
140.	Dendroctonus rhizophagus		North America		Atkinson (2018)
141.	Dendroctonus rufipennis		North America		Atkinson (2018)
142.	Dendroctonus simplex		North America		Atkinson (2018)
143.	Dendroctonus terebrans		North America		Atkinson (2018)
144.	Dendroctonus valens		North America, Central America, Asia		Atkinson (2018)
145.	Dendroctonus vitei		North America, Central America		Atkinson (2018)
146.	Dolurgus pumilus		North America		Atkinson (2018)
147.	Dryocoetes affaber		North America		Atkinson (2018)
148.	Dryocoetes baikalicus		Asia, European Russia,		Alonso‐Zarazaga et al. (2017)
149.	Dryocoetes brevipilosus		Asia		Alonso‐Zarazaga et al. (2017)
150.	Dryocoetes caryi		North America		Atkinson (2018)
151.	Dryocoetes confusus		North America		Atkinson (2018)
152.	Dryocoetes cristatus		Asia		Alonso‐Zarazaga et al. (2017)
153.	Dryocoetes granicollis		North America		Atkinson (2018)
154.	Dryocoetes indicus		Asia		Alonso‐Zarazaga et al. (2017)
155.	Dryocoetes infuscatus		Asia		Alonso‐Zarazaga et al. (2017)
156.	Dryocoetes karamatsu		Asia		Alonso‐Zarazaga et al. (2017)
157.	Dryocoetes niijimai		Asia		Alonso‐Zarazaga et al. (2017)
158.	Dryocoetes pilosus		Asia		Alonso‐Zarazaga et al. (2017)
159.	Dryocoetes pini		Asia		Alonso‐Zarazaga et al. (2017)
160.	Dryocoetes quadrisulcatus		Asia		Alonso‐Zarazaga et al. (2017)
161.	Dryocoetes rugicollis		Asia		Alonso‐Zarazaga et al. (2017)
162.	Dryocoetes sechelti		North America		Atkinson (2018)
163.	Dryocoetes striatus		Asia		Alonso‐Zarazaga et al. (2017)
164.	Dryocoetes uniseriatus		Asia		Alonso‐Zarazaga et al. (2017)
165.	Euwallacea barbatus		Oceania		Wood and Bright (1992)
166.	Euwallacea interjectus		North America, Asia		Atkinson (2018)
167.	Euwallacea validus		North America, Asia		Atkinson (2018)
168.	Glostatus acaciae		Sub‐Saharan Africa		Wood and Bright (1992)
169.	Gnathotrichus deleoni		North America		Atkinson (2018)
170.	Gnathotrichus denticulatus		North America		Atkinson (2018)
171.	Gnathotrichus imitans		North America		Atkinson (2018)
172.	Gnathotrichus nitidifrons		North America, Central America		Atkinson (2018)
173.	Gnathotrichus omissus		Central America		Atkinson (2018)
174.	Gnathotrichus perniciosus		North America, Central America		Atkinson (2018)
175.	Gnathotrichus pilosus		North America		Atkinson (2018)
176.	Gnathotrichus retusus		North America		Atkinson (2018)
177.	Gnathotrichus sulcatus		North America, Central America		Atkinson (2018)
178.	Hylastes ambiguus		Asia		Alonso‐Zarazaga et al. (2017)
179.	Hylastes asperatus		North America		Atkinson (2018)
180.	Hylastes batnensis batnensis	Italy	Algeria, Morocco	Introduced species	Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
181.	Hylastes batnensis anatolicus		Turkey		Alonso‐Zarazaga et al. (2017)
182.	Hylastes flohri		North America		Atkinson (2018)
183.	Hylastes fulgidus		North America, Central America		Atkinson (2018)
184.	Hylastes gracilis		North America, Central America		Atkinson (2018)
185.	Hylastes longicollis		North America		Atkinson (2018)
186.	Hylastes lowei		North Africa		Alonso‐Zarazaga et al. (2017)
187.	Hylastes macer		North America		Atkinson (2018)
188.	Hylastes mexicanus		North America		Atkinson (2018)
189.	Hylastes niger		North America		Atkinson (2018)
190.	Hylastes nigrinus		North America		Atkinson (2018)
191.	Hylastes obscurus		Asia		Wood and Bright (1992)
192.	Hylastes parallelus		Asia		Alonso‐Zarazaga et al. (2017)
193.	Hylastes porculus		North America		Atkinson (2018)
194.	Hylastes retifer		North America		Atkinson (2018)
195.	Hylastes ruber		North America		Atkinson (2018)
196.	Hylastes salebrosus		North America		Atkinson (2018)
197.	Hylastes subopacus		North America		Atkinson (2018)
198.	Hylastes substriatus		Asia		Alonso‐Zarazaga et al. (2017)
199.	Hylastes techangensis		Asia		Alonso‐Zarazaga et al. (2017)
200.	Hylastes tenuis		North America, Central America		Atkinson (2018)
201.	Hyledius kesiyae		Asia		Wood and Bright (1992)
202.	Hyleops glabratus		Oceania		Wood and Bright (1992)
203.	Hylocurus beckeri		Central America		Wood and Bright (1992)
204.	Hylocurus rivalis		North America		Atkinson (2018)
205.	Hylurdrectonus araucariae		Oceania		Wood and Bright (1992)
206.	Hylurdrectonus corticinus		Oceania		Wood and Bright (1992)
207.	Hylurdrectonus nanus		Oceania		Wood and Bright (1992)
208.	Hylurdrectonus piniarius		Oceania		Wood and Bright (1992)
209.	Hylurgonotus antipodus		South America		Atkinson (2018)
210.	Hylurgonotus armaticeps		South America		Atkinson (2018)
211.	Hylurgonotus solidus		South America		Atkinson (2018)
212.	Hylurgonotus tuberculatus		South America		Atkinson (2018)
213.	Hylurgops bonvouloiri		North Africa		Alonso‐Zarazaga et al. (2017)
214.	Hylurgops eusulcatus		Asia		Alonso‐Zarazaga et al. (2017)
215.	Hylurgops incomptus		North America, Central America		Atkinson (2018)
216.	Hylurgops inouyei		Asia		Alonso‐Zarazaga et al. (2017)
217.	Hylurgops interstitialis		Asia		Alonso‐Zarazaga et al. (2017)
218.	Hylurgops junnanicus		Asia		Alonso‐Zarazaga et al. (2017)
219.	Hylurgops knausi		North America		Atkinson (2018)
220.	Hylurgops longipillus		Asia		Alonso‐Zarazaga et al. (2017)
221.	Hylurgops major		Asia		Alonso‐Zarazaga et al. (2017)
222.	Hylurgops pinifex		North America		Atkinson (2018)
223.	Hylurgops planirostris		North America, Central America		Atkinson (2018)
224.	Hylurgops porosus		North America		Atkinson (2018)
225.	Hylurgops reticulatus		North America		Atkinson (2018)
226.	Hylurgops rugipennis		North America		Atkinson (2018)
227.	Hylurgops spessiwzeffi		Asia		Alonso‐Zarazaga et al. (2017);
228.	Hylurgops sulcatus		Asia		Alonso‐Zarazaga et al. (2017)
229.	Hylurgops tuberculatus		Asia		Alonso‐Zarazaga et al. (2017)
230.	Hylurgus indicus		Asia		Alonso‐Zarazaga et al. (2017)
231.	Hypocryphalus caplandicus		Sub‐Saharan Africa		Wood and Bright (1992)
232.	Hypothenemus crudiae		North America, Central America, South America, Sub‐Saharan Africa		Wood (2007)
233.	Hypothenemus interstitialis		North America, Central America, South America		Atkinson (2018) and Wood and Bright (1992)
234.	Hypothenemus seriatus		North America, Central America, South America, Asia, North Africa, Sub‐Saharan Africa, Oceania		Atkinson (2018)
235.	Indocryphalus intermedius		North America, Asia		Alonso‐Zarazaga et al. (2017)
236.	Ips apache		North America, Central America		Atkinson (2018)
237.	Ips avulsus		North America, Central America		Atkinson (2018)
238.	Ips bonanseai		North America, Central America		Atkinson (2018)
239.	Ips borealis borealis		North America		Atkinson (2018)
240.	Ips borealis lanieri		North America		Atkinson (2018)
241.	Ips borealis swainei		North America		Atkinson (2018)
242.	Ips borealis thomasi		North America		Atkinson (2018)
243.	Ips calligraphus		North America, Central America		Atkinson (2018)
244.	Ips chinensis		Asia		Alonso‐Zarazaga et al. (2017)
245.	Ips confusus		North America		Atkinson (2018)
246.	Ips cribricollis		North America, Central America		Atkinson (2018)
247.	Ips emarginatus		North America		Atkinson (2018)
248.	Ips grandicollis		North America, Central America		Atkinson (2018)
249.	Ips hauseri		Asia		Alonso‐Zarazaga et al. (2017)
250.	Ips hoppingi		North America		Atkinson (2018)
251.	Ips hunteri		North America		Atkinson (2018)
252.	Ips integer		North America, Central America		Atkinson (2018)
253.	Ips knausi		North America		Atkinson (2018)
254.	Ips lecontei		North America, Central America		Atkinson (2018)
255.	Ips longifolia		Asia		Alonso‐Zarazaga et al. (2017)
256.	Ips montanus		North America		Atkinson (2018)
257.	Ips nitidus		Asia		Alonso‐Zarazaga et al. (2017)
258.	Ips paraconfusus		North America		Atkinson (2018)
259.	Ips perroti		North America		Atkinson (2018)
260.	Ips perturbatus		North America		Atkinson (2018)
261.	Ips pilifrons pilifrons		North America		Atkinson (2018)
262.	Ips pilifrons sulcifrons		North America		Atkinson (2018)
263.	Ips pilifrons thatcheri		North America		Atkinson (2018)
264.	Ips pilifrons utahensis		North America		Atkinson (2018)
265.	Ips pini		North America		Atkinson (2018)
266.	Ips plastographus maritimus		North America		Atkinson (2018)
267.	Ips plastographus plastographus		North America		Atkinson (2018)
268.	Ips schmutzenhoferi		Asia		Alonso‐Zarazaga et al. (2017)
269.	Ips shangrila		Asia		Alonso‐Zarazaga et al. (2017)
270.	Ips stebbingi		Asia		Alonso‐Zarazaga et al. (2017)
271.	Ips subelongatus		Asia, European Russia,	Introduced into European Russia from Asia	Alonso‐Zarazaga et al. (2017)
272.	Ips tridens engelmanni		North America		Atkinson (2018)
273.	Ips tridens tridens		North America		Atkinson (2018)
274.	Ips woodi		North America		Atkinson (2018)
275.	Lanurgus oleaeformis		Sub‐Saharan Africa		Wood and Bright (1992)
276.	Lanurgus podocarpi		Sub‐Saharan Africa		Wood and Bright (1992)
277.	Lanurgus spathulatus		Sub‐Saharan Africa		Wood and Bright (1992)
278.	Lanurgus widdringtoniae		Sub‐Saharan Africa		Wood and Bright (1992)
279.	Liparthrum longifolia		Asia		Alonso‐Zarazaga et al. (2017)
280.	Microperus eucalypticus		Oceania		Bright (2014)
281.	Microperus intermedius		Oceania		Bright (2014)
282.	Monarthrum nevermanni		Central America		Atkinson (2018)
283.	Orthotomicus angulatus		Asia, Oceania		Alonso‐Zarazaga et al. (2017)
284.	Orthotomicus caelatus		North America, Central America		Atkinson (2018)
285.	Orthotomicus chaokhao		Asia		Bright (2014)
286.	Orthotomicus golovjankoi		Asia		Alonso‐Zarazaga et al. (2017)
287.	Orthotomicus kuniyoshii		Asia		Alonso‐Zarazaga et al. (2017)
288.	Orthotomicus latidens		North America		Atkinson (2018)
289.	Orthotomicus multidentatus		Asia		Alonso‐Zarazaga et al. (2017)
290.	Orthotomicus nankinensis		Asia		Alonso‐Zarazaga et al. (2017)
291.	Orthotomicus nobilis		North Africa		Alonso‐Zarazaga et al. (2017)
292.	Orthotomicus pinivorus		Asia		Alonso‐Zarazaga et al. (2017)
293.	Orthotomicus spinifer		North America		Atkinson (2018)
294.	Orthotomicus tosaensis		Asia		Alonso‐Zarazaga et al. (2017)
295.	Orthotomicus tridentatus	Austria	Turkey		Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
296.	Pachycotes araucariae		Oceania		Wood and Bright (1992)
297.	Pachycotes australis		Oceania		Wood and Bright (1992)
298.	Pachycotes clavatus		Oceania		Wood and Bright (1992)
299.	Pachycotes engelsi		Oceania		Bright (2014)
300.	Pachycotes grandis		Oceania		Bright (2014)
301.	Pachycotes kuscheli		Oceania		Wood and Bright (1992)
302.	Pachycotes minor		Oceania		Wood and Bright (1992)
303.	Pachycotes peregrinus		Oceania		Wood and Bright (1992)
304.	Pachycotes villosus		Oceania		Wood and Bright (1992)
305.	Pachysquamus subcostulatus		North America		Mercado‐Velez and Negron (2014)
306.	Pagiocerus punctatus		South America		Atkinson (2018)
307.	Phloeosinus abietis		Asia		Alonso‐Zarazaga et al. (2017)
308.	Phloeosinus acatayi		Asia		Alonso‐Zarazaga et al. (2017)
309.	Phloeosinus antennatus		North America		Atkinson (2018)
310.	Phloeosinus arisanus		Asia		Alonso‐Zarazaga et al. (2017)
311.	Phloeosinus arizonicus		North America		Atkinson (2018)
312.	Phloeosinus armatus	Greece, Italy, Cyprus	Turkey, Libya, Iran, Israel, Jordan, Lebanon, Syria		Alonso‐Zarazaga et al. (2017), de Jong et al. (2014) and Pennachio et al. (2013)
313.	Phloeosinus baumanni		North America, Central America		Atkinson (2018)
314.	Phloeosinus canadensis		North America		Atkinson (2018)
315.	Phloeosinus cedri	Spain	Turkey, Algeria, Morocco, India	Introduced species	Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
316.	Phloeosinus cristatus		North America		Atkinson (2018)
317.	Phloeosinus cupressi		North America, Central America		Atkinson (2018)
318.	Phloeosinus deleoni		North America		Atkinson (2018)
319.	Phloeosinus dentatus		North America		Atkinson (2018)
320.	Phloeosinus frontalis		North America		Atkinson (2018)
321.	Phloeosinus fulgens		North America		Wood and Bright (1992)
322.	Phloeosinus furnissi		North America		Atkinson (2018)
323.	Phloeosinus gifuensis		Asia		Alonso‐Zarazaga et al. (2017)
324.	Phloeosinus gillerforsi	Azores (Portugal)	Canary Islands(a)		Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
325.	Phloeosinus hoferi		North America, Asia		Atkinson (2018)
326.	Phloeosinus hopehi		Asia		Wood and Bright (1992)
327.	Phloeosinus hoppingi		North America		Atkinson (2018)
328.	Phloeosinus jubatus		Asia		Alonso‐Zarazaga et al. (2017)
329.	Phloeosinus keeni		North America		Atkinson (2018)
330.	Phloeosinus kinabaluensis		Asia		Wood and Bright (1992)
331.	Phloeosinus lewisi		Asia		Alonso‐Zarazaga et al. (2017)
332.	Phloeosinus osumiensis		Asia		Alonso‐Zarazaga et al. (2017)
333.	Phloeosinus palearis		North America		Atkinson (2018)
334.	Phloeosinus perlatus		Asia		Alonso‐Zarazaga et al. (2017)
335.	Phloeosinus pertuberculatus		Asia		Alonso‐Zarazaga et al. (2017)
336.	Phloeosinus phyllocladus		Asia		Wood and Bright (1992)
337.	Phloeosinus pini		North America		Atkinson (2018)
338.	Phloeosinus podocarpi		Asia		Wood and Bright (1992)
339.	Phloeosinus punctatus		North America		Atkinson (2018)
340.	Phloeosinus sannohensis		Asia		Alonso‐Zarazaga et al. (2017)
341.	Phloeosinus scopulorum neomexicanus		North America		Atkinson (2018)
342.	Phloeosinus scopulorum scopulorum		North America		Atkinson (2018)
343.	Phloeosinus sequoiae		North America		Atkinson (2018)
344.	Phloeosinus seriatus		Asia		Wood and Bright (1992)
345.	Phloeosinus serratus		North America, Central America		Atkinson (2018)
346.	Phloeosinus setosus		North America		Atkinson (2018)
347.	Phloeosinus shensi		Asia		Alonso‐Zarazaga et al. (2017)
348.	Phloeosinus sinensis		Asia		Bright and Skidmore (2002)
349.	Phloeosinus spinosus		North America		Atkinson (2018)
350.	Phloeosinus swainei		North America		Atkinson (2018)
351.	Phloeosinus tacubayae		North America, Central America		Atkinson (2018)
352.	Phloeosinus taxodii		North America		Atkinson (2018)
353.	Phloeosinus turkestanicus		Asia		Alonso‐Zarazaga et al. (2017)
354.	Phloeosinus variolatus		North America		Atkinson (2018)
355.	Phloeotribus argentinensis		South America		Atkinson (2018)
356.	Phloeotribus atavus		Central America		Atkinson (2018)
357.	Phloeotribus cylindricus		South America		Atkinson (2018)
358.	Phloeotribus lecontei		North America		Atkinson (2018)
359.	Phloeotribus piceae		North America		Wood and Bright (1992)
360.	Pityoborus comatus		North America, Central America		Atkinson (2018)
361.	Pityoborus frontalis		North America		Atkinson (2018)
362.	Pityoborus hirtellus		North America		Atkinson (2018)
363.	Pityoborus hondurensis		Central America		Atkinson (2018)
364.	Pityoborus rubentis		North America		Atkinson (2018)
365.	Pityoborus secundus		North America		Atkinson (2018)
366.	Pityoborus velutinus		North America		Atkinson (2018)
367.	Pityogenes carinulatus		North America		Atkinson (2018)
368.	Pityogenes fossifrons		North America		Atkinson (2018)
369.	Pityogenes foveolatus		Asia		Alonso‐Zarazaga et al. (2017)
370.	Pityogenes hopkinsi		North America		Atkinson (2018)
371.	Pityogenes japonicus		Asia		Alonso‐Zarazaga et al. (2017)
372.	Pityogenes knechteli		North America		Atkinson (2018)
373.	Pityogenes meridianus		North America		Atkinson (2018)
374.	Pityogenes mexicanus		North America		Atkinson (2018)
375.	Pityogenes pennidens	Greece, Cyprus	Russia, Israel, Syria		Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
376.	Pityogenes plagiatus		North America		Atkinson (2018)
377.	Pityogenes rudnevi		Asia		Alonso‐Zarazaga et al. (2017)
378.	Pityogenes scitus		Asia		Alonso‐Zarazaga et al. (2017)
379.	Pityogenes seirindensis		Asia		Alonso‐Zarazaga et al. (2017)
380.	Pityogenes spessivtsevi		Asia		Alonso‐Zarazaga et al. (2017)
381.	Pityokteines elegans		North America		Atkinson (2018)
382.	Pityokteines lasiocarpi		North America		Atkinson (2018)
383.	Pityokteines marketae		Asia		Alonso‐Zarazaga et al. (2017)
384.	Pityokteines minutus		North America		Atkinson, 2018;
385.	Pityokteines mystacinus		North America		Atkinson (2018)
386.	Pityokteines ornatus		North America		Atkinson (2018)
387.	Pityokteines sparsus		North America		Atkinson (2018)
388.	Pityophthorus abiegnus		North America		Atkinson (2018)
389.	Pityophthorus abietinus		Asia		Alonso‐Zarazaga et al. (2017)
390.	Pityophthorus absonus		North America		Atkinson (2018)
391.	Pityophthorus abstrusus		North America		Atkinson (2018)
392.	Pityophthorus aciculatus		North America, Central America		Atkinson (2018)
393.	Pityophthorus acuminatus		North America, Central America		Atkinson (2018)
394.	Pityophthorus acutus		North America		Atkinson (2018)
395.	Pityophthorus alpinensis		North America		Atkinson (2018)
396.	Pityophthorus amoenus		Central America		Atkinson (2018)
397.	Pityophthorus amplus		North America		Atkinson (2018)
398.	Pityophthorus angustus		North America		Atkinson (2018)
399.	Pityophthorus annectens		North America, Central America		Atkinson (2018)
400.	Pityophthorus anthracinus		North America		Atkinson (2018)
401.	Pityophthorus anticus		South America		Atkinson (2018)
402.	Pityophthorus apachae		North America		Atkinson (2018)
403.	Pityophthorus aquilus		North America		Atkinson (2018)
404.	Pityophthorus arakii		Asia		Alonso‐Zarazaga et al. (2017)
405.	Pityophthorus arcanus		North America		Atkinson (2018)
406.	Pityophthorus aztecus		North America		Atkinson (2018)
407.	Pityophthorus balsameus		North America		Atkinson (2018)
408.	Pityophthorus barberi		North America		Atkinson (2018)
409.	Pityophthorus bassetti		North America		Atkinson (2018)
410.	Pityophthorus biovalis		North America		Atkinson (2018)
411.	Pityophthorus blackmani		Central America		Atkinson (2018)
412.	Pityophthorus blandulus		North America, Central America		Atkinson (2018)
413.	Pityophthorus blandus		North America		Atkinson (2018)
414.	Pityophthorus boycei		North America		Atkinson (2018)
415.	Pityophthorus bravoi		North America		Atkinson (2018)
416.	Pityophthorus brevicomatus		North America		Atkinson (2018)
417.	Pityophthorus brevis		North America		Atkinson (2018)
418.	Pityophthorus brighti		North America		Atkinson (2018)
419.	Pityophthorus briscoei		North America		Atkinson (2018)
420.	Pityophthorus cacuminatus		North America, Central America		Atkinson (2018)
421.	Pityophthorus californicus		North America		Atkinson (2018)
422.	Pityophthorus carinatus carinatus		North America		Atkinson (2018)
423.	Pityophthorus carinatus monticolae		North America		Atkinson (2018)
424.	Pityophthorus cariniceps		North America		Atkinson (2018)
425.	Pityophthorus carinulatus		North America		Atkinson (2018)
426.	Pityophthorus carmeli		North America		Atkinson (2018)
427.	Pityophthorus cascoensis		North America		Atkinson (2018)
428.	Pityophthorus cavatus		North America		Atkinson (2018)
429.	Pityophthorus cedri		Asia		Alonso‐Zarazaga et al. (2017)
430.	Pityophthorus chalcoensis		North America		Atkinson (2018)
431.	Pityophthorus chilgoza		Asia		Alonso‐Zarazaga et al. (2017)
432.	Pityophthorus ciliatus		North America		Atkinson (2018)
433.	Pityophthorus clarus		North America		Atkinson (2018)
434.	Pityophthorus clivus		North America		Atkinson (2018)
435.	Pityophthorus comosus		North America		Atkinson (2018)
436.	Pityophthorus concavus		North America		Atkinson (2018)
437.	Pityophthorus confertus		North America		Atkinson (2018)
438.	Pityophthorus confinis		North America		Atkinson (2018)
439.	Pityophthorus confusus		North America, Central America		Atkinson (2018)
440.	Pityophthorus consimilis		North America		Atkinson (2018)
441.	Pityophthorus cortezi		North America		Atkinson (2018)
442.	Pityophthorus crassus		North America		Atkinson (2018)
443.	Pityophthorus cristatus		North America		Atkinson (2018)
444.	Pityophthorus culminicolae		North America		Atkinson (2018)
445.	Pityophthorus cuspidatus		North America		Atkinson (2018)
446.	Pityophthorus declivisetosus		North America		Atkinson (2018)
447.	Pityophthorus deleoni		North America		Atkinson (2018)
448.	Pityophthorus deletus		North America		Atkinson (2018)
449.	Pityophthorus delicatus		North America, Central America		Atkinson (2018)
450.	Pityophthorus dentifrons		North America		Atkinson (2018)
451.	Pityophthorus deodara		Asia		Alonso‐Zarazaga et al. (2017)
452.	Pityophthorus digestus		North America		Atkinson (2018)
453.	Pityophthorus diglyphus		North America, Central America		Atkinson (2018)
454.	Pityophthorus discretus		North America		Atkinson (2018)
455.	Pityophthorus dispar		North America		Atkinson (2018)
456.	Pityophthorus dolus		North America		Atkinson (2018)
457.	Pityophthorus durus		North America		Atkinson (2018)
458.	Pityophthorus elatinus		North America		Atkinson (2018)
459.	Pityophthorus electus		North America		Atkinson (2018)
460.	Pityophthorus euterpes		North America		Atkinson (2018)
461.	Pityophthorus festus		North America, Central America		Atkinson (2018)
462.	Pityophthorus flavimaculatus		Asia		Alonso‐Zarazaga et al. (2017)
463.	Pityophthorus furnissi		North America		Atkinson (2018)
464.	Pityophthorus fuscus		North America		Atkinson (2018)
465.	Pityophthorus glabratulus		North America, Central America		Atkinson (2018)
466.	Pityophthorus grandis		North America		Atkinson (2018)
467.	Pityophthorus immanis		North America		Atkinson (2018)
468.	Pityophthorus impexus		North America		Atkinson (2018)
469.	Pityophthorus indigus		North America		Atkinson (2018)
470.	Pityophthorus ineditus		North America		Atkinson (2018)
471.	Pityophthorus infulatus		North America		Atkinson (2018)
472.	Pityophthorus ingens		North America		Atkinson (2018)
473.	Pityophthorus intentus		North America		Atkinson (2018)
474.	Pityophthorus intextus		North America		Atkinson (2018)
475.	Pityophthorus jeffreyi		North America		Atkinson (2018)
476.	Pityophthorus jucundus		Asia		Alonso‐Zarazaga et al. (2017)
477.	Pityophthorus keeni		North America		Atkinson (2018)
478.	Pityophthorus kirgisicus		Asia		Alonso‐Zarazaga et al. (2017)
479.	Pityophthorus lapponicus		Asia, Asian Russia	NPPO of Finland confirmed that the species does not occur in Finland	Alonso‐Zarazaga et al. (2017)
480.	Pityophthorus laticeps		North America		Atkinson (2018)
481.	Pityophthorus lautus		North America		Atkinson (2018)
482.	Pityophthorus lecontei		North America		Atkinson (2018)
483.	Pityophthorus leechi		North America		Atkinson (2018)
484.	Pityophthorus leiophyllae		North America		Atkinson (2018)
485.	Pityophthorus lepidus		North America		Atkinson (2018)
486.	Pityophthorus levis		North America		Atkinson (2018)
487.	Pityophthorus litos		North America		Atkinson (2018)
488.	Pityophthorus malleatus		North America		Atkinson (2018)
489.	Pityophthorus mauretanicus	France	Algeria, Egypt, Libya, Morocco, Tunisia	FR NPPO: species from North Africa, mistakenly cited from France in the Catalogue of Palaearctic Coleoptera.	Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
490.	Pityophthorus megas		North America		Atkinson (2018)
491.	Pityophthorus mesembria		Central America		Atkinson (2018)
492.	Pityophthorus micans		North America		Atkinson (2018)
493.	Pityophthorus micrographus sibiricus		Asia		Alonso‐Zarazaga et al. (2017)
494.	Pityophthorus miniatus		North America, Central America		Atkinson (2018)
495.	Pityophthorus modicus		North America		Atkinson (2018)
496.	Pityophthorus montezumae		North America		Atkinson (2018)
497.	Pityophthorus montivagus		North America		Atkinson (2018)
498.	Pityophthorus mormon		North America		Atkinson (2018)
499.	Pityophthorus murrayanae		North America		Atkinson (2018)
500.	Pityophthorus nigricans		North America, Central America		Atkinson (2018)
501.	Pityophthorus nitidulus		North America		Atkinson (2018)
502.	Pityophthorus nitidus		North America		Atkinson (2018)
503.	Pityophthorus nocturnus		North America, Central America		Atkinson (2018)
504.	Pityophthorus obtusipennis		North America, Central America		Atkinson (2018)
505.	Pityophthorus occidentalis		North America		Atkinson (2018)
506.	Pityophthorus occlusus		North America, Central America		Atkinson (2018)
507.	Pityophthorus opaculus		North America		Atkinson (2018)
508.	Pityophthorus orarius		North America		Atkinson (2018)
509.	Pityophthorus ornatus		North America		Atkinson (2018)
510.	Pityophthorus parfentievi		Asia		Alonso‐Zarazaga et al. (2017)
511.	Pityophthorus pellitus		North America, Central America		Atkinson (2018)
512.	Pityophthorus perotei		North America		Atkinson (2018)
513.	Pityophthorus pinavorus		North America, Central America		Atkinson (2018)
514.	Pityophthorus pinguis		North America		Atkinson (2018)
515.	Pityophthorus pini		Asia		Alonso‐Zarazaga et al. (2017)
516.	Pityophthorus pityographus cribratus	Greece	Russia, Turkey		Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
517.	Pityophthorus podocarpi		South America		Atkinson (2018)
518.	Pityophthorus pseudotsugae		North America		Atkinson (2018)
519.	Pityophthorus puberulus		North America		Atkinson (2018)
520.	Pityophthorus pubifrons		North America		Atkinson (2018)
521.	Pityophthorus pulchellus		North America		Atkinson (2018)
522.	Pityophthorus pulicarius		North America, Central America		Atkinson (2018)
523.	Pityophthorus pullus		North America		Atkinson (2018)
524.	Pityophthorus punctifrons		North America		Atkinson (2018)
525.	Pityophthorus ramiperda		North America		Atkinson (2018)
526.	Pityophthorus recens		North America		Atkinson (2018)
527.	Pityophthorus rubidus		North America		Atkinson (2018)
528.	Pityophthorus rudis		North America		Atkinson (2018)
529.	Pityophthorus sachalinensis		Asia		Alonso‐Zarazaga et al. (2017)
530.	Pityophthorus sapineus		North America		Atkinson (2018)
531.	Pityophthorus scabridus		North America, Central America		Atkinson (2018)
532.	Pityophthorus scalptor		North America		Atkinson (2018)
533.	Pityophthorus scalptus		North America		Atkinson (2018)
534.	Pityophthorus schwarzi		North America		Atkinson (2018)
535.	Pityophthorus schwerdtfegeri		North America, Central America		Atkinson (2018)
536.	Pityophthorus segnis		North America		Atkinson (2018)
537.	Pityophthorus seiryuensis		Asia		Alonso‐Zarazaga et al. (2017)
538.	Pityophthorus serratus		North America		Atkinson (2018)
539.	Pityophthorus setosus		North America		Atkinson (2018)
540.	Pityophthorus sichotensis		Asia		Alonso‐Zarazaga et al. (2017)
541.	Pityophthorus sierrensis		North America		Atkinson (2018)
542.	Pityophthorus solatus		North America		Atkinson (2018)
543.	Pityophthorus solers		North America		Atkinson (2018)
544.	Pityophthorus solus	Spain	North America	Introduced species	Alonso‐Zarazaga et al. (2017)
545.	Pityophthorus spadix		North America		Atkinson (2018)
546.	Pityophthorus speculum		North America		Atkinson (2018)
547.	Pityophthorus subopacus		North America		Atkinson (2018)
548.	Pityophthorus subsimilis		North America, Central America		Atkinson (2018)
549.	Pityophthorus thomasi		North America		Atkinson (2018)
550.	Pityophthorus toralis		North America		Atkinson (2018)
551.	Pityophthorus trepidus		North America		Atkinson (2018)
552.	Pityophthorus tuberculatus		North America		Atkinson (2018)
553.	Pityophthorus tumidus		North America		Atkinson (2018)
554.	Pityophthorus venustus		North America		Atkinson (2018)
555.	Pityophthorus vespertinus		North America		Atkinson (2018)
556.	Pityophthorus viminalis		North America		Atkinson (2018)
557.	Pityophthorus woodi		North America		Atkinson (2018)
558.	Pityophthorus zonalis		North America		Atkinson (2018)
559.	Pityotrichus barbatus		North America		Atkinson (2018)
560.	Pityotrichus hesperius		North America		Atkinson (2018)
561.	Pityotrichus turkmenicus		Asia		Alonso‐Zarazaga et al. (2017)
562.	Polygraphus abietis		Asia		Alonso‐Zarazaga et al. (2017)
563.	Polygraphus angustus		Asia		Alonso‐Zarazaga et al. (2017)
564.	Polygraphus aterrimus		Asia		Alonso‐Zarazaga et al. (2017)
565.	Polygraphus convexifrons		North America		Atkinson (2018)
566.	Polygraphus difficilis		Asia		Alonso‐Zarazaga et al. (2017)
567.	Polygraphus formosanus		Asia		Wood and Bright (1992)
568.	Polygraphus fulvipennis		Asia		Alonso‐Zarazaga et al. (2017)
569.	Polygraphus gracilis		Asia		Alonso‐Zarazaga et al. (2017)
570.	Polygraphus hoppingi		North America		Atkinson (2018)
571.	Polygraphus horyurensis		Asia		Alonso‐Zarazaga et al. (2017)
572.	Polygraphus japonicus		Asia		Alonso‐Zarazaga et al. (2017)
573.	Polygraphus jezoensis		Asia		Alonso‐Zarazaga et al. (2017)
574.	Polygraphus junnanicus		Asia		Alonso‐Zarazaga et al. (2017)
575.	Polygraphus kisoensis		Asia		Alonso‐Zarazaga et al. (2017)
576.	Polygraphus longifolia		Asia		Maiti and Saha (2004)
577.	Polygraphus major		Asia		Alonso‐Zarazaga et al. (2017)
578.	Polygraphus meakanensis		Asia		Alonso‐Zarazaga et al. (2017)
579.	Polygraphus nobuchii		Asia		Alonso‐Zarazaga et al. (2017)
580.	Polygraphus pini		Asia		Alonso‐Zarazaga et al. (2017)
581.	Polygraphus proximus		Russia, Asia	Introduced species	Alonso‐Zarazaga et al. (2017)
582.	Polygraphus rudis rudis		Asia		Alonso‐Zarazaga et al. (2017)
583.	Polygraphus rudis hexiensis		Asia		Alonso‐Zarazaga et al. (2017)
584.	Polygraphus rufipennis		North America		Atkinson (2018)
585.	Polygraphus setosus		Asia		Alonso‐Zarazaga et al. (2017)
586.	Polygraphus sinensis		Asia		Alonso‐Zarazaga et al. (2017)
587.	Polygraphus squamulatus		Asia		Alonso‐Zarazaga et al. (2017)
588.	Polygraphus sumatranus		Asia		Wood and Bright (1992)
589.	Polygraphus szemaoensis		Asia		Alonso‐Zarazaga et al. (2017)
590.	Polygraphus taiwanensis		Asia		Alonso‐Zarazaga et al. (2017)
591.	Polygraphus trenchi		Asia		Alonso‐Zarazaga et al. (2017)
592.	Polygraphus verrucifrons		Asia		Alonso‐Zarazaga et al. (2017)
593.	Pseudips concinnus		North America		Atkinson (2018)
594.	Pseudips orientalis (b)		Asia		Alonso‐Zarazaga et al. (2017)
595.	Pseudips mexicanus		North America, Central America		Atkinson (2018)
596.	Pseudohylesinus dispar dispar		North America		Atkinson (2018)
597.	Pseudohylesinus dispar pullatus		North America		Atkinson (2018)
598.	Pseudohylesinus granulatus		North America		Atkinson (2018)
599.	Pseudohylesinus maculosus		North America		Atkinson (2018)
600.	Pseudohylesinus magnus		North America		Atkinson (2018)
601.	Pseudohylesinus nebulosus nebulosus		North America		Atkinson (2018)
602.	Pseudohylesinus nebulosus serratus		North America		Atkinson (2018)
603.	Pseudohylesinus nobilis		North America		Atkinson (2018)
604.	Pseudohylesinus pini		North America		Atkinson (2018)
605.	Pseudohylesinus sericeus		North America, Asia		Atkinson (2018)
606.	Pseudohylesinus sitchensis		North America		Atkinson (2018)
607.	Pseudohylesinus tsugae		North America		Atkinson (2018)
608.	Pseudohylesinus variegatus		North America		Atkinson (2018)
609.	Pseudothysanoes coniferae		North America		Atkinson (2018)
610.	Pseudothysanoes pini		North America		Atkinson (2018)
611.	Pseudoxylechinus piceae		Asia		Alonso‐Zarazaga et al. (2017)
612.	Scierus annectans		North America		Atkinson (2018)
613.	Scierus pubescens		North America		Atkinson (2018)
614.	Scolytodes gennaeus		South America		Atkinson (2018)
615.	Scolytoplatypus daimio		Asia		Alonso‐Zarazaga et al. (2017)
616.	Scolytoplatypus kivuensis		Sub‐Saharan Africa		Wood and Bright (1992)
617.	Scolytoplatypus nitidus		Asia		Wood and Bright (1992)
618.	Scolytoplatypus pusillus		Asia		Wood and Bright (1992)
619.	Scolytoplatypus raja		Asia		Alonso‐Zarazaga et al. (2017)
620.	Scolytoplatypus shogun		Asia		Wood and Bright (1992)
621.	Scolytoplatypus siomio		Asia		Maiti and Saha (2004)
622.	Scolytoplatypus tycoon		Asia		Alonso‐Zarazaga et al. (2017)
623.	Scolytus aztecus		North America		Atkinson (2018)
624.	Scolytus dentatus		North America		Atkinson (2018)
625.	Scolytus fiskei		North America		Atkinson (2018)
626.	Scolytus hermosus		North America		Atkinson (2018)
627.	Scolytus laricis		North America		Atkinson (2018)
628.	Scolytus major		Asia		Alonso‐Zarazaga et al. (2017)
629.	Scolytus monticolae		North America		Atkinson (2018)
630.	Scolytus morawitzi		Russia, Asia		Alonso‐Zarazaga et al. (2017)
631.	Scolytus mundus		North America		Atkinson (2018)
632.	Scolytus numidicus		North Africa		Alonso‐Zarazaga et al. (2017)
633.	Scolytus obelus		North America		Atkinson (2018)
634.	Scolytus oregoni		North America		Atkinson (2018)
635.	Scolytus piceae		North America		Atkinson (2018)
636.	Scolytus praeceps		North America		Atkinson (2018)
637.	Scolytus reflexus		North America		Atkinson (2018)
638.	Scolytus robustus		North America		Atkinson (2018)
639.	Scolytus sinopiceus		Asia		Alonso‐Zarazaga et al. (2017)
640.	Scolytus subscaber		North America		Atkinson (2018)
641.	Scolytus tsugae		North America		Atkinson (2018)
642.	Scolytus unispinosus		North America		Atkinson (2018)
643.	Scolytus ventralis		North America		Atkinson (2018)
644.	Scolytus virgatus		North America		Atkinson (2018)
645.	Sphaerotrypes magnus		Asia		Alonso‐Zarazaga et al. (2017)
646.	Sphaerotrypes tsugae		Asia		Alonso‐Zarazaga et al. (2017)
647.	Sternobothrus costatus		South America		Atkinson (2018)
648.	Sternobothrus suturalis		South America		Atkinson (2018)
649.	Tomicus armandii		Asia		Alonso‐Zarazaga et al. (2017)
650.	Tomicus brevipilosus		Asia		Alonso‐Zarazaga et al. (2017)
651.	Tomicus pilifer		Asia		Alonso‐Zarazaga et al. (2017)
652.	Tomicus puellus		Asia		Alonso‐Zarazaga et al. (2017)
653.	Tomicus yunnanensis		Asia		Alonso‐Zarazaga et al. (2017)
654.	Traglostus brevisetosus		Sub‐Saharan Africa		Wood and Bright (1992)
655.	Trypodendron dorjitenzingi		Asia		Alonso‐Zarazaga et al. (2017)
656.	Trypodendron gaimaensis		Asia		Alonso‐Zarazaga et al. (2017)
657.	Trypodendron proximum		Asia		Alonso‐Zarazaga et al. (2017)
658.	Trypodendron rufitarsus		North America		Atkinson (2018)
659.	Trypodendron scabricollis		North America		Atkinson (2018)
660.	Xyleborinus gracilis	Azores (Portugal)	North America, Central America, South America	Introduced species	Atkinson (2018) and Alonso‐Zarazaga et al. (2017)
661.	Xyleborinus linearicollis		South America		Atkinson (2018)
662.	Xyleborinus sentosus		South America		Wood and Bright (1992)
663.	Xyleborinus sharpae		Sub‐Saharan Africa		Wood and Bright (1992)
664.	Xyleborinus spinifer		Sub‐Saharan Africa		Wood and Bright (1992)
665.	Xyleborus adelographus		South America		Atkinson (2018)
666.	Xyleborus agathis		Asia		Wood and Bright (1992)
667.	Xyleborus apertus		Asia, Oceania		Wood and Bright (1992)
668.	Xyleborus aquilus		Asia		Alonso‐Zarazaga et al. (2017)
669.	Xyleborus bidentatus		Asia, Sub‐Saharan Africa, Oceania		Wood and Bright (1992)
670.	Xyleborus catharinensis		South America		Atkinson (2018)
671.	Xyleborus compressus		Oceania		Wood and Bright (1992)
672.	Xyleborus detectus		Asia		Wood and Bright (1992)
673.	Xyleborus emarginatus		Asia		Alonso‐Zarazaga et al. (2017)
674.	Xyleborus ferrugineus		North America, Central America, South America, Asia, Sub‐Saharan Africa, Oceania		Atkinson (2018)
675.	Xyleborus festivus		Asia		Alonso‐Zarazaga et al. (2017)
676.	Xyleborus gratiosus		Oceania		Wood and Bright (1992)
677.	Xyleborus intrusus		North America, Central America		Atkinson (2018)
678.	Xyleborus muticus		Asia		Alonso‐Zarazaga et al. (2017)
679.	Xyleborus neivai		South America		Atkinson (2018)
680.	Xyleborus perforans	Azores (Portugal)	North America, Central America, South America, Asia, North Africa, Sub‐Saharan Africa, Oceania		Alonso‐Zarazaga et al. (2017)
681.	Xyleborus perplexus		Oceania		Wood and Bright (1992)
682.	Xyleborus pinicola		Asia		Wood and Bright (1992)
683.	Xyleborus pubescens		North America, Central America		Atkinson (2018)
684.	Xyleborus septentrionalis		Asia		Alonso‐Zarazaga et al. (2017)
685.	Xyleborus seriatus		North America, Asia		Alonso‐Zarazaga et al. (2017)
686.	Xyleborus spinulosus		North America, Central America, South America		Atkinson (2018)
687.	Xyleborus squamulatus		North America, Central America, South America		Atkinson (2018)
688.	Xyleborus volvulus		North America, Central America, South America, Asia, Sub‐Saharan Africa, Oceania		Atkinson (2018)
689.	Xylechinosomus bicolor		South America		Atkinson (2018)
690.	Xylechinosomus brasiliensis		South America		Atkinson (2018)
691.	Xylechinosomus contractus		South America		Atkinson (2018)
692.	Xylechinosomus hirsutus		South America		Atkinson (2018)
693.	Xylechinosomus humilus		South America		Atkinson (2018)
694.	Xylechinosomus lucianae		South America		Atkinson (2018)
695.	Xylechinosomus minimus		South America		Atkinson (2018)
696.	Xylechinosomus paranaensis		South America		Atkinson (2018)
697.	Xylechinosomus pilosus		South America		Atkinson (2018)
698.	Xylechinosomus sachtlebeni		South America		Atkinson (2018)
699.	Xylechinosomus valdivianus		South America		Atkinson (2018)
700.	Xylechinus americanus		South America		Atkinson (2018)
701.	Xylechinus araucariae		Oceania		Bright (2014)
702.	Xylechinus montanus		North America		Atkinson (2018)
703.	Xylosandrus compactus	Italy, France, Greece	North America, Central America, South America, Asia, Sub‐Saharan Africa		Wood and Bright (1992), Wood (2007), Garonna et al. (2012), Anses (2017) and Spanou et al. (2019)
704.	Xylosandrus pseudosolidus		Oceania		Wood and Bright (1992)
705.	Xyloterinus politus		North America		Wood and Bright (1992)

In the sense of phytosanitary terms, in line with Article 1 point 3 of Regulation (EU) 2016/2031, the Canary Islands are considered as Third Countries though they are part of Spain.

Cognato A.I., 2000. Phylogenetic reveals new genus of Ipini bark beetle (Scolytidae). Annals of the Entomological Society of America 93:(362‐366.) and the species Ips (=Orthotomicus) orientalis Wood & Yin, 1986 was moved under Pseudips orientalis (Wood & Yin, 1986). The recent catalog of the palaearctic species of Alonso‐Zarazaga et al. (2017) adopts this new classification.

IID	Species	List of EU MS where species is present	List of non‐EU European countries where pest is present	Presence outside EU (continents)	Comments/Uncertainties	Reference
1.	Carphoborus cholodkovskyi	Estonia, Finland, Poland, Sweden	Norway, Russia	Asia		Alonso‐Zarazaga et al. (2017)
2.	Carphoborus minimus	Austria, Bulgaria, Croatia, Czech Republic, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Netherlands, Poland, Romania, Slovakia, Slovenia, Spain	Albania, Belarus, Bosnia‐Herzegovina, North Macedonia, Montenegro, Russia, Switzerland, Ukraine	Asia, North Africa		Alonso‐Zarazaga et al. (2017)
3.	Carphoborus pini	Croatia, Cyprus, France, Greece, Italy, Spain	Bosnia‐Herzegovina	Asia, North Africa		Alonso‐Zarazaga et al. (2017)
4.	Carphoborus rossicus	Austria, Finland, Sweden	Russia		FI NPPO: occurred in Finland before 1960. Endemic EU species SE NPPO: first observed in Sweden already around 1920. Listed as vulnerable on Swedish red‐list. Should be on EU list.	Alonso‐Zarazaga et al. (2017), de Jong et al. (2014), Hyvärinen et al. (2019), ArtDatabanken (2015), ArtDatabanken SLU (2019a), and Voolma et al. (2004)
5.	Carphoborus teplouchovi	Sweden	Russia	China, Mongolia	FI NPPO: the species has not been reported in Finland, though it may occur SE NPPO: first observed in Sweden before 1960. Red‐listed in Sweden.	Alonso‐Zarazaga et al. (2017), de Jong et al. (2014), ArtDatabanken (2015), ArtDatabanken SLU (2019b) and Voolma et al. (2004)
6.	Cryphalus asperatus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Netherlands, Poland, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Montenegro, Norway, Russia, Switzerland	Asia, North Africa		Alonso‐Zarazaga et al. (2017) and Wood and Bright (1992)
7.	Cryphalus intermedius	Austria, Czech Republic, France, Germany, Hungary, Italy, Poland, Romania, Slovakia, Slovenia	Switzerland, Ukraine			Alonso‐Zarazaga et al. (2017)
8.	Cryphalus numidicus	Bulgaria, France, Greece, Italy, Spain	Switzerland	Asia, North Africa		Alonso‐Zarazaga et al. (2017)
9.	Cryphalus piceae	Austria, Bulgaria, Croatia, Czech Republic, Denmark, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Poland, Romania, Slovakia, Slovenia, Spain	Belarus, Bosnia‐Herzegovina, North Macedonia, Montenegro, Russia, Switzerland, Ukraine	Asia, North Africa		Alonso‐Zarazaga et al. (2017) and Hansen (1996)
10.	Cryphalus saltuarius	Austria, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Hungary, Italy, Poland, Slovakia, Sweden	Belarus, Montenegro, Norway, Russia, Switzerland	Asia		Alonso‐Zarazaga et al. (2017)
11.	Crypturgus cinereus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Luxembourg, Poland, Romania, Slovakia, Slovenia, Spain, Sweden	Belarus, North Macedonia, Montenegro, Norway, Russia, Switzerland, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
12.	Crypturgus cribrellus	Bulgaria, Croatia, France, Italy, Portugal, Slovenia, Spain	North Macedonia, Montenegro, Ukraine			Alonso‐Zarazaga et al. (2017)
13.	Crypturgus cylindricollis	Bulgaria, Croatia, Greece, Malta	Bosnia‐Herzegovina	Asia		Alonso‐Zarazaga et al. (2017)
14.	Crypturgus hispidulus	Austria, Bulgaria, Czech Republic, Denmark, Estonia, Finland, Germany, Hungary, Italy, Latvia, Lithuania, Netherlands, Poland, Slovakia, Sweden	Belarus, Norway, Russia, Switzerland	Asia		Alonso‐Zarazaga et al. (2017), Vorst et al. (2008) and Vorst (2010)
15.	Crypturgus mediterraneus	Croatia, Cyprus, France, Greece, Italy, Portugal	Russia, Ukraine	Asia, North Africa		Alonso‐Zarazaga et al. (2017)
16.	Crypturgus numidicus	Bulgaria, Croatia, France, Greece, Italy, Malta, Spain	Bosnia‐Herzegovina, Montenegro, Russia	Asia, North Africa		Alonso‐Zarazaga et al. (2017)
17.	Crypturgus parallelocollis	Greece	Turkey			Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
18.	Crypturgus pusillus	Austria, Belgium, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Netherlands, Poland, Portugal, Romania, Slovakia, Slovenia, Spain, Sweden	Belarus, Bosnia‐Herzegovina, North Macedonia, Montenegro, Norway, Russia, Switzerland, Ukraine	North America, Asia, North Africa		Alonso‐Zarazaga et al. (2017)
19.	Crypturgus subcribrosus	Austria, Czech Republic, Denmark, Estonia, Finland, Germany, Latvia, Lithuania, Netherlands, Poland, Slovakia, Slovenia, Sweden	Belarus, Bosnia‐Herzegovina, Norway, Russia, Ukraine	Asia		Alonso‐Zarazaga et al. (2017), Vorst (2010) and Heijerman and Noordijk (2016)
20.	Dendroctonus micans	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Poland, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Montenegro, Norway, Russia, Switzerland, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
21.	Dryocoetes autographus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Netherlands, Poland, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, Moldova, Montenegro, Norway, Russia, Switzerland, Ukraine	North America, South America, Asia, North Africa		Atkinson (2018)
22.	Dryocoetes hectographus	Belgium, Bulgaria, Czech Republic, Finland, France, Germany, Hungary, Italy, Latvia, Lithuania, Poland, Slovakia, Slovenia, Sweden	Belarus, Montenegro, Norway, Russia, Switzerland	Asia		Alonso‐Zarazaga et al. (2017)
23.	Gnathotrichus materiarius	Belgium, Czech Republic, Finland, France, Germany, Italy, Netherlands, Poland, Slovenia, Spain, Sweden	Switzerland	North America, Central America	Introduced into Europe	Atkinson (2018), Geister (2004) and Kirkendall and Faccoli (2010)
24.	Hylastes angustatus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Netherlands, Poland, Portugal, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, North Macedonia, Montenegro, Russia, Switzerland, Ukraine	Asia, Sub‐Saharan Africa		Alonso‐Zarazaga et al. (2017)
25.	Hylastes ater	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Netherlands, Poland, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Montenegro, Russia, Switzerland, Ukraine	South America, Asia, North Africa, Oceania	FI NPPO: the species does not occur in Finland	Atkinson (2018)
26.	Hylastes attenuatus	Austria, Belgium, Croatia, Czech Republic, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Netherlands, Poland, Portugal, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Norway, Russia, Switzerland, Ukraine	Asia, North Africa		Alonso‐Zarazaga et al. (2017), Heijerman and Noordijk, 2017, 2018, 2019 and Vorst (2010)
27.	Hylastes brunneus	Austria, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, Germany, Greece, Hungary, Ireland, Italy, Latvia, Netherlands, Poland, Slovakia, Slovenia, Sweden, UK	North Macedonia, Norway, Russia, Switzerland	Asia		Alonso‐Zarazaga et al. (2017) and Vorst (2010)
28.	Hylastes cunicularius	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Netherlands, Poland, Portugal, Slovakia, Slovenia, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Montenegro, Norway, Russia, Switzerland, Ukraine	Asia, North Africa		Alonso‐Zarazaga et al. (2017) and Vorst (2010)
29.	Hylastes fallax	Austria, Czech Republic, Italy, Romania, Slovakia				Alonso‐Zarazaga et al. (2017)
30.	Hylastes linearis	Croatia, Cyprus, Czech Republic, France, Germany, Greece, Hungary, Italy, Netherlands, Poland, Portugal, Slovenia, Spain, UK	North Macedonia, Montenegro, Switzerland	South America, Asia, North Africa, Sub‐Saharan Africa		Atkinson (2018) and Vorst (2010)
31.	Hylastes opacus	Austria, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Netherlands, Poland, Slovakia, Slovenia, Sweden, UK	Belarus, North Macedonia, Montenegro, Norway, Russia, Switzerland	North America, Asia		Atkinson (2018) and Vorst (2010)
32.	Hylurgops glabratus	Austria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Poland, Slovakia, Slovenia, Sweden	Belarus, Bosnia‐Herzegovina, Norway, Russia, Switzerland	Asia		Alonso‐Zarazaga et al. (2017)
33.	Hylurgops palliatus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Hungary, Ireland, Italy, Latvia, Lithuania, Netherlands, Poland, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, North Macedonia, Montenegro, Norway, Russia, Switzerland, Ukraine	North America, Asia, North Africa		Atkinson (2018)
34.	Hylurgus ligniperda	Austria, Belgium, Bulgaria, Croatia, Cyprus, Czech Republic, Denmark, Estonia, Finland, France, Germany, Hungary, Italy, Latvia, Lithuania, Netherlands, Poland, Portugal, Romania, Slovakia, Slovenia, Spain, Sweden	Bosnia‐Herzegovina, North Macedonia, Moldova, Montenegro, Russia, Serbia, Switzerland, Ukraine	North America, South America, Asia, North Africa, Sub‐Saharan Africa, Oceania		Atkinson (2018)
35.	Hylurgus micklitzi	Croatia, France, Germany, Greece, Hungary, Italy, Malta, Spain	Russia	Asia, North Africa		Alonso‐Zarazaga et al. (2017)
36.	Hypothenemus eruditus	Croatia, France, Italy, Malta, Spain		North America, Central America, South America, Asia, North Africa, Sub‐Saharan Africa, Oceania		Atkinson (2018)
37.	Ips acuminatus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Luxembourg, Poland, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Moldova, Montenegro, Norway, Russia, Switzerland, Ukraine	Asia	NL NPPO: the reported presence of I. acuminatus in NL is questionable. All historical findings are associated with wood imports and it is, thus, questionable whether a viable population has established (Heijerman, 2010).	Alonso‐Zarazaga et al. (2017)
38.	Ips amitinus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Poland, Romania, Slovakia, Slovenia, Sweden	Bosnia‐Herzegovina, North Macedonia, Montenegro, Russia, Switzerland, Ukraine	Asia	NL NPPO: No valid records of the species exist in the Netherlands	Alonso‐Zarazaga et al. (2017) and Jurc and Bojović (2004)
39.	Ips cembrae	Austria, Belgium, Czech Republic, Denmark, France, Germany, Greece, Hungary, Italy, Netherlands, Poland, Slovakia, Slovenia, Sweden, UK	Russia, Switzerland	Asia		Alonso‐Zarazaga et al. (2017) and ArtDatabanken SLU (2019e)
40.	Ips duplicatus	Austria, Czech Republic, Estonia, Finland, France, Germany, Hungary, Latvia, Lithuania, Poland, Slovakia, Sweden	Belarus, Norway, Russia	Asia		Alonso‐Zarazaga et al. (2017)
41.	Ips sexdentatus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Poland, Portugal, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Moldova, Montenegro, Norway, Russia, Switzerland, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
42.	Ips typographus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Poland, Portugal, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, Moldova, Montenegro, Norway, Russia, Switzerland, Ukraine	Asia, North Africa		Alonso‐Zarazaga et al. (2017), Vorst et al. (2008), Vorst (2010) and Heijerman and Noordijk (2017, 2019)
43.	Orthotomicus erosus	Austria, Bulgaria, Croatia, Cyprus, France, Greece, Hungary, Italy, Malta, Netherlands, Portugal, Slovenia, Spain, UK	North Macedonia, Montenegro, Norway, Russia, Switzerland, Ukraine	North America, South America, Asia, North Africa, Sub‐Saharan Africa	CY NPPO: The species is present in Cyprus	Atkinson (2018), Vorst et al. (2008) and Vorst (2010)
44.	Orthotomicus laricis	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Poland, Portugal, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Moldova, Montenegro, Norway, Russia, Switzerland, Ukraine	South America, Asia, North Africa		Atkinson (2018)
45.	Orthotomicus longicollis	Bulgaria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Poland, Slovakia, Slovenia, Spain, Sweden	Belarus, Bosnia‐Herzegovina, North Macedonia, Norway, Russia, Serbia, Switzerland, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
46.	Orthotomicus mannsfeldi	Austria, Bulgaria, Croatia, France, Greece, Hungary, Italy, Poland, Romania, Spain	Bosnia‐Herzegovina, North Macedonia, Montenegro	Asia		Alonso‐Zarazaga et al. (2017)
47.	Orthotomicus proximus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Netherlands, Poland, Portugal, Slovakia, Slovenia, Spain, Sweden	Belarus, Bosnia‐Herzegovina, Moldova, Montenegro, Norway, Russia, Serbia, Switzerland, Ukraine	Asia, North Africa, Sub‐Saharan Africa		Alonso‐Zarazaga et al. (2017) and Artsdatabanken (2019)
48.	Orthotomicus robustus	Austria, Bulgaria, Czech Republic, Greece, Hungary, Slovakia	Bosnia‐Herzegovina	Asia, North Africa		Alonso‐Zarazaga et al. (2017)
49.	Orthotomicus starki	Lithuania, Poland	Belarus, Russia		FI NPPO: the species does not occur in Finland. The species was found in Poland before 1960 and Karpinski (1931) described it from Poland	Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
50.	Orthotomicus suturalis	Austria, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Netherlands, Poland, Portugal, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Moldova, Montenegro, Norway, Russia, Serbia, Switzerland, Ukraine			Alonso‐Zarazaga et al. (2017)
51.	Phloeosinus aubei	Austria, Belgium, Bulgaria, Croatia, Cyprus, Czech Republic, France, Greece, Hungary, Italy, Netherlands, Poland, Portugal, Romania, Slovakia, Slovenia, Spain, UK	Albania, Bosnia‐Herzegovina, North Macedonia, Montenegro, Russia, Switzerland, Ukraine	Asia, North Africa	SI NPPO: The presence of the species was confirmed based on the entomological collection of the Biotechnical Faculty, Department of Forestry and Renewable Forest Resources	Alonso‐Zarazaga et al. (2017), Moraal (2005) and Vorst (2010)
52.	Phloeosinus henschi	Bulgaria, Croatia, Greece, Slovenia, Italy	Bosnia and Herzegovina, North Macedonia, Ukraine	Asia	The species is associated with pathogenic fungi (Geosmithia spp.)	Alonso‐Zarazaga et al. (2017), de Jong et al. (2014) and Kolařík et al. (2007)
53.	Phloeosinus laricionis	Italy				Alonso‐Zarazaga et al. (2017)
54.	Phloeosinus pfefferi	Cyprus				Alonso‐Zarazaga et al. (2017)
55.	Phloeosinus rudis	France, Netherlands, Italy, Belgium		Asia		Alonso‐Zarazaga et al. (2017), de Jong et al. (2014) and Moucheron et al. (2019)
56.	Phloeosinus thujae	Austria, Belgium, Bulgaria, Croatia, France, Germany, Greece, Hungary, Italy, Malta, Netherlands, Poland, Portugal, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Albania, Bosnia‐Herzegovina, North Macedonia, Montenegro, Russia, Switzerland, Ukraine	Asia, North Africa		Alonso‐Zarazaga et al. (2017), Lindelöw (2013) and ArtDatabanken SLU (2019f)
57.	Phloeotribus spinulosus	Austria, Belgium, Czech Republic, Denmark, Estonia, Finland, France, Germany, Italy, Latvia, Netherlands, Poland, Romania, Slovakia, Spain, Sweden	Belarus, Norway, Russia, Switzerland, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
58.	Pityogenes bidentatus	Austria, Belgium, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Lithuania, Luxembourg, Netherlands, Poland, Portugal, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Moldova, Montenegro, Norway, Russia, Switzerland, Ukraine	North America, Asia, Sub‐Saharan Africa		Atkinson (2018)
59.	Pityogenes bistridentatus	Austria, Bulgaria, Croatia, Cyprus, Czech Republic, France, Germany, Greece, Hungary, Italy, Poland, Romania, Slovakia, Slovenia, Spain	Albania, Bosnia‐Herzegovina, North Macedonia, Montenegro, Russia, Switzerland, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
60.	Pityogenes calcaratus	Croatia, France, Greece, Italy, Malta, Portugal, Slovenia, Spain	Russia, Ukraine	Asia, North Africa	SI NPPO: The species was confirmed as present based on the entomological collection of the Biotechnical Faculty, Department of Forestry and Renewable Forest Resources	Alonso‐Zarazaga et al. (2017)
61.	Pityogenes chalcographus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Netherlands, Poland, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Moldova, Montenegro, Norway, Russia, Switzerland	Asia		Alonso‐Zarazaga et al. (2017), Vorst et al. (2008), Vorst (2010) and Heijerman and Noordijk (2018, 2019)
62.	Pityogenes conjunctus	Austria, Bulgaria, Czech Republic, France, Germany, Hungary, Italy, Romania, Slovakia, Slovenia	Switzerland, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
63.	Pityogenes irkutensis irkutensis	Finland, Sweden	Norway, Russia, Turkey	Asia	FI NPPO: occurred in Finland before 1960. Vulnerable in Finland according to the Finnish red‐list. SE NPPO: established in Sweden with old records e.g. from 1937. Red‐listed in Sweden. Should be on the EU list.	Alonso‐Zarazaga et al. (2017), de Jong et al. (2014), Hyvärinen et al. (2019), ArtDatabanken (2015), ArtDatabanken SLU (2019c,g), and Voolma et al. (2004)
64.	Pityogenes irkutensis monacensis	Austria, Bulgaria, Czech Republic, Germany, Lithuania, Poland, Romania	Montenegro, Serbia, Switzerland, Ukraine			Alonso‐Zarazaga et al. (2017)
65.	Pityogenes porifrons	Cyprus, Greece				Alonso‐Zarazaga et al. (2017)
66.	Pityogenes quadridens	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Poland, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Moldova, Montenegro, Norway, Russia, Switzerland, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
67.	Pityogenes saalasi	Finland, Poland, Sweden	Norway, Russia	Asia	FI NPPO: occurred in Finland before 1960. SE NPPO: established in Sweden with old records e.g. from 1944. Should be on the EU list.	Alonso‐Zarazaga et al. (2017), de Jong et al. (2014), Hyvärinen et al. (2019), ArtDatabanken SLU (2019d), and Voolma et al. (2004)
68.	Pityogenes trepanatus	Austria, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Hungary, Italy, Latvia, Netherlands, Poland, Portugal, Romania, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, Montenegro, Norway, Russia, Switzerland, Ukraine		SI NPPO: The presence of the species was confirmed based on the entomological collection of the Biotechnical Faculty, Department of Forestry and Renewable Forest Resources	Alonso‐Zarazaga et al. (2017), Vorst et al. (2008), Vorst (2010) and Heijerman and Noordijk (2016)
69.	Pityokteines curvidens	Austria, Belgium, Bulgaria, Croatia, Czech Republic, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Luxembourg, Poland, Portugal, Romania, Slovenia, Spain	Belarus, Bosnia‐Herzegovina, North Macedonia, Montenegro, Russia, Serbia, Switzerland, Ukraine	South America, Asia, Sub‐Saharan Africa		Atkinson (2018)
70.	Pityokteines spinidens	Austria, Belgium Bulgaria, Croatia, Czech Republic, France, Germany, Greece, Italy, Netherlands, Poland, Romania, Slovakia, Slovenia, Spain	Bosnia‐Herzegovina, Montenegro, Russia, Serbia, Switzerland	Asia	BE NPPO: the species is present in Belgium. SI NPPO: the species is present in Slovenia	Alonso‐Zarazaga et al. (2017) and Heijerman and Noordijk (2017)
71.	Pityokteines vorontzowi	Austria, Belgium, Bulgaria, Croatia, Czech Republic, France, Germany, Greece, Hungary, Italy, Poland, Romania, Slovakia, Slovenia, Spain	Bosnia‐Herzegovina, North Macedonia, Montenegro, Russia, Switzerland, Ukraine	Asia		Alonso‐Zarazaga et al. (2017) and Moucheron et al. (2018)
72.	Pityophthorus balcanicus	Bulgaria, Czech Republic, Romania, Slovakia, Slovenia	Albania, Bosnia‐Herzegovina, North Macedonia, Montenegro, Serbia			Alonso‐Zarazaga et al. (2017) and Titovšek (1983)
73.	Pityophthorus buyssoni angeri	France (Mainland and Corsica)			FR NPPO: endemic species in Corsica.	Alonso‐Zarazaga et al. (2017) and de Jong et al. (2014)
74.	Pityophthorus buyssoni buyssoni	Bulgaria, France, Greece, Italy, Spain	Switzerland		FR NPPO: the species is widespread in France and Europe	Alonso‐Zarazaga et al. (2017)
75.	Pityophthorus carniolicus	Austria, Croatia, Czech Republic, France, Germany, Hungary, Italy, Poland, Slovakia, Slovenia				Alonso‐Zarazaga et al. (2017)
76.	Pityophthorus cephalonicae	Greece, Poland				Alonso‐Zarazaga et al. (2017)
77.	Pityophthorus exsculptus	Austria, Bulgaria, Czech Republic, France, Germany, Hungary, Italy, Poland, Slovakia	Bosnia‐Herzegovina, Switzerland			Alonso‐Zarazaga et al. (2017)
78.	Pityophthorus glabratus	Austria, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Italy, Lithuania, Luxembourg, Netherlands, Poland, Slovakia, Spain, Sweden	Belarus, Montenegro, Norway, Russia, Serbia, Switzerland, Ukraine	Asia, North Africa		Alonso‐Zarazaga et al. (2017) and Vorst (2010)
79.	Pityophthorus henscheli	Austria, Bulgaria, France, Germany, Greece, Italy, Romania	Bosnia, Montenegro, Switzerland, Ukraine			Alonso‐Zarazaga et al. (2017)
80.	Pityophthorus lichtensteinii	Austria, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Poland, Slovakia, Slovenia, Spain, Sweden, UK	Bosnia‐Herzegovina, North Macedonia, Montenegro, Norway, Russia, Switzerland, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
81.	Pityophthorus micrographus micrographus	Austria, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Luxembourg, Poland, Romania, Spain, Sweden, UK	Belarus, Montenegro, Norway, Russia, Serbia, Switzerland	Asia		Alonso‐Zarazaga et al. (2017)
82.	Pityophthorus morosovi	Austria, Czech Republic, Finland, Latvia, Poland, Sweden	Russia	Asia		Alonso‐Zarazaga et al. (2017)
83.	Pityophthorus pinsapo	Spain				Alonso‐Zarazaga et al. (2017)
84.	Pityophthorus pityographus pityographus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, France, Germany, Greece, Hungary, Italy, Poland, Romania, Slovakia, Slovenia, Spain, Sweden	Bosnia‐Herzegovina, North Macedonia, Montenegro, Switzerland, Ukraine			Alonso‐Zarazaga et al. (2017) and Ericson (2010)
85.	Pityophthorus pubescens	Austria, Belgium, Croatia, Cyprus, Czech Republic, Denmark, France, Germany, Greece, Hungary, Ireland, Italy, Netherlands, Poland, Slovakia, Slovenia, Spain, Sweden, UK	Norway, Switzerland	Asia, North Africa		Alonso‐Zarazaga et al. (2017)
86.	Pityophthorus traegardhi	Austria, Estonia, Finland, Poland, Sweden	Norway, Russia	Asia		Alonso‐Zarazaga et al. (2017)
87.	Polygraphus grandiclava	Austria, Belgium, Bulgaria, Croatia, Czech Republic, France, Germany, Hungary, Italy, Netherlands, Poland, Romania, Slovakia, Slovenia	Bosnia‐Herzegovina, Moldova, Montenegro, Russia, Serbia, Switzerland, Ukraine			Alonso‐Zarazaga et al. (2017), Heijerman and Noordijk (2019), Vorst et al. (2008) and Vorst (2010)
88.	Polygraphus poligraphus	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Poland, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, Moldova, Montenegro, Norway, Russia, Switzerland, Ukraine	Asia, Sub‐Saharan Africa		Alonso‐Zarazaga et al. (2017), Vorst (2010), Titovšek (1988) and Martikainen et al. (1999)
89.	Polygraphus punctifrons	Czech Republic, Estonia, Finland, Latvia, Poland, Slovakia, Sweden	Norway, Russia, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
90.	Polygraphus subopacus	Austria, Bulgaria, Czech Republic, Estonia, Finland, France, Germany, Hungary, Latvia, Poland, Slovakia, Sweden	Montenegro, Norway, Russia, Switzerland	Asia		Alonso‐Zarazaga et al. (2017)
91.	Tomicus destruens	Croatia, Cyprus, France, Greece, Italy, Portugal, Slovenia, Spain	Russia, Ukraine	Asia, North Africa		Alonso‐Zarazaga et al. (2017)
92.	Tomicus minor	Austria, Belgium, Bulgaria, Croatia, Cyprus, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Poland, Portugal, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Montenegro, Norway, Russia, Switzerland, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
93.	Tomicus piniperda	Austria, Belgium, Bulgaria, Croatia, Cyprus, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Poland, Portugal, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Montenegro, Norway, Russia, Switzerland, Ukraine	North America, Asia, North Africa		Atkinson (2018)
94.	Trypodendron laeve	Austria, Czech Republic, Estonia, Finland, Germany, Latvia, Poland, Romania, Slovakia, Sweden	Norway, Russia, Switzerland	Asia		Alonso‐Zarazaga et al. (2017)
95.	Trypodendron lineatum	Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Ireland, Italy, Latvia, Lithuania, Netherlands, Poland, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Belarus, Bosnia‐Herzegovina, North Macedonia, Montenegro, Norway, Russia, Switzerland	North America, Asia, North Africa		Atkinson (2018)
96.	Xyleborinus saxesenii	Austria, Belgium, Bulgaria, Croatia, Denmark, Czech Republic, France, Germany, Greece, Hungary, Italy, Netherlands, Poland, Slovenia, Spain, Sweden, UK	Albania, Russia, Ukraine	North America, Sout h America, Asia, North Africa, Sub‐Saharan Africa, Oceania		Atkinson (2018), Heijerman and Noordijk (2016, 2017, 2018, 2019), Vorst et al. (2008), Vorst (2010) and Wood and Bright (1992)
97.	Xyleborus eurygraphus	Austria, Croatia, Czech Republic, France, Germany, Greece, Hungary, Italy, Luxembourg, Netherlands, Poland, Portugal, Slovakia, Spain	Bosnia‐Herzegovina, North Macedonia, Moldova, Montenegro, Russia, Serbia, Switzerland, Ukraine	Asia, North Africa	NL NPPO: the presence of X. eurygraphus in the NL is questionable. All historical findings are associated with wood imports and it is, thus, questionable whether a viable population has established (Heijerman, 2010).	Alonso‐Zarazaga et al. (2017)
98.	Xylechinus pilosus	Austria, Belgium, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Italy, Latvia, Poland, Romania, Slovakia, Slovenia, Spain, Sweden, UK	Montenegro, Norway, Russia, Ukraine	Asia		Alonso‐Zarazaga et al. (2017)
99.	Xylosandrus germanus	Austria, Belgium, Croatia, Czech Republic, Denmark, France, Germany, Hungary, Italy, Netherlands, Poland, Romania, Slovenia, Spain, UK	Russia, Switzerland, Ukraine	North America, Asia	Introduced into Europe	Atkinson (2018), Jurc et al. (2010) and Björklund and Boberg (2017)

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1. Commodity risk assessment of Persea americana from Israel.

Authors: Claude Bragard; Katharina Dehnen-Schmutz; Francesco Di Serio; Paolo Gonthier; Marie-Agnès Jacques; Josep Anton Jaques Miret; Annemarie Fejer Justesen; Alan MacLeod; Christer Sven Magnusson; Panagiotis Milonas; Juan A Navas-Cortes; Stephen Parnell; Roel Potting; Philippe Lucien Reignault; Hans-Hermann Thulke; Wopke Van der Werf; Antonio Vicent Civera; Lucia Zappalà; Pedro Gómez; Andrea Lucchi; Gregor Urek; Sara Tramontini; Olaf Mosbach-Schulz; Eduardo de la Peña; Jonathan Yuen
Journal: EFSA J Date: 2021-02-03

1 in total