Zhiyong Li1, Ute Hentschel2, Nicole Webste3, Julie Olson4, Max Häggblom5. 1. State Key Laboratory of Microbial Metabolism, School of Life Sciences & Biotechnology, Shanghai Jiao Tong University, Dongchuan Road 800, Shanghai 200240, P.R. China. 2. RD3 Marine Symbioses, GEOMAR Helmholtz Centre for Ocean Research Kiel, Düsternbrooker Weg 20, 24105 Kiel, Germany. 3. Australian Institute of Marine Science, PMB 3, Townsville MC QLD 4810, Australia. 4. Department of Biological Sciences, University of Alabama, Tuscaloosa, AL 35487, USA. 5. Department of Biochemistry and Microbiology, Rutgers University, School of Environmental and Biological Sciences, 76 Lipman Drive, New Brunswick NJ 08901, USA.