Lepidopteran biodiversity of Ethiopia: current knowledge and future perspectives.

Lepidoptera is the second largest order of insects. Encompassing moths and butterflies, it is regarded as one of the most important components of biodiversity. Here, an updated comprehensive overview of Lepidoptera recorded in Ethiopia is presented, composed of 2,438 taxa in 48 families, of which 664 are endemic. Records were compiled from various literature sources and website databases. Although still being far from complete, this review provides important baseline data for understanding zoogeographic patterns and thus for undertaking effective conservation action. Further research on Ethiopian Lepidoptera is encouraged. Tesfu Fekensa Tujuba, Andrea Sciarretta, Axel Hausmann, Getnet Atenafu Abate.

Introduction

Ethiopia is among the largest countries in the African continent, located in the horn of Africa, covering a total area of 1,127,127 km2 (Gordon and Carillet 2003; EBI 2015; Tesfu et al. 2018). It belongs to the Afro-tropical Region (former Ethiopian Region) and, based on the bioclimatic classification of Burgess et al. (2004), comprises the zones “Sahelian Savanna”, “Somalian Xeric Bushland and Shrubland” and “Ethiopian Montane forest and Alpine Moorland” (Hacker 2019). The country’s topography is very diverse, with 20 mountains peaks above 4,000 meters. The highest mountain, Ras Dashen, peaks 4,620 m above sea-level, the fourth-highest in Africa, whilst the third-lowest point in Africa, the Danakil Depression, reaches down to 125 m below sea level. The dominating topographic element is the vast and fertile central highland that accounts for 37% of the land area of the country with an average elevation from 1,500 to 2,400 m that deserved the country to be known as ‘roof of Africa’. It is the largest block of land above 1,500 m in Africa (Clausnitzer and Dijkstra 2005), dissected by the Great Rift Valley and surrounded by lowlands along the periphery (Gordon and Carillet 2003). The mean annual rainfall ranges from 500 mm to 2,800 mm and the mean annual temperatures range from around 10° to above 30 °C. Because of these diverging abiotic parameters, the country is endowed with an amazingly diversity of plant, animal and microbial organisms (EBI 2015). According to Clausnitzer (2014), the rate of endemism in Ethiopia’s flora and fauna is exceptionally high as a result of vast highlands being isolated by the surrounding dry lowlands. Only the most eurytopic and mobile species (usually those of the lowlands) tend to be found in both Ethiopia and the rest of tropical Africa. In the same manner, Kravchenko et al. (2007) stated that the territory of Ethiopia hosts an extraordinarily diverse landscape including high mountains, lowlands, deserts and tropical rain forests that resulted in a hyperdiverse fauna and flora. Likewise, in consequence of its rich biodiversity, Ethiopia is acknowledged as one of the 20-mega-biodiverse countries in the world (Mittermeier et al. 2011; Tesfu et al. 2018).

represent the second largest insect order, which consists of approximately 140 different families and 160,000 species that have been described and recognised worldwide, so far (Biodiversity Institute of Ontario 2006; Kristensen et al. 2007; Nieukerken et al. 2011). comprise nearly 17% of all insect species, and some recent estimates suggest that the real number of species would set up to 500,000 species (Brandao et al. 2009).

The aims of this paper are to give an updated comprehensive presentation of the actual knowledge of Ethiopian and to provide some estimates for the expected biodiversity of this major insect order in the country.

Materials and methods

The present review is based on all pertinent published scientific papers. In addition, records from up to date and relevant online databases were also included, particularly, records from the Natural History Museum of London website (“NHMUK”: Beccaloni et al. 2003), the Barcode of Life Data Systems (“BOLD”: Ratnasingham and Hebert 2007), the African moth website (Goff 2008), LepiMap (Navarro 2007), the African Butterfly Database (Sáfián et al. 2009), the Afromoth website (De Prins and De Prins 2019) and the Afrotropical Butterflies and Skippers digital encyclopaedia (Williams 2018). In all cases, records were included only when sample identifications were made at specific (or subspecific) level, and the provenience from Ethiopia was clearly indicated. Data from entomological collections but not publicly accessible were not considered.

We followed the classification system and nomenclature (valid names and synonymies) used in De Prins and De Prins (2019) with some updates coming from more recent publications. For , the Afrotropical Butterflies and Skippers digital encyclopaedia (Williams 2018) served as reference. These two outstanding references have also represented the fundamental database and resource for our compilation of the lepidopteran fauna of Ethiopia.

exploration in Ethiopia: from early explorers to present

Many entomologists have contributed to our current knowledge of the Ethiopian fauna. The following selection provides the most significant contributions made by past pioneers and current explorers.

Johann Christoph Friedrich Klug in 1829 was the first to mention Abyssinia, the former name of Ethiopia, in the description of a new species, the butterfly (Klug, 1829) now (Nazari et al. 2011), indicating as locus typicus “in Arabia deserta, in Sinai monte, in Dongala et Habessinia”.

From the mid-nineteenth century, additional descriptions came from few authors such as Félix Edouard Guérin-Méneville (1849), Louis Reiche (1850), and Hippolyte Lucas (1852). However, the most significant advance in the nineteenth century was made by the French entomologist Achille Guenée, who published various contributions between 1852 and 1858. He described 31 new species belonging to the and , based on material collected mainly by Georg Wilhelm Schimper in 1850. In all cases, the locus typicus was indicated as “Abyssinia” (Guénée 1852).

Other important contributions to the study of Ethiopian were made subsquently, many of which have reported the description of new species from specimens collected in the country. For instance, George Hampson described 23 species from different families in the period between 1896 and 1930 (Hampson 1896, 1898, 1899, 1905, 1909, 1910, 1913, 1916, 1918, 1919, 1926, 1930). Edward Meyrick firstly reported Microlepidoptera from the country, with 40 new species, from the material collected during the expeditions carried out by Hugh Scott and Omer-Coper in 1926–1927 (Meyrick 1932). The most important contribution to the study of butteflies was made by Lionel Walter Rothschild and Karl Jordan, during the first decades of the twentieth century, with 34 new taxa (Rothschild 1902, 1926; Rothschild and Jordan 1900, 1903, 1905). Debauche (1937) reported 42 geometrid species from Ethiopia with eight new descriptions. Likewise, Emilio Berio published many papers dedicated to the and of East Africa, describing from Ethiopia 12 and 37 species, respectively (Berio 1939a, 1940a, 1943, 1944, 1945, 1947, 1954, 1962, 1975, 1977), some of them from the localities of Adu-Abuna and Metema, at that time part of Eritrea, but now in Tigray, Northern Ethiopia (Berio 1937, 1939b, 1939c, 1940b, 1973, 1976). Pierre-Claude Rougeot has explored the country several times in 1970s and described 55 new species belonging to various families (Rougeot 1974, 1975, 1977, 1984, Plantrou and Rougeot 1979; Laporte and Rougeot 1981; Rougeot and Laporte 1983). The two French entomologists Bernard Laporte and Claude Herbulot in their publications described from Ethiopia 137 new noctuid (specifically, eight species of , two species of , and 127 ) and 22 new geometrid taxa, respectively (Herbulot 1983, 1993, 2002; Laporte 1974, 1975, 1976, 1978; Rougeot 1977, 1984; Laporte and Rougeot 1981; Rougeot and Laporte 1983; Rougeot et al. 1991).

With the new millennium, the country has awakened a renewed interest from entomologists, which led to the description of 255 new taxa in 18 years. In particular, major contributions to Ethiopian were made by Hermann H. Hacker, with various colleagues, for , and (178 new taxa); David Agassiz for (five new taxa); Jósef Razowski and Pasquale Trematerra for (34 new taxa described); Axel Hausmann, Andrea Sciarretta and Francesco Parisi for (27 new taxa); Ulf Eitschberger and Tomas Melichar for , with eleven new taxa (Hacker and Fibiger 2007; Hacker and Zilli 2007; Haxaire and Melichar 2008; Hausmann et al. 2014, 2016; Hacker et al. 2008, 2012; Razowski and Trematerra 2010, 2012; Hacker and Mey 2010; Hacker 2011, 2013, 2014, 2016, 2019; Eitschberger and Ströhle 2011; Melichar and Řezáč 2015; Eitschberger and Melichar 2016Melichar et al. 2016; Razowski et al. 2018; Agassiz 2019).

Many of these and other minor contributions resulted from dedicated expeditions, such as the “Joint Ethiopian-Russian Biological Expedition” lead by Vasiliy Kravchenko from Tel Aviv University, Israel; the “Ethiopian Insects Project”, between the Ethiopian Wildlife Conservation Authority (EWCA), the Bavarian State Collection of Zoology (ZSM) and the Museum Thomas Witt (MWM) in Munich, Germany; the projects carried out by the Italian entomologists of the University of Molise with EWCA and Ethiopian Biodiversity Institute (Kravchenko et al. 2007; Sciarretta et al. 2014; Hausmann et al. 2016).

Current state of knowledge on Ethiopian

Based on the results of our current review, 2,438 taxa (species or subspecies) are known to occur in Ethiopia hitherto, belonging to 48 families (Table 1; full list at: https://doi.org/10.5281/zenodo.3234617). This number includes 170 taxa which are not reported by the scientific literature but have been extracted from the above-mentioned websites.

Table 1.

Ethiopian families and number of taxa (species and subspecies) reported.

No.	Family	Total number of taxa	Common name
1	Bedelliidae	1	Narrow-winged moths
2	Blastobasidae	2	Scavenger moths
3	Bombycidae	2	Silkworm moths
4	Brahmaeidae	2	Brahmin moths
5	Carposinidae	1	Fruitworm moths
6	Choreutidae	2	Metalmark moths
7	Cosmopterigidae	4	Cosmet moths
8	Cossidae	17	Carpenterworm moths
9	Crambidae	109	Grass moths
10	Depressariidae	2	Flat-bodied moths
11	Drepanidae	1	Hook-tips
12	Elachistidae	1	Grass miner moths
13	Epermeniidae	1	Fringe-tufted moths
14	Erebidae	523	Tiger moths
15	Eupterotidae	8	Snout moths
16	Euteliidae	10	Euteliid moths
17	Gelechiidae	10	Twirler moths
18	Geometridae	306	Geometer moths
19	Glyphipterigidae	1	Sedge moths
20	Gracillariidae	13	Leafminer moths
21	Hesperiidae	36	Skipper butterflies
22	Hyblaeidae	1	Teak moths
23	Lasiocampidae	38	Lappet moths
24	Limacodidae	15	Slug caterpillar moths
25	Lycaenidae	116	Gossamer-winged butterflies
26	Lyonetiidae	2	Lyonet moths
27	Metarbelidae	4	Wood-borer moths
28	Noctuidae	471	Owlet moths
29	Nolidae	85	Tuft moths
30	Notodontidae	28	Prominent moths
31	Nymphalidae	178	Brush-footed butterflies
32	Oecophoridae	1	Concealer boths
33	Papilionidae	17	Swallowtail butterflies
34	Pieridae	79	Yellows, Whites, & Sulphurs
35	Plutellidae	5	Diamondback moths
36	Psychidae	6	Bagworm moths
37	Pterophoridae	39	Plume moths
38	Pyralidae	31	Snout moths
38	Saturniidae	53	Emperor moths
40	Scythrididae	7	Flower moths
41	Sesiidae	6	Clearwing moths
42	Sphingidae	81	Hawk moths
43	Thyrididae	9	Picture-winged leaf moths
44	Tineidae	38	Fungus moths
45	Tortricidae	60	Leafroller moths
46	Uraniidae	3	Swallowtail moths
47	Yponomeutidae	6	Ermine moths
48	Zygaenidae	8	Burnet moths
Total		2438

In particular, 929 species or subspecies were described from type specimens collected in Ethiopia, 131 of them, mostly butterflies, at subspecific level (Table 2). It is interesting to note that endemic taxa number 664, approximately 27% of the total . This high number can be explained by the particular physical and biogeographical history of the country and a broad range of different ecosystems with great diversity of habitats.

Table 2.

List of taxa originally described from Ethiopia (only valid names are listed). An asterisk (*) denotes that the town is in Sudan, but the river originates in Ethiopia. The type locality is recorded with the corrected spelling or current locality name in square brackets. The endemic taxa from Ethiopia are indicated with E in the last column. Synonymies are not reported. De Prins and De Prins (2019) and Williams (2018) have been used as a basic reference for the preparation of the list.

	Family	Taxon	Author	Type Locality
1	Blastobasidae	Blastobasis eridryas	Meyrick, 1932	Mt Chillálo	E
2	Brahmaeidae	Dactyloceras richinii	Berio, 1940	Adi Abuna [in Tigray, Ethiopia]
3	Carposinidae	Carposina candace	Meyrick, 1932	Jem-Jem Forest	E
4	Choreutidae	Choreutis argyrastra	Meyrick, 1932	Mt Zukwala/Cuqala	E
5		Telosphrantis aethiopica	Meyrick, 1932	Mt Chillálo	E
6	Cosmopterigidae	Ascalenia secretifera	Meyrick, 1932	Mt Chillálo	E
7		Cosmopterix derrai	Koster, 2016	14 km S of Debre Tabor, Alemsaga Forest	E
8		Cosmopterix epismaragda	Meyrick, 1932	Jem-Jem Forest	E
9	Cossidae	Aethalopteryx obscurascens	(Gaede, 1930)	Centr. Abyss., Maraquo
10		Afroarabiella strohlei	Yakovlev & Witt, 2016	Turmi, Mango Lodge	E
11		Azygophleps brehmi	Yakovlev & Witt, 2016	Bale Mountains, Karcha near Rira	E
12		Camellocossus abyssinica	(Hampson, 1910)	Abyssinia [Ethiopia]
13		Camellocossus lalibela	Yakovlev & Witt, 2017	Arba Minch	E
14		Camellocossus strohlei	Yakovlev & Witt, 2017	Arba Minch	E
15		Macrocossus sidamo	Rougeot, 1977	near Kébré-Mengist [Kebre Mengist]	E
16		Oreocossus ungemachi	Rougeot, 1977	Ioubdo, Birbir	E
17		Strigocossus kushit	Yakovlev, 2011	Ethiopia SE, Bale, 11 km SW Goba, Bale Mts	E
18	Crambidae	Adelpherupa aethiopicalis	Maes, 2002	SW Abyssinia [Ethiopia], Djimma [Jimma]	E
19		Agathodes dufayi	Rougeot, 1977	Koffolé [Kofale]	E
20		Alphacrambus cristatus	Bassi, 1995	Maraqo	E
21		Ancylolomia jacquelinae	Rougeot, 1984	Arba Minch	E
22		Ancylolomia shafferi	Rougeot, 1977	Koffolé [Kofale]	E
23		Ancylolomia shefferialis	Rougeot, 1984	Bahar Dar	E
24		Chilo luniferalis	Hampson, 1896	Abyssinia [Ethiopia]	E
25		Classeya aphrodite	Błeszyński, 1964	Dire Dawa	E
26		Crambus arnaudiae	Rougeot, 1977	Koffolé [Kofale]	E
27		Crambus bachi	Bassi, 2012	Bahar Dar, Lake Tana	E
28		Crambus bellinii	Bassi, 2014	Bale Mts, Sanetti Plateau
29		Crambus boislamberti	Rougeot, 1977	Dinsho Reserve	E
30		Crambus dedalus	Bassi, 2000	Karsan, Kollubi	E
31		Crambus descarpentriesi	(Rougeot, 1977)	Koffolé [Koffale]	E
32		Crambus jupiter	Błeszyński, 1963	Ethiopia SW, Gamu-Gofa, Konso	E
33		Crambus netuncus	Bassi, 2012	Near Debra Libanos	E
34		Crambus richteri	Błeszyński, 1963	Kaffa, Ghimira	E
35		Dembea venulosella	Ragonot, 1888	Abyssinia [Ethiopia]
36		Euchromius donum	Schouten, 1988	Haro-Ali, Gurra	E
37		Euclasta sidamona	Rougeot, 1977	Koffolé [Koffale]	E
38		Euctenospila castalis	Warren, 1892	Abyssinia [Ethiopia]
39		Leucinodes ethiopica	Mally, Korycinska, Agassiz, Hall, Hodgetts & Nuss, 2015	Dire Dawa Region, Dire Dawa District, Dire Dawa
40		Lygropia nigricornis	Hampson, 1898	Abyssinia [Ethiopia]
41		Noorda trimaculalis	Amsel, 1965	Ethiopia SW, Gammu-Gofa, Konso	E
42	Crambidae	Noorda unipunctalis	Amsel, 1963	Konso	E
43		Pagyda pulvereiumbralis	(Hampson, 1918)	Diré Daouá [Dire Dawa]
44		Pediasia ferruginea	Błeszyński, 1963	Kaffa, Gembi
45		Pediasia simiensis	Błeszyński, 1962	Soddu Province, Wolamo [Walita]	E
46		Prionapteryx selenalis	(Hampson, 1919)	Taddecha Mullka	E
47		Prionotalis friesei	Błeszyński, 1963	Ethiopia SW, Gamu-Gofa, Konso	E
48		Tegostoma richteri	Amsel, 1963	Awash	E
49	Depressariidae	Odites aethiopicus	Lvovsky, 2001	Kaffa, Gembi
50	Elachistidae	Elachista delocharis	Meyrick, 1932	Jem-Jem Forest	E
51	Erebidae	Achaea monodi	Laporte, 1975	near Kebré-Mengist [Kibre Mengist]	E
52		Afrasura rivulosa ethiopica	Durante, 2009	Menegesha-Suba state Forest	E
53		Afrasura indecisa orientalis	Durante, 2009	Menegesha-Suba state Forest	E
54		Afrasura terlinea	Durante, 2009	Langano Lake	E
55		Afrojavanica kostlani	(Gaede, 1923)	Adis-Abeba
56		Alpenus geminipuncta	(Hampson, 1916)	Abyssinia [Ethiopia]	E
57		Amata alicia	(Butler, 1876)	Abyssinia [Ethiopia]
58		Amata magrettii	Berio, 1937	Metema [in Tigray, Ethiopia]	E
59		Amata rufina	(Oberthür, 1878)	Abyssinia [Ethiopia]
60		Amata shoa	(Hampson, 1898)	Abyssinia [Ethiopia]
61		Amata velatipennis	Walker, 1865	Marako
62		Amphicallia kostlani	Strand, 1911	Gipfel des Sugyala	E
63		Amsacta nigrisignata	Gaede, 1923	Addis Ababa	E
64		Amsactarctia radiosa	(Pagenstecher, 1903)	Darassum
65		Anomis sabulifera	(Guenée, 1852)	Abyssinia [Ethiopia]
66		Antiophlebia bourgognei	Laporte, 1975	Arba Minch	E
67		Aroa quadriplagata	Pagenstecher, 1903	Galata	E
68		Asura xanthophaea	Toulgoët, 1977	Ethiopia	E
69		Beriodesma smithii	(Holland, 1897)	River Darde
70		Brunia birketsmithi	(Toulgoët, 1977)	Kébré-Mengist [Kibre Mengist]	E
71		Brunia dorsti	(Toulgoët, 1977)	Kébré-Mengist [Kibre Mengist]	E
72		Callophisma viettei	Laporte, 1975	Arba Minch	E
73		Carcinarctia rougeoti	Toulgoët, 1977	Bale Reserve, Dinsho	E
74		Casama impura	(Hering, 1926)	Abyssinia [Ethiopia]
75		Cautatha abyssinia	Hacker, Fiebig & Stadie, 2019	Reg. South Nations, Bonga Guesthouse
76		Cautatha bifasciata	Hacker, Fiebig & Stadie, 2019	Reg. South Nations, road Shishinda-Bonga, 6 km, w Wushwush	E
77		Cerocala confusa	Warren, 1913	Abyssinia [Ethiopia]	E
78		Clytie thibauti	Laporte, 1991	Kibre Mengist	E
79		Corgatha hyperxantha	Hacker, Fiebig & Stadie, 2019	Reg. South Nations, Bonga Guesthouse	E
80		Corgatha minutulana	Hacker, 2019	Southern Prov., 6 km ENE Weyto, Segen river
81		Cortyta canescens septentrionalis	Hacker, 2016	12 km W of Jinka, near border of Mago National Park
82		Crambiforma leucostrepta	Hampson, 1926	Harrar	E
83		Crypsotidia digitata	Kühne, 2005	Harar	E
84		Crypsotidia gigantea	Kühne, 2005	Harar
85		Ctenusa curvilinea	Hampson, 1913	Taddecha Mullka
86		Cyana abyssinica	Karisch, 2003	Akaki River, Addis Ababa	E
87	Erebidae	Cyana ethiopica	Karisch, 2013	near Kebré-Mengist [Kibre Mengist]	E
88		Dasychira grisea	Pagenstecher, 1903	Bone	E
89		Dasychira plesia	Collenette, 1938	Abyssinia [Ethiopia]
90		Digama meridionalis deliae	Berio, 1939	Adu-Abuna [in Tigray, Ethiopia]	E
91		Donuctenusa fiorii	Berio, 1940	Ogaden, Uarder [Warder]	E
92		Enargeiosia elegans	(Butler, 1877)	Atbara*
93		Eublemma accedens aethiopica	Hacker, 2019	Ethiopia, 3 km N Turmi, Mango Camping Site
94		Eublemma aethiopiana	Hacker, 2019	Jinka, Mago Nat. Park, Magoriverside
95		Eublemma baccatrix	Hacker, 2019	Southern Prov., 2.6 km EE Wondo Genet
96		Eublemma collacteana	Hacker, 2019	12 km W Jimma, border Mago Nat. Park
97		Eublemma costivinata	Berio, 1945	Borana Nagelli [Borena Nagelle]	E
98		Eublemma diredaoua	Hacker, 2019	Dire Daoua, Abyssinia	E
99		Eublemma ferruginata	Hacker, 2019	20 ESE Sashamane, Wendo Genet
100		Eublemma heteropaura	Hacker, 2019	Oromia, 7 km NW Yabelo
101		Eublemma joergmuelleri	Hacker & Schrier, 2019	Ethiopia, Awash N.P., Headquarter	E
102		Eublemma perturbata	Hacker, 2019	Oromia prov., 6.5 km ne Shebe
103		Eublemma plectoversa	Hacker, 2019	8 km N Turmi
104		Eublemma schreieri	Hacker, 2019	Oromia, 1km W vill. Aluweya
105		Eublemma sidamonia	Hacker, Fiebig & Stadie, 2019	Sidamo, Yabello, vic. 6km SO near Deritu village	E
106		Eublemma siticulina	Hacker, 2019	Dire Daoua, Abyssinia	E
107		Eublemma uhlenhuthi	Wiltshire, 1988	Abyssinia, Dire Daoua [dire-dawa]
108		Euproctis chrysophaea	(Walker, 1865)	Abyssinia [Ethiopia]
109		Eyralpenus scioana	(Oberthür, 1880)	Scioa [Shoa]
110		Galtara doriae	(Oberthür, 1880)	Mahal Uonz, between Harrar and Addis Abeba [Awash River]
111		Kenyarctia melanogastra	(Holland, 1897)	Gof [Gofa]
112		Hypena abyssinialis	Guenée, 1854	Abyssinia [Ethiopia]
113		Hypena padelekorum	Lödl, 1995	Djem-Djem [Jem Jem] Forest	E
114		Hypena philippi	Laporte, 1991	Arba Minch	E
115		Hyposada zavattarii	Berio, 1944	Gondaraba
116		Hypotacha fiorii	Berio, 1943	Diredaua [Dire Dawa]
117		Hypotacha glaucata	(Holland, 1897)	Sjeikh Husein [Shek Hussein]
118		Ischnarctia cinerea	(Pagenstecher, 1903)	Gogoru
119		Laelia dabano	Collenette, 1934	Dabano River
120		Lithacodia awassensis	Berio, 1984	Awassa Lake	E
121		Lithacodia persubtilis	Berio, 1984	Kebré-Mengist	E
122		Marcipa rougeoti	Pelletier, 1975	Kebré-Mengist	E
123		Metachrostis debivar	(Berio, 1947)	Ogaden, Ualual [Walwal]	E
124		Metachrostis phaeographa	Hacker, 2011	12 km W of Jinka, border Mago National Park
125		Metarctia carmel	Kiriakoff, 1957	SW Abyssinia [Ethiopia], Kambatta	E
126	Erebidae	Metarctia gada	Rougeot, 1977	Dinsho Reserve, Réserve de Balé	E
127		Metarctia haematricha	Hampson, 1905	Kutai Metha
128		Metarctia kumasina	Strand, 1920	Zegi Tsana [Zegie Tana]
129		Metarctia negusi	Kiriakoff, 1957	Abyssinia [Ethiopia]	E
130		Metarctia noctis	Druce, 1910	Diré Daouá [Dire Dawa]	E
131		Metarctia saalfeldi	Kiriakoff, 1960	Villagio	E
132		Metarctia unicolor	(Oberthür, 1880)	Oromo Country, Fin-Fekéré
133		Micralarctia punctulatum purus	(Butler, 1878)	Abyssinia [Ethiopia]
134		Oediblemma peregrina	Hacker, Fiebig & Stadie, 2019	Reg. South Nations, Sheiko Forest Road Teppi Mizan Teferi	E
135		Ophiusa dianaris	(Guenée, 1852)	Abyssinia [Ethiopia]
136		Pantydia dufayi	Laporte, 1975	Near Koffolé [Koffalé]
137		Paramarbla abyssinica	Collenette, 1956	Birbir, Joubdo [Yubdo]	E
138		Paraonagylla zavattarii	Berio, 1939	Neghelli [Nagelle]	E
139		Pericyma schreieri	Hacker, 2016	Gamu-Gofa Province, 3 km N of Turmi
140		Phytometra angensteini	Hacker, 2019	Arba Minch	E
141		Plecopterodes melliflua	(Holland, 1897)	Sjeikh Husein [Shek Hussein]
142		Plecopterodes molybdena	Berio, 1954	Gorgorà, Lake Tana	E
143		Podomachla antinorii	(Oberthür, 1880)	Mahal Uonz [Awash River]
144		Polymona rufifemur ellisoni	Collenette, 1938	Abyssinia [Ethiopia]
145		Proluta ethiopica	(Hacker, 2011)	Arba Minch Region, Omo Province, Gemu Gofa
146		Pseudomicrodes varia	Berio, 1944	Elolo
147		Pteredoa atripalpia	Hampson, 1910	Atbara River
148		Rhabdophera exarata	(Mabille, 1890)	Abyssinia [Ethiopia]
149		Ruanda nuda	(Holland, 1897)	River Darde
150		Seydelia geometrica	(Oberthür, 1883)	Scioa [Shoa]
151		Spilosoma mediopunctata	(Pagenstecher, 1903)	Arbarone
152		Spilosoma quadrimacula	Toulgoët, 1977	Lalokéli	E
153		Stenilema aurantiaca	Hampson, 1909	Abyssinia [Ethiopia]
154		Stenilema hailesellassiei	(Birket-Smith, 1965)	Addis Ababa, University College Campus	E
155		Stracena aegrota	Le Cerf, 1922	Harar	E
156		Stracilla translucida	(Oberthür, 1880)	Scioa [Shoa], Mahal Uonz
157		Syngatha eremita	Hacker, Fiebig & Stadie, 2019	Reg. South Nations, Bonga Guesthouse	E
158		Syngatha parascotoides	Hacker, 2019	12 km W Jinka, border Mago National Park
159		Syngatha simplicicata	Hacker, Fiebig & Stadie, 2019	Reg. South Nations, Sheiko Forest Road Teppi Mizan Teferi	E
160		Tegiapa ambiguosa	Hacker, Fiebig & Stadie, 2019	Reg. South Nations, road Shishinda-Bonga, 6 km W Wushwush
161		Tegiapa obliqua	Hacker, Fiebig & Stadie, 2019	Sidamo, Yabello vic., 10km W road to Konso
162		Tegiapa schreieri	Hacker, 2019	Oromia Prov., 6 km ESE Jimma
163		Teracotona abyssinica	(Rothschild, 1933)	Central Abyssinia [Ethiopia], Maraco [Marako]
164		Teracotona neumanni	Rothschild, 1933	SW Abyssinia [Ethiopia], Kambatta	E
165	Erebidae	Teracotona postalbida	(Gaede, 1926)	Abyssinia [Ethiopia]	E
166		Teracotona clara rubiginea	(Toulgoët, 1977)	Fisha Genet	E
167		Teracotona seminigra	(Hampson, 1905)	Zegi Tsana [Tana]	E
168		Thyretes negus	Oberthür, 1878	Abyssinia [Ethiopia]
169		Tigreana nathaliannae	Laporte, 1991	Wollo, Ataye	E
170		Tigreana sandrae	Laporte, 1991	Wollo Ataye	E
171		Trigonodes exportata	Guenée, 1852	Abyssinia [Ethiopia]
172		Tytroca alabuensis alabuensis	Wiltshire, 1970	Alabu
173		Tytroca balnearia mutabilis	Hacker, 2016	15 km N of Arba Minch, 2 km after junction to Chencha	E
174		Tytroca heterophysa	Hacker, 2016	Omo Region, Gemu Gofa Province, Arba Minch	E
175		Ulotrichopus phaeoleucus griseus	Kühne, 2005	Addis Ababa
176		Utetheisa amhara	Jordan, 1939	Abyssinia [Ethiopia]
177		Zekelita heteroleuca	Hacker, 2016	Southern Province, 11.2 km W of Bonga	E
178		Zekelita lehmanni magnificaria	Hacker, 2016	10.5 km W of Weyto	E
179		Zekelita nilotica	Hacker, 2016	30 km SE of Bahir Dar, Tisisat above Blue Nile falls	E
180	Eupterotidae	Phiala abyssinica	Aurivillius, 1904	Zegi Tsana [Tana]	E
181		Phiala bergeri	Rougeot, 1975	Bale	E
182		Rhodopteriana abyssinica	(Rothschild, 1917)	Harrar [Harar]
183		Rhodopteriana sidamoensis	Darge, 2013	Sidamo Province, near Mega	E
184	Euteliidae	Eutelia favillatrix	(Guenée, 1852)	Abyssinia [Ethiopia]
185		Stenosticta schreieri	Hacker, 2010	3 km N Turmi, Mango Camping Site	E
186	Gelechiidae	Aphanostola maxima	Bidzilya & Mey, 2016	Lake Tana, Bahir Dar	E
187		Chrysoesthia parilis	(Vári, 1963)	Little Akaki River, near Addis Ababa	E
188		Stegasta sattleri	Bidzilya & Mey, 2011	Addis Ababa
189		Stomopteryx ochrosema	Meyrick, 1932	Addis Alam [Alem], ca. 20 miles W. of Addis Ababa	E
190		Chiasmia abyssinica	Krüger, 2001	Harrar [Harar]	E
191		Chiasmia procidata	(Guenée, 1858)	Abyssinia [Ethiopia]
192		Chiasmia streniata	(Guenée, 1858)	Abyssinia [Ethiopia]
193		Chiasmia trinotatula	Krüger, 2001	Kabarutar, 56 miles W of Lake Tana	E
194		Cleora oculata sidamo	Herbulot, 1977	Kébré-Mengist [Kibre Mengist]	E
195		Cleora pavlitzkiae etesiae	Fletcher, 1967	Harar
196		Coenina dentataria	Swinhoe, 1904	Abyssinia [Ethiopia]
197		Comibaena theodori	Hausmann & Parisi, 2014	Kaffa Province, 10 km N of Bonga	E
198		Drepanogynis nigerrima	(Swinhoe, 1904)	Abyssinia [Ethiopia]	E
199		Epigynopteryx flavedinaria	(Guenée, 1857)	Abyssinia [Ethiopia]
200		Epigynopteryx rougeoti	Herbulot, 1977	Dinsho Marshes	E
201		Epigynopteryx scotti	Fletcher, 1959	Ethiopia N, Simien, near Mindigabsa	E
202		Erastria marginata	(Swinhoe, 1904)	Abyssinia [Ethiopia]	E
203		Eupithecia angulata	Fletcher, 1951	Harar	E
204		Eupithecia dinshoensis	Herbulot, 1983	Dinsho Col	E
205		Eupithecia incommoda	Herbulot, 1983	Dinsho Reserve	E
206		Eupithecia inquinata	Fletcher, 1950	Lekamti [Naqamte]
207	Geometridae	Eupithecia ochralba	Herbulot, 1983	Dinsho Reserve	E
208		Eupithecia pseudoabbreviata	Fletcher, 1951	Harar	E
209		Eupithecia rougeoti	Herbulot, 1983	Dinsho Reserve	E
210		Eupithecia urbanata	Fletcher, 1956	Harar	E
211		Geodena brunneomarginata	Karisch, 2003	Shoa, 50 km W of Adis Ababa	E
212		Hemistola aetherea	Debauche, 1937	Addis Ababa	E
213		Henicovalva negus	Krüger, 2017	Dire Daoua [Dawa]	E
214		Heterostegane serrata	(Fletcher, 1958)	Diré Daouá [Dire Dawa]
215		Hydrelia candace	Prout, 1929	Addis Ababa	E
216		Hypochrosis chiarinii	(Oberthür, 1883)	Scioa [Shoa]
217		Idaea glomerata	(Prout, 1937)	Abyssinia [Ethiopia]	E
218		Lomographa indularia	(Guenée, 1858)	Abyssinia [Ethiopia]
219		Mimoclystia pudicata cecchii	(Oberthür, 1883)	Scioa [Shoa], Let-Marefia [Jet Marafia]	E
220		Nothofidonia xenoleuca	Prout, 1928	Wolisso, between Hauash [Awash] and Omo	E
221		Odontopera briela	(Debauche, 1937)	Mt Chillálo [Chilalo]	E
222		Odontopera integraria	Guenée, 1858	Abyssinia [Ethiopia]
223		Odontopera protecta	Herbulot, 1983	Dinsho Reserve	E
224		Omphacodes pulchrifimbria pulchritacta	Prout, 1923	Abyssinia [Ethiopia] Central Moraqui [Marako]
225		Oreometra ras	Herbulot, 1983	near Mount Batu	E
226		Piercia zukwalensis	Debauche, 1937	Mt Zukwala/Cuqala
227		Pingasa abyssiniaria	(Guenée, 1858)	Harar
228		Platypepla bifida	Herbulot, 1984	near Kébré-Mengist [Kibre Mengist]	E
229		Platypepla uhlenhuthi	Krüger, 2001	Diré Daouá [Dire Dawa]	E
230		Prasinocyma aquamarina	Hausmann, Sciarretta & Parisi, 2016	Bale Mts, 10 km S Rira	E
231		Prasinocyma aetheraea	(Debauche, 1937)	Addis Ababa	E
232		Prasinocyma albivenata	Herbulot, 1983	Dinsho Marsh	E
233		Prasinocyma amharensis	Hausmann, Sciarretta & Parisi, 2016	SW Debre Sina & Sembo, Umg. Debre Sina	E
234		Prasinocyma angolica pseudopedicata	Hausmann, Sciarretta & Parisi, 2016	7 km NW Yabello	E
235		Prasinocyma angulifera	Hausmann, Sciarretta & Parisi, 2016	southern Bale Mts, Harenna Forest	E
236		Prasinocyma batesi distans	Hausmann, Sciarretta & Parisi, 2016	Addis Ababa	E
237		Prasinocyma baumgaertneri	Hausmann, Sciarretta & Parisi, 2016	Harenna Forest	E
238		Prasinocyma beryllaria	Hausmann, Sciarretta & Parisi, 2016	13 km W Yabello Motel	E
239		Prasinocyma bongaensis	Hausmann, Sciarretta & Parisi, 2016	Bonga, 12 km E	E
240		Prasinocyma discipuncta	Hausmann, Sciaretta & Parisi, 2016	16 km SW Kibre Mengist	E
241		Prasinocyma fallax	Hausmann, Sciarretta & Parisi, 2016	SW. Debre Sina & Sembo, Umg. Debre Sina	E
242		Prasinocyma fusca	Hausmann, Sciarretta & Parisi, 2016	Harenna Forest	E
243		Prasinocyma gajdacsi	Prout, 1930	Adis Abeba [Addis Ababa]	E
244		Prasinocyma gemmifera	Hausmann, Sciaretta & Parisi, 2016	Wushwush, 7.4 km w	E
245	Geometridae	Prasinocyma germinaria	(Guenée, 1857)	Abyssinia [Ethiopia]
246		Prasinocyma getachewi	Hausmann, Sciarretta & Parisi, 2016	Arba Minch	E
247		Prasinocyma hailei	Debauche, 1937	Addis Ababa	E
248		Prasinocyma immaculata thiaucourti	Herbulot, 1993	Debre Zeit	E
249		Prasinocyma leveneorum	Hausmann, Sciarretta & Parisi, 2016	Harenna Forest, Karcha clearing	E
250		Prasinocyma lutulenta	Hausmann, Sciarretta & Parisi, 2016	Arba Minch	E
251		Prasinocyma magica	Hausmann, Sciarretta & Parisi, 2016	Mago National park	E
252		Prasinocyma monikae	Hausmann, Sciarretta & Parisi, 2016	13 km W Yabello, Motel	E
253		Prasinocyma pedicata aethiopica	Hausmann, Sciarretta & Parisi, 2016	16 km SW Kibre Mengist	E
254		Prasinocyma robusta	Hausmann, Sciarretta & Parisi, 2016	13 km W Yabello, Motel	E
255		Prasinocyma septentrionalis	Hausmann, Sciarretta & Parisi, 2016	Arba Minch	E
256		Prasinocyma shoa shoa	Herbulot, 1993	Debre Zeit	E
257		Prasinocyma shoa yabellensis	Hausmann, Sciarretta & Parisi, 2016	13 km W Yabello, Motel	E
258		Prasinocyma stefani	Hausmann, Sciarretta & Parisi, 2016	Bonga, 12 km E	E
259		Prasinocyma tranquilla	Prout, 1917	NW of Harar, Diredaua [Dire Dawa]	E
260		Prasinocyma trematerrai simienensis	Hausmann, Sciarretta & Parisi, 2016	Semien Mountains, chennek Camp	E
261		Prasinocyma trematerrai trematerrai	Hausmann, Sciarretta & Parisi, 2016	Dinsho	E
262		Problepsis fiebigi	Stadie & Stadie, 2016	Omo Region, Province of Gemu Gofa, Arba Minch	E
263		Problepsis neumanni	Prout, 1932	Djiren, Djimma [Jimma]	E
264		Problepsis sihvoneni	Stadie & Stadie, 2016	Sidamo, 13 km W of Yabello, Motel	E
265		Protosteira decolorata	Herbulot, 1984	Semyen, Sankaber	E
266		Rhodometra labdoides	Herbulot, 1997	Choa [Shoa], Debré Zeit	E
267		Rhodometra plectaria	(Guenée, 1857)	Abyssinia [Ethiopia]
268		Rougeotiella pseudonoctua	Herbulot, 1983	Kébré-Mengist [Kibre Mengist]	E
269		Scopula erymna	Prout, 1928	Gurra, Dagaje	E
270		Scopula scotti	Debauche, 1937	Addis Ababa
271		Scopula silonaria	(Guenée, 1858)	Abyssinia [Ethiopia]
272		Scopula simplificata	Prout, 1928	NE Africa, Ganale River	E
273		Sesquialtera lonchota	Prout, 1931	Diré Daouá [Dire Dawa], NW of Harrar	E
274		Somatina pythiaria	(Guenée, 1857)	Abyssinia [Ethiopia]
275		Tephronia aethiopica	Herbulot, 1983	Shoa, Menagesha Forest	E
276		Traminda neptunaria	(Guenée, 1857)	Abyssinia [Ethiopia]
277		Trimetopia aetheraria	Guenée, 1858	Abyssinia [Ethiopia]	E
278		Xanthisthisa copta	Herbulot, 1977	Boré Forest	E
279		Xanthisthisa terna	Herbulot, 1984	Shoa, Menagesha Forest	E
280		Xanthorhoe abyssinica	Herbulot, 1983	Chensha	E
281		Xanthorhoe alta	Debauche, 1937	Mt Chillálo, Albaso	E
282	Geometridae	Xanthorhoe cadra	(Debauche, 1937)	Mt Chillálo, from forest of Kosso-trees	E
283		Xanthorhoe cuneosignata	Debauche, 1937	Mt Chillálo, Albaso	E
284		Xanthorhoe excelsissima	Herbulot, 1977	Mt Batu	E
285		Xenimpia sabae amarei	Hausmann, 2006	Arba Minch, Region of Omo, Gemu Gofa,	E
286		Xylopteryx emunctaria	(Guenée, 1858)	Abyssinia [Ethiopia]
287		Xylopteryx gada	Herbulot, 2000	Balé, Harena Forest	E
288		Xylopteryx raphaelaria	(Oberthür, 1880)	Scioa [Shoa]	E
289		Zamarada excavata pollex	Fletcher, 1974	Jlubador [Ilubabor] Gore	E
290		Zamarada hyalinaria	(Guenée, 1857)	Abyssinia [Ethiopia]
291		Zamarada melasma	Fletcher, 1974	Dire Daoua [Dire Dawa]
292		Zamarada secutaria	(Guenée, 1857)	Abyssinia [Ethiopia]
293		Zamarada shoa	Herbulot, 2002	Shoa, 50 km W of Addis Ababa	E
294		Zamarada torrida	Fletcher, 1974	Dire Daoua [Dire Dawa]
295	Glyphipterigidae	Ussara semicoronis	Meyrick, 1932	Jem-Jem Forest	E
296	Gracillariidae	Acrocercops heteroloba	Meyrick, 1932	Jem-Jem Forest	E
297		Acrocercops orianassa	Meyrick, 1932	Mt Zukwala/Cuqala
298		Caloptilia macropleura	(Meyrick, 1932)	Jem Jem Forest
299		Metacercops hexactis	(Meyrick, 1932)	Jem-Jem Forest	E
300		Metriochroa carissae	Vári, 1963	Addis Ababa, Little Akaki River	E
301		Metriochroa scotinopa	Vári, 1963	Dabra Zeit [Debre Zeit]	E
302	Gracillariidae	Porphyrosela homotropha	Vári, 1963	Addis Ababa, Little Akaki River	E
303		Stomphastis heringi	Vári, 1963	Near Addis Ababa, Little Akaki River	E
304		Stomphastis horrens	(Meyrick, 1932)	Jem-Jem Forest	E
305	Hesperiidae	Abantis meneliki	Berger, 1979	Harrar
306		Apallaga menageshae	Libert, 2014	Mt Menagesha, NW Addis Abeba
307		Coeliades chalybe immaculata	Carpenter, 1935	Alanga River	E
308		Coeliades menelik menelik	(Ungemach, 1932)	Lilmo, dans la pays de Sayo
309		Eretis mixta	Evans, 1937	Dire Daouna [Dire Dawa]
310		Metisella formosus mittoni	Carcasson, 1961	Mega	E
311		Sarangesa lucidella helena	Evans, 1947	Harar	E
312	Lasiocampidae	Beralade perobliqua monostrigata	Berio, 1940	Adi-Abuna [in Tigray, Ethiopia]	E
313		Bombycopsis abyssinica	Joannou & Krüger, 2009	Addis Abeba	E
314		Mallocampa toulgoeti	Rougeot, 1977	Kébré-Mengist [Kibre Mengist]	E
315		Odontocheilopteryx eothina	Tams, 1931	Djoubdo [Yubdo], Birbir	E
316		Odontocheilopteryx lajonquieri	Rougeot, 1977	near Kébré-Mengist [Kibre Mengist]	E
317		Pallastica hararia	Zolotuhin & Gurkovich, 2009	Harar	E
318		Sena donaldsoni rougeoti	Lajonquière, 1977	Arba Minch	E
319		Sena scotti	(Tams, 1931)	Djem-Djem [Jem-Jem] Forest
320		Stoermeriana abbayensis	(Rougeot, 1984)	Bahar-Dar, marais du Nil Bleu, Abbay	E
321		Stoermeriana chavailloni	(Rougeot, 1984)	Melka-Kontouré	E
322		Stoermeriana das	(Hering, 1928)	Eli	E
323		Stoermeriana laportei	(Rougeot, 1977)	Kébré-Mengist [Kibre Mengist]	E
324		Stoermeriana murinuscolor	(Rougeot, 1984)	Shoa, Menagesha Forest	E
325		Stoermeriana saanayetae	(Rougeot, 1984)	Awassa Lake	E
326		Stoermeriana tamsi	(Rougeot, 1977)	Dinsho Marshes, Balé	E
327		Stoermeriana viettei	(Rougeot, 1977)	Dinsho Marshes	E
328	Limacodidae	Crothaema flava	Berio, 1940	Adi-Abuna [in Tigray, Ethiopia]	E
329		Hamartia johanni	Rougeot, 1977	Kébré-Mengist [Kibre Mengist]	E
330		Hamartia medora moulini	Rougeot, 1977	Kébré-Mengist [Kibre Mengist]	E
331		Jordaniana lactea	(Pagenstecher, 1903)	Ganale
332	Lycaenidae	Anthene amarah	(Guérin-Méneville, 1849)	Dire Dawa
333		Anthene butleri butleri	(Oberthür, 1880)	Mantek; Mahal-Uonz
334		Anthene chojnackii	Libert, 2010	10 km NW of Neghelli	E
335		Anthene confusa	Libert, 2010	Touloudimtou [Tullu Dimtu]
336		Anthene contrastata	(Ungemach, 1932)	Bedelle	E
337		Anthene definita nigrocaudata	(Pagenstecher, 1902)	Ginir	E
338		Anthene dulcis	(Pagenstecher, 1902)	Gambe beim Abasse-See
339		Anthene hodsoni	(Talbot, 1935)	Kibish River
340		Anthene opalina janna	Gabriel, 1949	Fich-Babile Road
341		Anthene opalina opalina	Stempffer, 1946	Callafo[Kalafo], Webi Shebeli, Ogaden
342		Anthene pitmani aethiopana	Libert, 2010	Ghibe River, Addis Abeba-Jimma road
343		Anthene princeps	(Butler, 1876)	Atbara*
344		Anthene saddacus	(Talbot, 1935)	Ethiopia	E
345		Anthene suquala	(Pagenstecher, 1902)	Suquala
346		Axiocerses maureli	Dufrane, 1954	Harrar	E
347		Azanus jesous	(Guérin-Méneville, 1849)	Abyssinie [Ethiopia]
348		Cacyreus ethiopicus	(Tite, 1961)	25 km north of Quiha	E
349		Cacyreus fracta ghimirra	Talbot, 1935	Shoa Ghimirra province	E
350		Chilades elicola	(Strand, 1911)	Eli, Ethiopia
351		Deudorix lorisona baronica	Ungemach, 1932	Baro River	E
352		Deudorix ungemachi	Libert, 2004	Ethiopia	E
353		Eicochrysops antoto	(Strand, 1911)	Umgebung unterhalb Antotos [Entoto]	E
354		Eicochrysops meryamae	Rougeot, 1983	Province de Gondar, environs de Debarek	E
355		Eicochrysops messapus sebagadis	(Guérin-Méneville, 1849)	Abyssinie [Ethiopia]
356		Euchrysops abyssinicus	(Aurivillius, 1922)	Tchafianani; Debasso	E
357		Euchrysops cyclopteris	(Butler, 1876)	Atbara*
358		Euchrysops mauensis Abyssinia [Ethiopia]e	Storace, 1950	Bahrdàr [Bahar Dar] sulle rive meridionali del Lago Tana	E
359		Euchrysops nandensis	(Neave, 1904)	Lake Tana
360		Hypolycaena ogadenensis	Stempffer, 1946	Dagahbur, Ogaden	E
361		Iolaus crawshayi maureli	Dufrane, 1954	Harrar [Harar]
362		Iolaus piaggiae	Oberthür, 1883	Kolla di Giagaguè-Agher	E
363		Lachnocnema abyssinica	Libert, 1996	Dire Daouna [Dawa]
364		Lepidochrysops abyssiniensis abyssiniensis	(Strand, 1911)	Eli	E
365		Lepidochrysops abyssiniensis oculus	(Ungemach, 1932)	Ouama	E
366		Lepidochrysops guichardi	Gabriel, 1949	10 miles West of Addis Ababa	E
367		Lepidochrysops lunulifer	(Ungemach, 1932)	Didessa	E
368		Lepidochrysops negus	(Felder & Felder, 1865)	Africa septentrionali-orientalis: Bogo
369		Lepidochrysops pterou lilacina	(Ungemach, 1932)	Didessa	E
370	Lycaenidae	Lepidochrysops subvariegata	Talbot, 1935	Dirre Dawa	E
371		Leptomyrina boschi	Strand, 1911	Abyssinie [Ethiopia]n	E
372		Lycaena phlaeas pseudophlaeas	(Lucas, 1866)	Abyssinie [Ethiopia]	E
373		Myrina silenus nzoiae	Stoneham, 1937	Western Kenya to Ethiopia and Eritrea
374		Pentila pauli ras	Talbot, 1935	S.W. Abyssinia [Ethiopia], Pokodi [Bokoji]	E
375		Stugeta bowkeri ethiopica	(Stempffer & Bennett, 1958)	Harrar [Harar]	E
376		Tarucus ungemachi	Stempffer, 1942	Rivière Baro Abyssinie [Ethiopia] occidentale
377		Thermoniphas colorata	(Ungemach, 1932)	Youbdo
378		Tuxentius cretosus	(Butler, 1876)	Atbara*
379		Tuxentius kaffana	(Talbot, 1935)	Nado’s Province, Yeki; Mocha District, Gamadura	E
380		Uranothauma antinorii	(Oberthür, 1883)	Torrente di Sciotalit
381		Uranothauma nubifer distinctesignatus	(Strand, 1911)	[Ethiopia]	E
382		Zintha hintza resplendens	(Butler, 1876)	Atbara*	E
383	Metarbelidae	Aethiopina semicirculata	Gaede, 1929	Abyssinia [Ethiopia]	E
384		Salagena fetlaworkae	Rougeot, 1977	near Koffolé [Koffale]	E
385		Teragra lemairei	Rougeot, 1977	Dinsho Marches	E
386		Teragra villiersi	Rougeot, 1977	near Koffolé [Koffale]	E
387	Noctuidae	Abrostola obliqua	Dufay, 1958	Abyssinia [Ethiopia]	E
388		Abrostola rougeoti	Rougeot, 1977	near Koffolé [Koffale]	E
389		Acontia albatrigona	Hacker, Legrain & Fibiger, 2008	Arba Minch Region, Omo, Province Gemu, Gofa
390		Acontia amarei	Hacker, Legrain & Fibiger, 2010	Gamu-Gofa Province, 10.5 km W of Weyto	E
391		Acontia amhara	Hacker, Legrain & Fibiger, 2008	Gamu-Gofa Province, 8 km N of Turmi	E
392		Acontia proesei	Hacker, Legrain & Fibiger, 2008	Valley of the river Tekezé, 30 km N of Gashena	E
393		Acontia robertbecki	Hacker, Legrain & Fibiger, 2010	Arba Minch Region, Gemu Gofa Province	E
394		Acontia ruficincta	Hampson, 1910	Atbara*	E
395		Acontia secta	Guenée, 1852	Abyssinia [Ethiopia]
396		Acontia uhlenhuthi	Hacker, Legrain & Fibiger, 2008	Diré Daouá [Dire Dawa]	E
397		Acontiola boursini	(Berio, 1940)	Lekemti [Naqamte]
398		Acrapex abbayei	Laporte, 1984	Dinsho Reserve	E
399		Acrapex apexangula	Laporte, 1984	near Koffolé [Koffale]	E
400		Acrapex ausseili	Laporte, 1984	Fisha Genet	E
401		Acrapex franeyae	Laporte, 1984	Dinsho Reserve	E
402		Acrapex genrei	Laporte, 1984	Dinsho Reserve	E
403		Acrapex girardi	Laporte, 1984	Dinsho Reserve	E
404		Acrapex guiffrayorum	Laporte, 1984	Dinsho Reserve	E
405		Acrapex mastawatae	Laporte, 1984	Arba Minch	E
406		Acrapex matilei	Laporte, 1984	Dinsho Reserve	E
407		Acrapex satanas	Laporte, 1984	Dinsho Reserve	E
408		Acrapex soyema	Le Ru, 2017	Gibe, Soyema Bridge	E
409		Acrapex ulmii	Laporte, 1991	Koffole [Koffale]	E
410		Acrapex zaouditou	Laporte, 1991	Koffole [Koffale]	E
411		Aedia albirena	(Hampson, 1926)	Taddecha Mullha
412	Noctuidae	Aedia konsonata	Hacker, 2016	Konso	E
413		Aedia marmoreata	Hacker, 2016	12 km W of Jinka
414		Aegocera ferrugo	Jordan, 1926	Hora Daka	E
415		Agrotis baleense	Laporte, 1977	Dinsho, Bale Reserve
416		Agrotis cinchonina	Guenée, 1852	Abyssinia [Ethiopia]	E
417		Agrotis debivari	(Berio, 1962)	Africa Orientale Italiana, Debivar	E
418		Agrotis separata	Guenée, 1852	Abyssinia [Ethiopia]	E
419		Amazonides berioi	(Laporte, 1984)	Lekemti [Naqamte]	E
420		Amazonides berliozi	Laporte, 1974	Dinsho Col	E
421		Amazonides dubiomeodes	Laporte, 1977	Kébré-Mengist [Kibre Mengist]	E
422		Amazonides ezanai	(Laporte, 1984)	Kébré-Mengist [Kibre Mengist]	E
423		Amazonides fumigera	Laporte, 1977	Dinsho Marshes	E
424		Amazonides koffoleense	Laporte, 1977	Koffolé [Koffale]	E
425		Amazonides laheuderiae	Laporte, 1984	Abba Hoye-Gara	E
426		Amazonides pseudoberliozi	Rougeot & Laporte, 1983	Simyen, Sankaber	E
427		Amazonides putrefacta	(Guenée, 1852)	Abyssinia [Ethiopia]
428		Amazonides ungemachi	(Laporte, 1984)	Ioubdo, Birbir, Nole Kabe	E
429		Amazonides zarajakobi	Laporte, 1984	Dinsho Marshes	E
430		Amphia hepialoides	Guenée, 1852	Abyssinia [Ethiopia]antio	E
431		Aporophoba subaustralis	Berio, 1977	Addis Ababa	E
432		Apospasta albirenalis	Laporte, 1974	Mt Batu	E
433		Apospasta diffusa	Laporte, 1974	Dinsho Col	E
434		Apospasta erici	Laporte, 1984	Dinsho Reserve	E
435		Apospasta incongrua	Laporte, 1974	Dinsho Col	E
436		Apospasta maryamae	Laporte, 1974	Dinsho Marshes	E
437		Apospasta niger	Laporte, 1974	Dinsho Marshes	E
438		Apospasta rougeoti	Laporte, 1991	Boré Forest
439		Apospasta rufa	Laporte, 1991	Choa [Shoa], Menageshah [Menegasha] Forest	E
440		Apospasta sabulosa	Fletcher, 1959	Simien, Lori	E
441		Apospasta thomasi	Laporte, 1991	Addis Ababa	E
442		Ariathisa abyssinia	(Guenée, 1852)	Abyssinia [Ethiopia]
443		Aspidifrontia ungemachi	(Laporte, 1978)	Metti	E
444		Athetis aeschrioides	Berio, 1940	Adi-Abuni [in Tigray, Ethiopia]
445		Athetis carayoni	Laporte, 1977	Dinsho col	E
446		Athetis viettei	Laporte, 1991	Choa [Shoa], Melka-Kontoure [Melka Konture]	E
447		Axylia aregashae	Laporte, 1984	near Kébré-Mengist [Kibre Mengist]	E
448		Axylia bryi	Laporte, 1984	Dinsho Marshes	E
449		Axylia destefanii	Berio, 1944	El-Dire
450		Axylia gabriellae	Laporte, 1975	Boré Forest	E
451		Axylia marthae	Laporte, 1984	near Koffolé [Koffale]	E
452		Axylia orbicularis	Laporte, 1984	near Kébré-Mengist [Kibre Mengist]	E
453		Axylia sanyetiensis	Laporte, 1984	near Mt Batu	E
454		Axylia vespertina	Laporte, 1984	near Kébré-Mengist [Kibre Mengist]	E
455		Batuana abbahoyegarana	Rougeot, 1983	Abba Hoye-Gara, Wollo	E
456		Batuana exspectata	Laporte & Rougeot, 1981	Gojam, Mt Choke	E
457		Batuana lobeliarum	Laporte, 1976	near Dinsho	E
458		Batuana rougeoti	Laporte, 1976	near Mt Batu	E
459	Noctuidae	Berionycta beckroberti	Kiss, 2017	15 km E of Yabello	E
460		Berionycta behouneki	Kiss, 2017	13 km W of Yabello	E
461		Berionycta berioi	Kiss, 2017	12 km NNE of Arba Minch	E
462		Berionycta nigra	Kiss, 2017	15 km E of Yabello	E
463		Berionycta orbicularis	Kiss, 2017	15 km E of Yabello	E
464		Berionycta ponticamima	Kiss, 2017	15 km E of Yabello	E
465		Capillamentum gelleyi	Laporte, 1984	Addis Ababa	E
466		Caradrina atriluna	Guenée, 1852	Abyssinia [Ethiopia]
467		Caradrina torpens	Guenée, 1852	Abyssinia [Ethiopia]	E
468		Carcharoda erlangeri	Rothschild, 1924	Waute Merehan [Mreham]	E
469		Cirrodes rosaceus	Rothschild, 1924	Waute Merehan [Mreham]	E
470		Claudaxylia dinshoense	Laporte, 1984	Dinsho Reserve	E
471		Compsotata corneliae	Behounek & Beck, 2012	Bale Mountains, Province of Bale, Hangasso	E
472		Conservula ludocaroli	(Laporte, 1991)	Debre Zeit	E
473		Conservula scriptura	(Rougeot & Laporte, 1983)	Simyen, Sankaber	E
474		Cucullia simoneaui	Laporte, 1976	Bale Reserve, Dinsho	E
475		Cucullia tedjicolora	Laporte, 1977	Kébré-Mengist [Kibre Mengist]	E
476		Eucladodes achrorophilus	Laporte, 1976	Near Mt Batu	E
477		Eucladodes baleensis	Laporte, 1976	Bale Reserve	E
478		Euplexia imperator	Laporte, 1984	Dinsho Marshes	E
479		Euplexia mercieri	Laporte, 1984	Arussi, near Koffolé [Koffale]	E
480		Euplexia pinoni	Laporte, 1984	Kébré-Mengist [Kibre Mengist]	E
481		Euplexia shoana	Laporte, 1984	Shoa, near Hosana	E
482		Euxoa dodolaense	Laporte, 1984	Road to Dodola	E
483		Euxoa montigenarum	Rougeot & Laporte, 1983	Simyen, Sankaber	E
484		Euxoa semyenensis	Laporte, 1991	Sankaber	E
485		Euxoa waliarum	Rougeot & Laporte, 1983	Simyen, Sankaber	E
486		Feliniopsis duponti	Laporte, 1974	near Kebré-Mengist [Kibre Mengist]
487		Feliniopsis germainae	Laporte, 1975	near Kebré-Mengist [Kibre Mengist]	E
488		Feliniopsis insolita	Hacker & Fibiger, 2007	Addis Ababa, Sholla	E
489		Feliniopsis jinka	Hacker, 2010	Gamu-Gofa Province, 10 km W of Jinka
490		Feraxinia jemjemensis	(Laporte, 1984)	Kébré-Mengist [Kibre Mengist]	E
491		Heliophobus africana	Berio, 1977	Addis Ababa	E
492		Heliothis saskai	(Berio, 1975)	Addis Ababa	E
493		Hemituerta mahdi	(Pagenstecher, 1903)	Hanadscho [Dinsho district]
494		Heraclia viettei	Kiriakoff, 1973	Nole Kaba	E
495		Hermonassoides abyssinica	(Berio, 1975)	Addis Ababa	E
496		Hermonassoides dinshoensis	(Laporte, 1977)	Dinsho Marshes	E
497		Hermonassoides marmorata	(Laporte, 1977)	Fisha Genet	E
498		Hermonassoides mauricei	(Laporte, 1975)	Koffale
499		Hermonassoides mendeboense	(Laporte, 1984)	Dinsho	E
500		Hermonassoides minosi	(Laporte, 1991)	Managesha Forest	E
501		Hermonassoides scipioni	(Laporte, 1977)	Dinsho, Bale Reserve	E
502		Hiccoda clarae	Berio, 1947	Ogaden, Uarder [Warder]	E
503		Hyperfrontia direae	Berio, 1962	Dire-Daoua [Dire Dawa]	E
504		Hyperfrontia limbata	Berio, 1962	El-Dire	E
505		Koffoleania michaellae	Laporte, 1977	near Koffolé [Koffale]	E
506		Leucania aedesiusi	Rougeot & Laporte, 1983	Simyen, Sankaber	E
507		Leucania argyrina	Laporte, 1984	Bahar Dar	E
508	Noctuidae	Leucania claudicans	Guenée, 1852	Abyssinia [Ethiopia]	E
509		Leucania cyprium	(Laporte, 1984)	Dinsho Marshes	E
510		Leucania fasilidasi	(Laporte, 1984)	Dinsho Marshes	E
511		Leumicamia oreias	(Fletcher, 1959)	Simien, above Lori	E
512		Leumicamia palustris	Laporte, 1976	Dinsho Marshes	E
513		Leumicamia venustissima	(Laporte, 1974)	Bale Reserve	E
514		Lophotarsia girmai	Laporte, 1975	Arba Minch	E
515		Lophotarsia leucoplagoides	(Berio, 1941)	El-Dire	E
516		Lophotarsia theresae	Beck & Behounek, 2013	Bale Mountains National Park, region Oromia/Sidamo, Province of Bale, 4 km W of Sura	E
517		Maghadena ingridae	Laporte, 1977	Dinsho Reserve, Balé	E
518		Maliattha eburnea	Hacker, 2016	Oroma Province, 6 km ESE of Jimma
519		Matopo berhanoui	Laporte, 1984	Melka-Kontouré [Konture]	E
520		Mentaxya bruneli	Laporte, 1975	near Kebré-Mengist [Kibre Mengist]	E
521		Mentaxya fouqueae	Laporte, 1974	Boré Forest
522		Mentaxya inconstans	Laporte, 1984	Dinsho Marshes	E
523		Mentaxya lacteifrons	Laporte, 1984	Kébré-Mengist [Kibre Mengist]	E
524		Michelliana afroalpina	Laporte, 1976	near Mt Batu	E
525		Micraxylia antemedialis	Laporte, 1975	near Kebré-Mengist [Kibre Mengist]	E
526		Micraxylia hypericoides	Berio, 1962	Oromo e Sidamo, Neghelli [Neghelle]	E
527		Micraxylia lividoradiata	(Berio, 1940)	Adi-Abuna [in Tigray, Ethiopia]	E
528		Mythimna altiphila	Hreblay & Legrain, 1996	Addis Abeba [Ababa]
529		Mythimna amlaki	Laporte, 1984	Near Mt Batu	E
530		Mythimna bisetulata	(Berio, 1940)	Adi-Abuna [in Tigray, Ethiopia]	E
531		Mythimna germanae	Laporte, 1991	Melka Kontouré	E
532		Neostichtis teruworkae	Laporte, 1984	Near Hosana	E
533		Nocthadena griseoviridis	Laporte, 1976	Near Mt Batu	E
534		Numeniastes selenis	Fletcher, 1963	Harar
535		Nyodes biardi	Laporte, 1984	Shashemane	E
536		Ochropleura sidamona	Laporte, 1977	Fisha-Genet
537		Odontestra richinii	Berio, 1940	Adi-Abuna [in Tigray, Ethiopia]	E
538		Odontestra variegata	Berio, 1940	Adi-Abuna [in Tigray, Ethiopia]
539		Odontestra vitta	Berio, 1975	Addis Ababa	E
540		Oligia adactricula	Guenée, 1852	Abyssinie [Ethiopia]	E
541		Oligia arbaminchensis	Laporte, 1991	Arba Minch	E
542		Oligia genettae	Laporte, 1991	Kebre-Mengist [Kibre Mengist]	E
543		Omphalestra nellyae	(Berio, 1939)	Adua [in Ethiopia]	E
544		Ozarba alberti phaeoxantha	Hacker, 2016	Dire Dawa
545		Ozarba didymochra	Hacker, 2016	Gamu-Gofa Province, 8 km E of Weyto	E
546		Ozarba fuscundosa	Hacker, 2016	Oromia, 3 km NNE of Finchawa
547		Ozarba grisescens	Berio, 1947	Harrar [Harar], Dire Daua [Dire Dawa]
548		Ozarba latizonata	Hacker, 2016	Gamu-Gofa Province, 8 km E of Weyto
549		Ozarba naumanni	Hacker, 2016	Gamo Gofa Province, Konso	E
550	Noctuidae	Ozarba permutata	Hacker, 2016	20 km ESE of Sashemene, Wondo Genet
551		Ozarba rubrofusca	Berio, 1947	Ogaden, Uarder [Werder]	E
552		Ozarba tenuis	Hacker, 2016	Province of Gamo Gofa, 8 km N of Turmi	E
553		Ozarba uhlenhuthi	Hacker, 2016	Dire Dawa	E
554		Phyllophila corgatha	Berio, 1984	Arba Minch	E
555		Phyllophila richinii	Berio, 1940	Adi-Abuna [in Tigray, Ethiopia]	E
556		Pseudozarba nilotica	Hacker, 2016	30 km SE of Bahir Dar, Tisisat above Blue Nile Falls	E
557		Pusillathetis fiorii	Berio, 1976	Uarder [Warder in Ogaden]
558		Ramesodes oblonga	Berio, 1976	Adi Abuna [in Tigray, Ethiopia]
559		Rhodochlaena dinshoense	Laporte, 1974	Dinsho Marshes	E
560		Rougeotia abyssinica	(Hampson, 1918)	Kutai Mecha	E
561		Rougeotia aethiopica	Laporte, 1974	Dinsho swamp	E
562		Rougeotia ludovici	Laporte, 1974	Bale Reserve	E
563		Rougeotia ludovicoides	Laporte, 1977	Dinsho Marshes	E
564		Rougeotia obscura	Laporte, 1974	Dinsho Col	E
565		Rougeotia roseogrisea	Laporte, 1974	Near Mt Batu	E
566		Rougeotia rougeoti	Laporte, 1984	Mt Batu Forest	E
567		Schinia ennatae	(Laporte, 1984)	Addis Ababa	E
568		Schinia magdalenae	(Laporte, 1976)	Bale Reserve, Dinsho	E
569		Schinia ungemachi	(Berio, 1945)	Oromo Sidamo, Uollega [Wollega]	E
570		Schinia xanthiata	(Berio, 1940)	Adi-Abuna [in Tigray, Ethiopia]	E
571		Sciomesa boulardi	(Laporte, 1984)	near Koffolé [Koffale]	E
572		Sciomesa excelsa	(Laporte, 1976)	Near Mt Batu	E
573		Sciomesa franciscae	Laporte, 1991	Choa, Hosana	E
574		Sciomesa secata	Berio, 1977	Addis Ababa	E
575		Sesamia enanouae	Laporte, 1991	Gojam, Bahr-Dar, marais du Nil Bleu	E
576		Sesamia roumeti	Laporte, 1991	Gojam, Bahr-Dar	E
577		Solgaitiana petrosi	Laporte, 1984	Kébré-Mengist [Kibre Mengist]	E
578		Spodoptera excelsa	Rougeot & Laporte, 1983	Simyen, Sankaber	E
579		Subnoctua arbaminchensis	Laporte, 1984	Arba Minch	E
580		Thiacidas robertbecki	Hacker & Zilli, 2007	Awassa, Awassa Lake, Bale Region	E
581		Tholeropsis decimata	Berio, 1977	Addis Ababa	E
582		Tholeropsis uncinata	Berio, 1977	Addis Ababa	E
583		Thysanoplusia asapheia	(Dufay, 1977)	near Koffolé [Koffale]
584		Thysanoplusia dolera	Dufay, 1977	near Koffolé [Koffale]	E
585		Timora flavocarnea	Hampson, 1903	Abyssinia [Ethiopia]
586		Timora zavattarii	Berio, 1944	El-Dire	E
587		Tracheplexia annabellae	Laporte, 1991	Menagesha Forest	E
588		Tracheplexia colettae	Laporte, 1991	Gemu-Gofa, Arba-Minch	E
589		Tracheplexia leguerni	Laporte, 1984	Fort Wosha	E
590		Tracheplexia petryvesi	Laporte, 1991	Menagesha Forest	E
591		Tracheplexia richinii	Berio, 1973	Adiu Abuna [in Tigray, Ethiopia]
592		Tycomarptes adami	Laporte, 1974	Dinsho Col	E
593		Tycomarptes aethiopica	Laporte, 1974	Mt Batu	E
594		Tycomarptes berioi	Laporte, 1974	Boré Forest	E
595		Tycomarptes bipuncta	Laporte, 1974	Boré Forest	E
596		Tycomarptes bipunctatoides	Laporte, 1974	near Koffolé [Koffale]	E
597	Noctuidae	Tycomarptes gelladarum	Rougeot & Laporte, 1983	Simyen, Sankaber	E
598		Tycomarptes inferior	(Guenée, 1852)	Abyssinia [Ethiopia]
599		Tycomarptes journiaci	Laporte, 1977	Near Mt Batu	E
600		Tycomarptes limoni	Laporte, 1974	near Koffolé [Koffale]	E
601		Tycomarptes semyensis	Rougeot & Laporte, 1983	Simyen, Sankaber	E
602		Tycomarptes thibauti	Laporte, 1974	Boré Forest	E
603		Vietteania chojnackii	(Laporte, 1984)	Dinsho Marshes	E
604	Nolidae	Arcyophora zanderi	Felder & Rogenhofer, 1875	Abyssinia [Ethiopia]
605		Bryophilopsis martinae	Laporte, 1991	Gemu-Gofa, Konso	E
606		Characoma adiabunensis	Berio, 1940	Adi-Abuna [in Tigray, Ethiopia]	E
607		Earias richinii	Berio, 1940	Adi-Abuni [in Tigray, Ethiopia]
608		Eligma neumanni	Rothschild, 1924	Blue Nile, Abera Koritscha, Uata Dera	E
609		Escarpamenta damarana abyssinica	Hacker, 2013	6 km E of Weyto, Weyto River	E
610		Evonima littoralis abyssinica	Hacker, 2012	Southern Province, Jinka, Mago National Park, 350 m SW of Headquarter,	E
611		Gigantoceras villiersi	Laporte, 1975	Arba Minch	E
612		Meganola cerographa	Hacker, 2012	Oromia District, 6.5 km N of Bonga	E
613		Meganola coffeana	Hacker, 2012	Oromia Province, 6.5 km NE of Shebe
614		Meganola ethiopica	Hacker, 2012	Addis Ababa	E
615		Meganola harenna	Hacker, 2014	Harenna Forest, Karcha Camp Ground	E
616		Meganola leucometabola	Hacker, 2012	Oromia Province, 6.5 km N of Bonga
617		Meganola longisigna	Hacker, 2012	Oromia Region, 1km W. of village Aluweya
618		Meganola lupii	Hacker & Hausmann, 2012	Oromia Province, 13 km S. of Agere Maryam	E
619		Meganola pachygrapha	Hacker, 2012	Oromia Province, 6.5 km N of Bonga
620		Meganola poliovittata	Hacker, 2012	Oromia Province, 6 km ESE of Jimma	E
621		Meganola pyrrhomorpha	Hacker, 2012	Oromia Province, 6.5 km N of Bonga	E
622		Meganola simillima	Hacker, 2012	Oromia District, 13 km S of Agere Maryam	E
623		Meganola stadiensis	Hacker, 2014	Harenna Forest, Karcha Camp Ground	E
624		Meganola stigmatolalis	Hacker, 2012	Southern Province, 23 km WSW of Welkite, Gibe River
625		Meganola unilineata	Hacker, 2012	Southern Province, 11.2 km W of Bonga	E
626		Neaxestis mesogonia	Hampson, 1905	Atbara R.
627		Nola abyssinica	Hacker, 2012	Oromia Province, 13 km S of Agere Maryam
628		Nola afrotaeniata	Hacker, 2012	12 km W Jinka, border Mago National Park
629		Nola amhara	Hacker, 2012	Addis Ababa
630		Nola angensteini	Hacker, 2012	Afar Region, NE of Mile Serdo Wildlife Refuge, Tendaho
631	Nolidae	Nola balealpina	Hacker, 2012	Oromia Province, Bale Mountains National Park, Disho	E
632		Nola calochromata	Hacker, 2014	Harenna Forest, Harenna Forest Road	E
633		Nola destituta	Hacker, 2012	Oromia Province, 8 km W of Nazret	E
634		Nola jarzabekae	Hacker, 2012	Oromia Province, Abiyata-Shala-Hayak National Park	E
635		Nola omphalota euroetes	Hacker, 2012	Oromia Province, 6 km ESE of Jimma
636		Nola socotrensis vansoni	Hacker, 2012	12 km W Jinka, border Mago NP
637		Nola sphaeromorpha	Hacker, 2012	Oromia Province, 13 km S of Agere Maryam,	E
638		Nolidia platygrapha	Hacker, 2012	Amhara Region, W of Mirab, Gojam Zone, 15 km NW of Bahar Dar	E
639	Notodontidae	Afroplitis quadratus	(Viette, 1954)	River Baro	E
640		Antheua birbirana	Viette, 1954	middle course of Birbir, Youbdo	E
641		Antheua gaedei	Kiriakoff, 1962	Addis Ababa	E
642		Antheua trivitta	(Hampson, 1910)	Abyssinia [Ethiopia]	E
643		Antistaura decorata	Kiriakoff, 1965	Derdaua, North-East of Harrar	E
644		Boscawenia nora	(Pagenstecher, 1903)	Ganale	E
645		Desmeocraera kiriakoffi	Thiaucourt, 1977	near Kébré-Mengist [Kibre Mengist]	E
646		Eutimia smithii	Holland, 1897	Dombalok	E
647		Polelassothys callista abyssinica	Viette, 1954	Moy. Dedissa [Didessa]	E
648		Psalisodes saalfeldi	Kiriakoff, 1979	Al Abed	E
649		Scalmicauda azebae	Thiaucourt, 1977	near Kébré-Mengist [Kibre Mengist]	E
650		Thaumetopoea apologetica abyssinica	Strand, 1911	Addis Ababa
651		Tricholoba rougeoti	Thiaucourt, 1977	Arba Minch	E
652	Nymphalidae	Acraea aganice orientalis	(Ungemach, 1932)	Bouré
653		Acraea alcinoe nado	(Ungemach, 1932)	Bouré	E
654		Acraea chilo chilo	Godman, 1880	Kalamet, Sebka Valley
655		Acraea doubledayi	Guérin-Méneville, 1849	Abyssinie [Ethiopia]
656		Acraea epaea homochroa	(Rothschild & Jordan, 1905)	Banka, Malo	E
657		Acraea kakana	Eltringham, 1911	Adie Kaka, Kafa	E
658		Acraea oscari	Rothschild, 1902	Banka, Malo	E
659		Acraea poggei ras	(Ungemach, 1932)	Oullaga [Wollega]	E
660		Acraea zetes sidamona	Rothschild & Jordan, 1905	Alata, Sidamo	E
661		Acraea zoumi	Pierre, 1995	Ethiopia	E
662		Amauris echeria steckeri	Kheil, 1890	Abessynia
663		Amauris hecate stictica	Rothschild & Jordan, 1903	Anderatscha	E
664		Amauris niavius aethiops	Rothschild & Jordan, 1903	Anderatscha
665		Amauris ochlea darius	Rothschild & Jordan, 1903	Anderatscha
666		Antanartia abyssinica	(C. & R. Felder, [1867])	Ethiopia	E
667		Antanartia schaeneia diluta	Rothschild & Jordan, 1903	Kaffa	E
668		Argynnis hyperbius neumanni	Rothschild, 1902	Kaffa	E
669		Aterica galene incisa	Rothschild & Jordan, 1903	between Kankati and Djibbe, Djimma [Jimma]	E
670	Nymphalidae	Bicyclus pavonis	(Butler, 1876)	Abyssinia [Ethiopia]
671		Bicyclus safitza aethiops	(Rothschild & Jordan, 1905)	Lake Abassi	E
672		Charaxes etesipe abyssinicus	Rothschild, 1900	Sciotalit, Sxioa [Shoa]	E
673		Charaxes eurinome birbirica	(Ungemach, 1932)	Youbdo
674		Charaxes figini	van Someren, 1969	Eritaea, Setit, El Eghin [Ethiopia]
675		Charaxes galawadiwosi	Plantrou & Rougeot, 1979	Arba-Minch	E
676		Charaxes hansali hansali	Van Someren, 1971	Africa septentrionali-orientalis: Bogos
677		Charaxes jahlusa ganalensis	Carpenter, 1937	Salakle, Ganale river”
678		Charaxes junius junius	Oberthür, 1883	Scioa [Shoa]	E
679		Charaxes junius somalicus	Rothschild, 1900	Harrar Highlands, Somaliland
680		Charaxes kirki daria	Rothschild, 1903	Jabalo	E
681		Charaxes lactetinctus ungemachi	Le Cerf, 1927	Youbdo (Birir)
682		Charaxes larseni	Rydon, 1982	Jambo area, Nanji Hill	E
683		Charaxes numenes neumanni	Rothschild, 1902	Wori to Gamitscha, Kaffa	E
684		Charaxes pelias pagenstecheri	Poulton, 1926	S Ethiopia
685		Charaxes phoebus	Butler, 1866	Abyssinia [Ethiopia]	E
686		Charaxes rectans	Rothschild & Jordan, 1903	Upper Urga, Kollu, Schoa [Shoa]
687		Charaxes saturnus pagenstecheri	Poulton, 1926	S. Abyssinia [Ethiopia]	E
688		Charaxes sidamo	Plantrou & Rougeot, 1979	Kébré-Mengist [Kibre Mengist]	E
689		Charaxes tiridates marginatus	Rothschild & Jordan, 1903	Scheko	E
690		Eronia cleodora cleodora	Hübner, [1823]	Ethiopia
691		Eronia leda	(Boisduval, 1847)	Marako
692		Euphaedra caerulescens submarginalis	Hecq, 1997	[Ethiopia?]	E
693		Euphaedra castanoides deficiens	Hecq, 1997	West, Didessa River	E
694		Euphaedra medon abouna	Ungemach, 1932	Youbdo	E
695		Euphaedra neumanni	Rothschild, 1902	Scheko [Sheko]
696		Euphaedra sarita abyssinica	Rothschild, 1902	Kankati forest, Djimma	E
697		Eurytela hiarbas abyssinica	Rothschild & Jordan, 1903	Banka	E
698		Euxanthe eurinome birbirica	Ungemach, 1932	Youbdo
699		Hypolimnas salmacis platydema	Rothschild & Jordan, 1903	Scheko	E
700		Junonia terea fumata	(Rothschild & Jordan, 1903)	Gillet Mountains
701		Lasiommata maderakal	(Guérin-Méneville, 1849)	Abyssinie [Ethiopia]
702		Melitaea abyssinica	Oberthür, 1909	Abyssinie [Ethiopia]	E
703		Neptis nemetes obtusa	Rothschild & Jordan, 1903	Scheko	E
704		Phalanta eurytis microps	(Rothschild & Jordan, 1903)	Walenso [Woliso], Gillet Mts	E
705		Phalanta phalantha aethiopica	(Rothschild & Jordan, 1903)	Gillet Mts
706		Pseudacraea boisduvalii sayonis	Ungemach, 1932	Oumbi	E
707		Pseudacraea eurytus mimoras	Ungemach, 1932	Oumbi	E
708		Pseudacraea lucretia walensensis	(Sharpe, 1896)	Waenso [Woliso]	E
709		Sevenia boisduvali kaffana	(Rothschild & Jordan, 1903)	Godjeb to Bonga, Kaffa	E
710	Nymphalidae	Telchinia aurivillii schecana	Rothschild & Jordan, 1905	Scheko [Sheiko]	E
711		Telchinia bonasia banka	Eltringham, 1912	Banka, Malo
712		Telchinia guichardi	Gabriel, 1949	Lekempti	E
713		Telchinia jodutta aethiops	Rothschild & Jordan, 1905	Dereta Mts	E
714		Telchinia necoda	Hewitson, 1861	Abyssinia [Ethiopia]	E
715		Telchinia peneleos gelonica	(Rothschild & Jordan, 1905)	Upper Gelo River	E
716		Telchinia perenna kaffana	(Rothschild, 1902)	Kaffa	E
717		Telchinia pharsalus rhodina	Rothschild, 1902	Kaffa	E
718		Telchinia rangatana maji	Carpenter, 1935	Maji Province	E
719		Telchinia safie antinorii	(Oberthür, 1880)	Mahal-Uonz	E
720		Telchinia safie safie	(C. & R. Felder, 1865)	Abyssinia [Ethiopia] Meridionalis	E
721		Telchinia ungemachi	(Le Cerf, 1927)	Youbdo (Birbi)	E
722		Tirumala formosa neumanni	(Rothschild & Jordan, 1903)	Kaffa	E
723		Vanessa abyssinica abyssinica	Vane-Wright & Hughes, 2007	Ethiopia
724		Ypthima impura paupera	Ungemach, 1932	Soubé-Boro
725		Ypthima simplicia	Butler, 1876	Atbara*
726	Papilionidae	Graphium almansor birbiri	(Ungemach, 1932)	Baro	E
727		Graphium angolanus baronis	(Ungemach, 1932)	Baro
728		Papilio arnoldiana	Vane-Wright, 1995	S.W. Abyssinia [Ethiopia], Grine	E
729		Papilio dardanus antinorii	Oberthür, 1883	Abissinia, Feleklek and Sciotalit	E
730		Papilio echerioides leucospilus	Rothschild, 1902	Gara Mulata near Harar”	E
731		Papilio echerioides oscari	Rothschild, 1902	Kaffa and Djima [Jimma]	E
732		Papilio microps	Storace, 1951	Shoa, Abyssinia [Ethiopia] centrale
733		Papilio nireus pseudonireus	Felder & Felder, 1865	Africa Septentrionali Oriental, Bogos
734		Papilio rex abyssinicana	Vane-wright, 1995	S. W. Abyssinia [Ethiopia], Ganji River	E
735		Papilio wilsoni	Rothschild, 1926	Nubar Hills, Taldi	E
736	Pieridae	Appias sylvia abyssinica	Talbot, 1932	Joubda (Birbir)	E
737		Belenois gidica abyssinica	(Lucas, 1852)	Abyssinie [Ethiopia]	E
738		Belenois gidica hypoxantha	(Ungemach, 1932)	Gambela	E
739		Belenois raffrayi	(Oberthür, 1878)	Lac de Tzana [Lake Tana]
740		Belenois subeida hailo	(Ungemach, 1932)	Nolé Kaba [in Wollega]	E
741		Belenois thysa tricolor	Talbot, 1943	Abyssinia [Ethiopia]
742		Belenois zochalia gada	(Ungemach, 1932)	Nole-Kaba [in Wollega]	E
743		Colias electo meneliki	Berger, 1940	Gondar
744		Colias erate marnoana	Rogenhofer, 1884	Ethiopia
745		Colotis antevippe zera	(Lucas, 1852)	Abyssinie [Ethiopia]
746		Colotis celimene celimene	(Lucas, 1852)	Abyssinie [Ethiopia]
747		Colotis danae eupompe	(Klug, 1829)	in Arabia deserta, in Sinai monte, in Dongala et Habessinia
748		Colotis euippe exole	(Reiche, 1850)	Abyssinie [Ethiopia]
749		Colotis hetaera aspasia	(Ungemach, 1932)	Baro
750		Colotis phisadia ocellatus	(Butler, 1886)	Somali-land [False locality]	E
751		Colotis ungemachi	(Le Cerf, 1922)	N Ethiopia	E
752		Dixeia charina septentrionalis	(Bernardi, 1958)	Djemdjem	E
753	Pieridae	Eronia leda pupillata	Strand, 1911	Marako	E
754		Euchloe belemia abyssinica	Riley, 1928	Mt. Chillalo	E
755		Eurema desjardinsii regularis	(Butler, 1876)	Atbara*
756		Leptosia alcesta pseudonuptilla	Bernardi, 1959	Haute-Orguessa
757		Mylothris erlangeri	Pagenstecher, 1902	Gewidscha	E
758		Mylothris mortoni balkis	Ungemach, 1932	Alenga	E
759		Mylothris mortoni mortoni	Blachier, 1912	Kaffa, dans l’Abyssinie [Ethiopia] meridionale”	E
760		Mylothris rueppellii	(Koch, 1865)	Abessynica
761		Mylothris sagala swaynei	Butler, 1899	Harar Highlands	E
762		Mylothris yulei amhara	Ungemach, 1932	Alenga	E
763		Pieris brassicoides	Guérin-Méneville, 1849	Abyssinie [Ethiopia]
764		Pontia daplidice aethiops	(De Joannis & Verity, 1913)	Abyssinie [Ethiopia]	E
765	Plutellidae	Lepocnemis metapelista	Meyrick, 1932	Jem-Jem Forest	E
766		Plutella dryoxyla	Meyrick, 1932	Mt Chillálo	E
767		Plutella oxylopha	Meyrick, 1932	Mt Chillálo	E
768		Plutella stichocentra	Meyrick, 1932	Mt Chillálo	E
769	Psychidae	Acanthopsyche chrysora	Bourgogne, 1980	Arba Minch	E
770		Oiketicoides aethiopica	Bourgogne, 1991	Wollo, Lalibela	E
771		Taleporia aethiopica	Strand, 1911	Mahenge	E
772	Pterophoridae	Cosmoclostis gorbunovi	Ustjuzhanin & Kovtunovich, 2011	West Shewa, 2 km S of Ambo	E
773		Hellinsia aethiopicus	(Amsel, 1963)	Gembi
774		Hellinsia ambo	Ustjuzhanin & Kovtunovich, 2011	West Shewa, 2 km S of Ambo
775		Hellinsia bigoti	(Rougeot, 1983)	Simyen, Sankaber	E
776		Hellinsia negus	(Gibeaux, 1994)	Wondo-Genet	E
777		Merrifieldia lonnvei	Gielis, 2011	Amhara Region, S of Debub, Gondar zone, 8 km NW of Addis Zemen, Highway 3	E
778		Paracapperia esuriens	Meyrick, 1932	Jem Jem Forest
779		Platyptilia daemonica	Meyrick, 1932	Jem Jem Forest	E
780		Platyptilia gondarensis	Gibeaux, 1994	Gondar Province
781		Platyptilia implacata	Meyrick, 1932	Jem Jem Forest	E
782		Pterophorus lindneri	(Amsel, 1963)	Gore	E
783		Stenoptilia aethiopica	Gibeaux, 1994	Sidamo, Wondo-Genet
784		Stenoptilia amharae	Gielis, 2011	Amhara Region, Semien North, Gondar zone, 17 km NEE of Debark, Simien Mts National Park	E
785		Stenoptilia rougeoti	Gibeaux, 1994	Bale, marais de Dinsho	E
786		Stenoptilia tyropiesta	Meyrick, 1932	Mt Chillálo	E
787	Pyralidae	Aglossodes dureti	(Rougeot, 1977)	Arba Minch	E
788		Aglossodes navattae	Rougeot, 1977	Arba Minch	E
789		Bostra excelsa	Rougeot, 1984	near Mt Batu	E
790		Bostra pseudoexcelsa	Rougeot, 1984	Arba Minch	E
791		Dembea venulosella	Ragonot, 1888	Abyssinia [Ethiopia]
792		Ematheudes pollex	Shaffer, 1998	Kosogay Wagra	E
793		Emmalocera erythrinella	(Ragonot, 1888)	Abyssinia [Ethiopia]
794		Endotricha ellisoni	Whalley, 1963	Harar
795		Harraria rufipicta	Hampson, 1930	Harrar [Harar]	E
796		Loryma albilinealis	Hampson, 1917	Diré Daouá [Dire Dawa]	E
797	Pyralidae	Megarthridia christyi	Rougeot, 1984	Arba Minch	E
798		Nussia rougeoti	Leraut, 2015	Koffolé [Koffale]
799	Saturniidae	Aurivillius cadioui	Bouyer, 2008	100 kn E of Addis Ababa	E
800		Bunaeopsis birbiri	Bouvier, 1929	Joubdo (Birbir)	E
801		Bunaeopsis oubie	(Guérin-Méneville, 1849)	Abyssinia [Ethiopia]
802		Eosia digennaroi	Bouyer, 2008	Bale, S of Omar
803		Epiphora antinorii	(Oberthür, 1880)	Scioa [Shoa], Mahal Uonz [Awash River]
804		Epiphora bauhiniae atbarina	(Butler, 1877)	Atbara*
805		Epiphora fourneri	Rougeot, 1974	Road Koffolé-Arussi [Koffale- Arsi]
806		Gonimbrasia belina abayana	Rougeot, 1977	Arba Minch	E
807		Gonimbrasia belina felderi	Rothschild, 1895	Bogos	E
808		Gonimbrasia ellisoni	Lemaire, 1962	Harar	E
809		Gonimbrasia fletcheri	Rougeot, 1960	Ethiopia	E
810		Gonimbrasia fucata	Rougeot, 1978	Ethiopia	E
811		Goodia smithii	(Holland, 1897)	East Africa [Ethiopia]	E
812		Gynanisa arba	Darge, 2008	Arba Minch	E
813		Heniocha digennaroi	Bouyer, 2008	Sidamo, Neguele Borana	E
814		Holocerina digennariana	Darge, 2008	Shashemene (Arsi)	E
815		Ludia hansali	Felder, 1874	Bogos
816		Ludia pupillata	Strand, 1911	Antottos	E
817		Micragone leonardi	Bouyer, 2008	Sidamo, Dilla	E
818		Nudaurelia fasciata	Gaede, 1927	[Ethiopia]	E
819		Nudaurelia ungemachti	Bouvier, 1926	Djemdejm [Jem Jem]	E
820		Pseudobunaea heyeri citrinarius	Gaede, 1927	Harrar [Harar]
821		Pseudobunaea megana	Darge, 2012	Sidamo Province, near Mega	E
822		Urota melichari	Bouyer, 2008	Sidamo Province, 15 km S of Negele	E
823	Scythrididae	Scythris ethiopica	Bengtsson, 2014	Lake Tana, Bahir Dar	E
824	Sesiidae	Agriomelissa aethiopica	(Le Cerf, 1917)	Abyssinia [Ethiopia]	E
825		Jerbeia darkovi	Gorbunov, 2018	Oromia, 21.8 km NW (289.5°) of Dembi Dolo	E
826		Melittia abyssiniensis	Hampson, 1919	Harar	E
827		Melittia ambo	Gorbunov, 2015	West Shewa, 3 km S of Ambo	E
828	Sphingidae	Ceridia heuglini	(Felder C. & Felder R., 1874)	Abyssinia [Ethiopia]
829		Ceridia quirini	Sulak, Naumann & Witt, 2016	Oromia Region, road between Deritu and Dubuluk, near Deritu	E
830		Chaerocina ellisoni	Hayes, 1963	Harar	E
831		Covelliana berioi	Eitschberger & Melichar, 2016	near Debark Gondar	E
832		Covelliana robertbecki	Eitschberger & Melichar, 2016	Ethiopia Central, Oromia, southern Bale Mts, Harenna Forest	E
833		Dovania dargei	Pierre, 2000	Metu	E
834		Dovania neumanni	Jordan, 1926	SW Abyssinia [Ethiopia], Dhimma [Jimma]	E
835		Falcatula tamsi	Carcasson, 1968	Harrar [Harar]	E
836		Leucophlebia neumanni	Rothschild, 1902	Gelo River to Akobo River
837		Lophostethus dumolinii riedeli	Eitschberger & Ströhle, 2011	Arba Minch	E
838	Sphingidae	Lophostethus negus	Jordan, 1926	SW Abyssinia [Ethiopia], Kambatta	E
839		Macropoliana chrismonika	Eitschberger & Melichar, 2016	Ethiopia W, 12 km E of Bonga	E
840		Macropoliana haileselassiei	Eitschberger & Melichar, 2016	Sidamo Province, 20 km S of Angere Maryam	E
841		Macropoliana kingstoni	Eitschberger, 2016	Oromia Region, 25 km E of Bonga/Mera	E
842		Macropoliana stroehlei	Eitschberger, 2016	Near Dorze	E
843		Nephele xylina	Rothschild & Jordan, 1910	Abyssinia [Ethiopia]
844		Platysphinx dorsti	Rougeot, 1977	Kébré-Mengist [Kibre Mengist]	E
845		Praedora melichari	Haxaire, 2011	Sidamo Province, near Bitata	E
846		Pseudoclanis bianchii	(Oberthür, 1883)	Scioa [Shoa]	E
847		Rufoclanis numosae rostislavi	Haxaire & Melichar, 2008	Gamo Gofa Province, Dagabule National Park	E
848		Temnora arida	Melichar & Řezáč & Ilčíková, 2016	Dorze, Guge Mts
849		Temnora robusta	Melichar, Řezáč & Ilčíková, 2016	Kaffa Prov., 40 km SW Jima,
850		Theretra ankae	Melichar & Řezáč, 2015	Asosa	E
851	Thyrididae	Arniocera cyanoxantha	(Mabille, 1893)	Abyssinia [Ethiopia]
852		Arniocera guttulosa	Jordan, 1915	Harar	E
853		Lamprochrysa amata	(Druce, 1910)	Diré Daouá [Dire Dawa]	E
854	Tineidae	Afrocelestis minuta	(Gozmány, 1965)	Gamu-Gofa, Konso
855		Ateliotum convicta	(Meyrick, 1932)	Jem Jem Forest	E
856		Ceratophaga luridula	(Meyrick, 1932)	Mt Chillálo, moorland	E
857		Ceratophaga nephelotorna	(Meyrick, 1932)	Jem-Jem Forest	E
858		Criticonoma spinulosa	Gozmány, 1965	Gamu-Gofa, Konso	E
859		Crypsithyris stenovalva	(Gozmány, 1965)	Gamu-Gofa, Konso	E
860		Dryadaula glycinoma	(Meyrick, 1932)	Jem-Jem Forest	E
861		Ectabola pygmina	(Gozmány, 1965)	Marako
862		Edosa torrifacta	(Gozmány, 1965)	Harrar [Harar]	E
863		Hapsifera gypsophaea	Gozmány, 1965	Gamu-Gofa, Konso	E
864		Hapsifera pachypsaltis	Gozmány, 1965	Kaffa, Ghimira
865		Hapsifera richteri	Gozmány, 1965	Ethiopia SW, Gamu-Gofa, Konso	E
866		Leptozancla zelotica	(Meyrick, 1932)	Jem-Jem Forest	E
867		Monopis addenda	Gozmány, 1965	Kaffa, Gembi [Gimbi]
868		Monopis leopardina	Gozmány, 1965	Kaffa, Abaro	E
869		Monopis triplacopa	Meyrick, 1932	Jem-Jem Forest, 45 miles W. of Addis-Ababa,	E
870		Perissomastix lucifer	Gozmány, 1965	Muger Valley	E
871		Scalmatica separata	Gozmány, 1965	Konso, Gamu-Gofa	E
872		Silosca mariae	Gozmány, 1965	Djerrer Valley
873		Tinissa spaniastra	Meyrick, 1932	Jem-Jem Forest, , 45 miles from Addis Ababa
874	Tortricidae	Acleris baleina	Razowski & Trematerra, 2010	Bale Mountains, Sanetti Plateau	E
875		Acleris harenna	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest, Karcha Camp	E
876		Ancylis colaccii	Razowski & Trematerra, 2012	Wellega Zone, Didessa River	E
877		Bubonoxena alatheta	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest, Karcha Camp	E
878		Choristoneura palladinoi	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest	E
879		Coccothera carolae	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest
880	Tortricidae	Coccothera triorbis	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest	E
881		Coniostola separata	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest, Karcha Camp	E
882		Cosmetra anepenthes	(Razowski & Trematerra, 2010)	Bale Mountains, Harenna Forest, Karcha Camp	E
883		Cosmetra latiloba	(Razowski & Trematerra, 2010)	Bale Mountains, Harenna Forest, Karcha Camp	E
884		Cydia calliglypta	(Meyrick, 1932)	Jem-Jem Forest, edge of forest	E
885		Cydia dinshoi	Razowski & Trematerra, 2010	Bale Mountains, Dinsho Lodge	E
886		Cydia lathetica	Razowski & Trematerra, 2010	Bale Mountains, Dinsho Lodge	E
887		Cydia tytthaspis	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest, Karcha Camp	E
888		Eccopsis aegidia	(Meyrick, 1932)	Jem-Jem Forest
889		Eccopsis brunneopostica	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest, Karcha Camp	E
890		Eccopsis maschalista	(Meyrick, 1932)	Jem-Jem Forest	E
891		Eccopsis subincana	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest	E
892		Endothenia albapex	(Razowski & Trematerra, 2010)	Bale Mountains, Harenna Forest	E
893		Endothenia ethiopica	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest, Karcha Camp	E
894		Epichoristodes spilonoma	(Meyrick, 1932)	Jem Jem Forest
895		Eucosma vulpecularis	Meyrick, 1932	Jem-Jem Forest	E
896		Eucosmocydia zegieana	Razowski & Trematerra, 2018	Amhara, Zegie Peninsula	E
897		Grapholita insperata	Razowski & Trematerra, 2010	Bale Mountains, Dinsho Lodge	E
898		Gypsonoma giorgiae	Razowski & Trematerra, 2012	Ilubabor zone, Bedelle, Dabeda River	E
899		Lozotaenia karchana	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest, Karcha Camp	E
900		Lozotaenia sciarrettae	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest, Karcha Camp	E
901		Megaherpystis oromiae	Razowski & Trematerra, 2018	Oromia, Suba Forest	E
902		Megaherpystis subae	Razowski & Trematerra, 2018	Oromia, Suba Forest	E
903		Megalota lygaria	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest	E
904		Metamesia physetopa	(Meyrick, 1932)	Jem-Jem Forest and Mt Chillálo
905		Multiquaestia aequivoca	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest	E
906		Olethreutes didessae	Razowski & Trematerra, 2012	Wellega zone, Didessa River	E
907		Olethreutes polymorpha	(Meyrick, 1932)	Jem-Jem Forest	E
908		Parabactra addisalema	Razowski & Trematerra, 2018	Oromia, Addis Alem, Ambo Park	E
909		Paraeccopsis addis	Aarvik, 2014	Addis Ababa	E
910		Phtheochroa lonnvei	Aarvik, 2010	Oromia Province, Bale zone, 43 km SW of Goba, Bale Mts National Park, Darwin Camp	E
911		Plutographa xanthala	Razowski & Trematerra, 2010	Bale Mountains, Dinsho Lodge	E
912		Procrica dinshona	Razowski & Trematerra, 2010	Bale Mountains, Dinsho Lodge
913		Procrica ophiograpta	(Meyrick, 1932)	em-Jem Forest and Mt Chillálo
914		Procrica parisii	Razowski & Trematerra, 2010	Bale Mountains, Dinsho Lodge	E
915		Russograptis albulata	Razowski & Trematerra, 2010	Bale Mountains, Harenna Forest	E
916		Thaumatographa amarana	Razowski & Trematerra, 2018	Amhara, Zegie Peninsula	E
917		Thaumatovalva spinai	Razowski & Trematerra, 2010	Omo Valley, Dowro Zone, Tarcha
918		Tortrix diametrica	Meyrick, 1932	Jem-Jem Forest	E
919		Trachybyrsis chionochlaena	Meyrick, 1932	Mt Chillálo	E
920	Uraniidae	Arussiana herbuloti	Rougeot, 1977	near Koffolé [Koffale]
921	Yponomeutidae	Yponomeuta ocypora	(Meyrick, 1932)	Jem-Jem Forest	E
922		Yponomeuta ioni	Agassiz, 2019		E
923		Yponomeuta ocypora	Meyrick, 1932
924		Yponomeuta oromiensis	Agazzia, 2019		E
925	Zygaenidae	Alteramenelikia jordani	(Alberti, 1954)	Abyssinia [Ethiopia]
926		Astyloneura bicoloria	Röber, 1929	Abyssinia [Ethiopia]	E
927		Epiorna abessynica	(Koch, 1865)	Abyssinia [Ethiopia]
928		Saliunca anhyalina	Alberti, 1957	Abyssinia [Ethiopia]	E
929		Saliunca homochroa	(Holland, 1897)	Darde River

1 Rothschild, 1902 2 (Le Moult, 1945) 3 (De Joannis, 1913) 4 Herbulot, 1983 5 Hampson, 1901 6 (Rougeot, 1977). Photos credit: Alenuccio Palladino (1); Francesco Parisi (2); Dirk Stadie (3–6).

Given these numbers, knowledge on the Ethiopian butterflies and moths appear to be particularly unsatisfactory, when compared to their (estimated) potential total numbers with other countries. For instance, the two most diverse European Mediterranean countries, i.e., France and Italy, with a combined land surface comparable to Ethiopia, have ca. 5,109 and 5,086 species of , respectively (Stoch 2003; Wikipedia 2011).

To better evaluate the level of knowledge of the lepidopteran fauna in Ethiopia, and to roughly estimate the real biodiversity, we can compare it with neighboring Kenya, which for several aspects can be considered similar to Ethiopia, but probably it has been better investigated. So far, from Kenya approximately 4,815 lepidopteran taxa were reported, belonging to 63 families (Sáfián et al. 2009; De Prins and De Prins 2019). The currently known number of species in Kenya is almost twice that of Ethiopia, and 15 families are not recorded at all in the latter country. Is it really due to difference in faunal richness between the two coutries or because of the different level of investigation? A better idea can come from the differences observed within the single families. When considering most groups of the ‘Microlepidoptera’, very few investigations were made in Ethiopia and the difference in species numbers between the two countries is huge. Considering only the most species-rich families of Microlepidoptera, the percentage of species present in Ethiopia, compared to the species numbers in Kenya, is 10% for , 13% for , 17% for , 31% for , 34% for , 45% for , 46% of , up to 76% for . However, if we look at the ‘larger moths’ () and butterflies, which are better investigated in both countries, the difference is decreasing from 40% for , 41% for , 50% for , 53% for , 55% for up to 77% for , 79% for , 91% for , peaking to 132% in , where Ethiopia shows a higher number of species than Kenya.

Although the two countries certainly exhibit faunistic differences, due to biogeographic or climatic factors, it seems clear that the Ethiopian fauna is seriously understudied in many groups. By analysing comprehensive revisions of single genera or families accompanied by major collection campaigns in Ethiopia, we can have an idea of the potential biodiversity the country inhabits.

The geometrid genus Warren, 1897 was subject of an extensive review focused on Ethiopian species, based on an investigation carried out in 100 collection localities in the country for more than 15 years, which included an integrative taxonomic analysis based on morphology and DNA barcodes (Hausmann et al. 2016). As a result of this contribution, the species number was raised from eight previously known Ethiopian species to 40, of which 19 were new to science. After the publication, another seven new species for the Ethiopian fauna were described. Authors of the same article estimated the number of Ethiopian geometrids to exceed 700 species once the unidentified material in their hands is examined, which may suggest a more realistic total species number in excess of 1,000 for the whole country.

Another contribution came from the revision that Hacker carried out on the subfamily (; Hacker et al. 2012; Hacker 2014), where many of the published data concerned sub-Saharan Africa. For Ethiopia, only three species were previously reported. After Hacker’s monograph, the number was raised to 61 species, with 27 newly described taxa from Ethiopia. For Kenya, he raised the figure from 12 to 73, a number not far from that of Ethiopia.

Although these are two examples of taxonomically particularly difficult groups, we can assume similar multiplicators for the so called ‘Microlepidoptera’ resulting in an estimate for the entire order of in Ethiopia which may exceed 10,000 species, of which a number of species new for science. This estimate is based on, and in concordance with the usual ratio of geometrid species number versus lepidopteran species number of roughly 1:10, and on the usual ratio of the (400+ species in Ethiopia) versus lepidopteran species number of roughly 1:20, as it results from large museum material (e.g. ZSM) and from various fauna inventories (e.g. Bavaria: Haslberger and Segerer 2016; Europe: Karsholt and Razowski 1996; North America: Hodges et al. 1983). For the moth fauna of Africa, 38,988 species group names of them are listed by Afromoths (2019), of which 5510 (14%) are geometrids. The total number, however, does not include names, with 4405 species (Williams, 2018) and Microlepidoptera taxonomy is underrepresented, hence also here the “10%-rule” for the ratio seems to apply, at least roughly.

Data from DNA barcoding

In the framework of the international Barcode of Life initiative, DNA barcodes (658bp 5’ COI gene fragment, cf. Hebert et al. 2003) have been assembled for Ethiopian since 2006 with the aim to establish a national DNA reference library for integrated taxonomic studies. So far, 3160 DNA barcodes have been generated from Ethiopian (including many Ethiopian type specimens), belonging to 1012 genetic clusters (Barcode Index Numbers, ‘BINs’) which are a good proxy for real species numbers (Ratnasingham and Hebert 2013; Hausmann et al. 2013). Most DNA barcodes could be assembled in the (2290 barcodes, 571 BINs), (314 barcodes, 165 BINs) and (246 barcodes, 143 BINs). Species coverage is particulary good in the smaller families such as the (121 barcodes, 36 BINs) and (70 barcodes, 24 BINs), while it is still being very poor in the ‘Microlepidoptera’. All images and most metadata and molecular data are accessible in the public database BOLD (Ratnasingham and Hebert 2007).

Actual constraints and future perspectives of research on Diversity of Ethiopia

Butterflies and moths are a major component of biodiversity playing a crucial role in the ecosystem as primary consumers, essential part of food-chains and pollinators. However, humans are exerting unprecedented pressures on all of the earth’s ecosystems, and such pressures may affect all species (Sanchez-Bayo and Wyckhuyes 2019). Nature conservation strategies have focused most of their attention on the “charismatic megafauna”, i.e., on mammals, birds, and other vertebrates. The vast majority of invertebrate species – although accounting for more than 80% of the animal species - are too poorly known to allow an assessment of how they are affected by human activities, and what might be done to mitigate the damage that humans cause. In most cases, the best way that can be done is to conserve their habitats so that most inhabiting species will continue to thrive.

The greatest threats to butterflies and moths are habitat fragmentation and destruction, intensification of agricultural practice with over-use of pesticides and herbicides; climate change mainly affecting endemic species adapted to mountainous habitats, whereas scientific collecting is absolutely negligible (Hausmann 2001; Sanchez-Bayo and Wyckhuyes 2019). In general, human activity is enormously threatening the global diversity of life on the planet. Rough estimates suggest that we are currently undergoing not only unprecedented, but also accelerating rates of species extinction (UNEP 2006; Sanchez-Bayo and Wyckhuyes 2019).

In the same manner, Ethiopia is experiencing major biodiversity loss, mainly related to extensive destruction of habitats, deforestation, land degradation, intensive agricultural expansion, climate change, excessive pesticide and herbicide use, introduction of exotic plant species, among others (EBI 2015; Tesfu et al. 2018). The loss of primary or native forest areas, due to clearcutting and conversion into agroforests, farmland or settlements, are currently the major threat to the Ethiopian biodiversity in general and in particular.

Despite Ethiopia being known for its rich heritage of biological diversity and many diverse ecosystems, the conservation of its habitats have received scant attention. The system of protected areas so far established includes 21 national parks, four sanctuaries, eight wildlife reserves, 20 controlled hunting areas, six open hunting areas, six community conservation areas and 58 national forest priority areas (Young 2012), covering 14% of the country (EBI 2015). However, most of its biodiversity, including , is still unexplored because of significant lack of national research capacity. Hence, in parallel to conservation programs and sustainable utilisation of biological resources, efforts for the preparation of a comprehensive bio-inventory should receive highest priority. Such an instrument must be considered an essential baseline for policy makers, planners, donors and researchers working on biodiversity conservation in Ethiopia.

In order to upsurge biodiversity knowledge, capacity building in the area at various levels is needed. Lack of well organised natural history museums, specialists, and scientific societies providing support and fostering citizen science, international research networks and projects are among the identified gaps. Currently, most of the type specimens and reference collections are deposited outside the country of origin. In this context, the Nagoya Protocol (UNSG 2010), although intending to strengthen nations to conserve their genetic resources, to some extent could lead to the opposite effect by hampering international collaboration. Joint protocols and agreements between national actors (research institutes, governing agencies, universities, NGO’s) and international research bodies should be promoted in a collaborative way, favoring shared, non-commercial biodiversity research. Close collaboration with museums and universities possessing reference collections and skills, designing and organising projects are required to teach and train a generation of highly competent scientists and managers so that collections of Ethiopian insects could be built and properly managed. In absence of these minimum requirements, establishing a national entomological museum/collection could be ineffective in promoting the study and conservation of local biodiversity resources.