Gousi Li1,2,3, Yao-Guang Liu4,5,6, Yuanling Chen7,8,9. 1. State Key Laboratory for Conservation and Utilization of Subtropical Agro-Bioresources, South China Agricultural University, Guangzhou, 510642, China. 2. Key Laboratory of Plant Functional Genomics and Biotechnology of Guangdong Provincial Higher Education Institutions, South China Agricultural University, Guangzhou, 510642, China. 3. College of Life Sciences, South China Agricultural University, Guangzhou, 510642, China. 4. State Key Laboratory for Conservation and Utilization of Subtropical Agro-Bioresources, South China Agricultural University, Guangzhou, 510642, China. ygliu@scau.edu.cn. 5. Key Laboratory of Plant Functional Genomics and Biotechnology of Guangdong Provincial Higher Education Institutions, South China Agricultural University, Guangzhou, 510642, China. ygliu@scau.edu.cn. 6. College of Life Sciences, South China Agricultural University, Guangzhou, 510642, China. ygliu@scau.edu.cn. 7. State Key Laboratory for Conservation and Utilization of Subtropical Agro-Bioresources, South China Agricultural University, Guangzhou, 510642, China. pcb2000@scau.edu.cn. 8. Key Laboratory of Plant Functional Genomics and Biotechnology of Guangdong Provincial Higher Education Institutions, South China Agricultural University, Guangzhou, 510642, China. pcb2000@scau.edu.cn. 9. College of Life Sciences, South China Agricultural University, Guangzhou, 510642, China. pcb2000@scau.edu.cn.