Jon E Keeley1,2, Philip van Mantgem3, Donald A Falk4. 1. U.S. Geological Survey, Western Ecological Research Center, Sequoia-Kings Canyon Field Station, Three Rivers, CA, USA. jon_keeley@usgs.gov. 2. Department of Ecology and Evolutionary Biology, UCLA, Los Angeles, CA, USA. jon_keeley@usgs.gov. 3. U.S. Geological Survey, Western Ecological Research Center, Redwoods Field Station, Arcata, CA, USA. 4. School of Natural Resources and the Environment, University of Arizona, Tucson, AZ, USA.