Haijun Wen1, Hurng-Yi Wang2, Xionglei He1, Chung-I Wu1,3. 1. State Key Laboratory of Biocontrol, College of Ecology and Evolution, Sun Yat-Sen University, Guangzhou. 2. Graduate Institute of Clinical Medicine and Hepatitis Research Center, Taiwan University and Hospital, Taipei. 3. Department of Ecology and Evolution, University of Chicago, Chicago.