Chunlei Zhang1, Youming Zhang1, Zhaojun Ding2, Yang Bai3. 1. State Key Laboratory of Microbial Technology, Shandong University, Qingdao, Shandong 266237, China; The Key Laboratory of Plant Development and Environmental Adaptation Biology, Ministry of Education, College of Life Sciences, Shandong University, Qingdao, Shandong 266237, China. 2. State Key Laboratory of Microbial Technology, Shandong University, Qingdao, Shandong 266237, China; The Key Laboratory of Plant Development and Environmental Adaptation Biology, Ministry of Education, College of Life Sciences, Shandong University, Qingdao, Shandong 266237, China. Electronic address: dingzhaojun@sdu.edu.cn. 3. State Key Laboratory of Plant Genomics, Institute of Genetics and Developmental Biology, Chinese Academy of Science, Beijing 100101, China; CAS-JIC Centre of Excellence for Plant and Microbial Science, Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing 100101, China; University of Chinese Academy of Sciences, College of Advanced Agricultural Sciences, Beijing 100039, China. Electronic address: ybai@genetics.ac.cn.