| Literature DB >> 30450439 |
Tibebu Kassawmar1,2, Gete Zeleke3,2, Amare Bantider2, Gizaw Desta Gessesse2, Lemlem Abraha4.
Abstract
This paper demonstrates synoptic ways of presenting and characterizing land change processes across Ethiopia's large, complex Rainfed Agricultural Area (RAA). We translated pixel-level detected changes into neighbourhood-level changes that are useful to decision-makers. First, we identified pixel-level changes without and with type/direction of change, based on land cover maps from the years 1986 and 2010. For type-/direction-based characterization, we sorted observed transitions into four categories of prominent land change processes ("forest degradation", "deforestation", "afforestation", and "no change"). Adopting appropriate window sizes for identified ecoregions in the study area, we ran a focal statistics summation operator separately on the two change rasters (with/without consideration of direction of change). The results obtained by applying the approach can be described in relative terms as well as qualitative terms, using ranges of change values that can be further classified using qualitative terms, i.e. ranging from "no change" to "high/substantial change". Our non-directional change assessment result showed that approximately 6% of the RAA is characterized by substantial change, whereas 40% appears stable ("no change"). Based on the directional-change assessment results, 3% of deforestation, 4% of forest degradation, and 3% of revegetation processes were found to constitute "high/substantial change". The types and intensity of landscape transformations display distinct spatial patterns linked to agro-ecological belts and socio-economic dynamics. Minimal reverse changes were observed on some severely degraded lands in the highlands, but the overall per cent cover remains relatively small. Overall, vegetation degradation still exceeds regeneration by more than half a per cent. Relatively lower altitudes and middle altitudes exhibit higher transformation. The presented approach and resulting outputs can provide planners and decision-makers with a synoptic view of land change processes. It can support policy formulation of sustainable land management and rehabilitation activities of the agricultural ecosystem at national and regional scales.Entities:
Keywords: Geography; Geoscience
Year: 2018 PMID: 30450439 PMCID: PMC6226589 DOI: 10.1016/j.heliyon.2018.e00914
Source DB: PubMed Journal: Heliyon ISSN: 2405-8440
Fig. 1The study area boundaries showing Ethiopia's RAA and the ecoregions.
Identified and mapped ecoregions featured in the Rainfed Agricultural Area (RAA) of Ethiopia.
| Major ecoregions | Sub-ecoregions | Codes | Ecoregion divides |
|---|---|---|---|
| Forestry- and semi-agroforestry- dominated ecoregions | Natural high forest, protected areas, national forest reserves, areas of coffee cultivation and other silvo-cultural practices | 1a | Highlands |
| Agroforestry- dominated ecoregions | Distinct agroforestry system and mosaic of croplands with trees and high rainfall areas | 1b | |
| Mixed agricultural system (moderately cultivated) ecoregions | Disturbed forest, artificial forest, mosaic of crops with high trees, high-potential and high-rainfall areas | 2a | |
| Mixed agricultural system (intensively cultivated) ecoregions | Intensively cultivated, high-potential, and high-rainfall areas | 3a | |
| Intensively cultivated, low-potential, and moderate-rainfall areas | 3b | ||
| Intensively cultivated, low-potential, and low-rainfall areas | 3c | ||
| Agro-pastoralist system (lightly cultivated) ecoregions | Lightly cultivated, moderate-potential, lowland, and low-rainfall areas | 4a | Lowlands |
| Dominantly pastoralist ecoregion | Wooded land in the lowlands and moderate-rainfall areas where slash-and-burn cultivation is rarely practiced | 5a | |
| Wooded land in the lowlands and low rainfall areas where slash-and-burn cultivation is rarely practiced | 5b | ||
| Wooded land in the lowlands and high rainfall areas where slash-and-burn cultivation is rarely practiced | 5c |
Altitudinal ranges (meters) and corresponding traditional Agro-Ecological Belts (AEB) in the study area.
| Altitude range | 500–1,000 | 1,000–1,500 | 1,500–2,000 | 2,000–2,300 | 2,300–3,000 | 3,000–3,700 | >3,700 |
| Traditional agro-ecological names | Lower Kolla | Upper Kolla | Lower Woyna Dega | Upper Woyna Dega | Lower Dega | High Dega | Wurch |
| Proportional area coverage (%) | 17 | 26 | 30 | 11 | 14 | 2 | 0.2 |
The classification scheme applied to map the two categories of LULC (I and II) for the two periods.
| I level classes | SN | II level classes | Brief description of the classes | Code | I level classes | SN | II level classes | Description of the classes | Code |
|---|---|---|---|---|---|---|---|---|---|
| Water body | 1 | Water body | Lakes and Ponds | WB | Agroforestry | 24 | Agroforestry 7 | Dominantly Mango and other fruit system | AFma |
| Settlement and built-up | 2 | Settlement | All urban and dense rural settlements mixed with several lands cover and uses | ST | 25 | Agroforestry 8 | Dominantly Banana, mango and other fruit tree system | AFbm | |
| 3 | Homestead plantation | Dense plantations around rural settlements | HP | 26 | Agroforestry 9 | Enset-Banana and Mango mixed system | AFenbm | ||
| 4 | Built-up (concrete) | Only concert part of the settlement | BU | Grassland | 27 | Grassland (Drained) | Drained and dry grasslands found around farm borders and sloping landscapes | GD | |
| Forest | 5 | High forest | Moist ever green forests | HF | 28 | Grassland (Wet or un-drained) | Often found along water course, water bodies and upland depressions | GW | |
| 6 | Dry Forest | Deciduous dry region forests (lowlands of the country) | DF | 29 | Savanna grassland | Tall and extensive found in dry and flat terrain | SG | ||
| 7 | Riverine Vegetation | Any type of woody vegetation exist following the river courses | RV | Wetland | 30 | Swamp | All the time wet, often covered by grass, found along riversides, lakes | SW | |
| 8 | Mixed Forest | Enriched degraded natural forest | MF | 31 | Marshland | Intermittently flooded and muddy | ML | ||
| 9 | Church Forest | Protected patches of forests in religious sites | CF | Cropland | 32 | Large scale farm (commercial) | Agricultural lands owned by non-substance or smallholder farms | LSI | |
| 10 | Plantation forest | Private and community forests dominantly Eucalyptus | PF | 33 | Shifting Cultivation | Cleared/burnt woodland for crop production managed by smallholders | SC | ||
| 11 | Bamboo Forest | Perineal grass plant community having hollow circular stems | BF | 34 | Cropland with trees | Croplands with scattered trees managed by smallholders | CT | ||
| Woodland | 12 | Dense Woodland | Densely spaced trees but dominated by single species, largely found in the lowlands | WLd | 35 | Croplands (with no trees) | Open crop fields managed by smallholders | CWOT | |
| 13 | Open Woodland | A woodland but degraded and has very scattered trees | WLo | 36 | Croplands (in hilly terrain) | Intermittently cultivated often in a rugged terrain | CH | ||
| Shrubland/Bushland | 14 | Dense Shrub and Bush | Non-tree but woody plants often < 5m height, densely spaced, found in inaccessible highland upslope and arid areas | SBd | 37 | Irrigated fields Croplands | Managed for dry season crop production | IF | |
| 15 | Open Shrub and Bush | Exist as scattered plant community and often as mosaic with grass plants | SBo | Barren land | 38 | Degraded hills | Highly degraded lands with exposed rock or soil | DH | |
| Afro-alpine vegetation | 16 | Sub-afro Alpine Vegetation | Dominated by woody vegetation (Eg.Erica bush) | AAbs | 39 | Exposed Rock | Landscapes with exposed rocks and little bit covered by annual grass species | ER | |
| 17 | Afro Alpine Vegetation | Dominated by herbaceous vegetation (Grass) | AAg | 40 | Extensive unpalatable | Herbs A landscape covered by unpalatable herbaceous species | HS | ||
| Agroforestry | 18 | Agroforestry 1 | Enset dominating system | AFen | 41 | Salt Flat Deposition | Extensive salt depositions exposed on the surface | SFd | |
| 19 | Agroforestry 2 | Crop mixed with Khat-system | AFck | 42 | Construction sites | Mining, quarry and other construction site | MS | ||
| 20 | Agroforestry 3 | Enset mixed with Khat system | AFek | 43 | Exposed Sand | Exposed surface commonly sandy surface | ES | ||
| 21 | Agroforestry 4 | Dominantly Coffee system | AFcof | 44 | Exposed surface | Exposed surface but the type of surface cover is unknown | ESc | ||
| 22 | Agroforestry 5 | Enset and Coffee system | AFec | River bed | 45 | Wider river course | Intermittently covered by water, boulders and gravels | RC | |
| 23 | Agroforestry 6 | Dominantly Banana system | AFba | Others | 46 | Others | Any other LULC which is not identifiable by Landsat images | OT |
Fig. 2Detailed Level II LULC map produced for the present assessment.
Fig. 3Generalized (Level II) LULC maps used for the present assessment.
Fig. 4Schematic representation of the workflow
Independent LULC change scientific studies conducted across the study area (RAA) (AP refers aerial photographs and SI refers satellite images).
| Validation site codes | Authors | Study area/location | Method of study | Area (km2) | Period of analysis |
|---|---|---|---|---|---|
| 1 (1a) | ( | The southwest natural forest of Ethiopia (Metu) Dizi catchment | AP of 1957, 1976, 1982, and 1990 | 7 | 1957–1990 |
| 2 (1a) | ( | The southwest natural forest of Ethiopia (Metu) Gey catchment | AP of 1957, 1976, 1982 and 1990 | 10 | 1957–1990 |
| 3 (1a, 5c, 2a) | ( | Southwest/Baro Akobo Basin | AP of 1971 & 1975 and SI of 1986 & 1990 | 30,000 | 1973–1990 |
| 4 (1a) | ( | Fincha Watershed/Blue Nile Basin | AP for 1957 &1980 and SI of 2001 | 1318 | 1957–2001 |
| 5 (3a) | ( | Gilgel Abay/Blue Nile Basin | AP of 1957 & 1975 and SI of 1986 & 2009 | 4570 | 1957–2009 |
| 6 (3a) | ( | Jedeb/Blue Nile Basin | AP of 1957 & 1975 and SI of 1986 & 2009 | 296 | 1957–2009 |
| 7 (3a) | ( | Demebecha woreda/Blue Nile Basin | AP of 1957 & 1982 and SI of 1995 | 271 | 1957–1995 |
| 8 (3b) | ( | Blue Nile Basin, Tara Gedam catchment | AP of 1957 & 1975 and SI of 1986 & 2009 | 9 | 1957–2009 |
| 9 (3c) | ( | Gilgel Tekeze catchment | SI of 1986, 1995 & 2010 | 352 | 1976–2008 |
| 10 (3c) | ( | Maileba and Gumselassa catchment, Tigray | AP of 1964 &1994 and field survey of 2006 | 40.8 | 1964–2006 |
| 11 (3c) | ( | Maileba and Gumselassa catchment, Tigray | AP of 1964 &1994 and field survey of 2006 | 40.8 | 1964–2006 |
| 12 (4a and 5c) | ( | The western Ethiopia | SI of 1986, 1995 & 2010 | 3806, 6328 & 3518 | 1986–2010 |
| 13 (4a and 5b) | ( | The western Ethiopia | SI of 1986, 1995 & 2010 | 3806, 6328 & 3518 | 1986–2010 |
| 14 (5b) | ( | The western Ethiopia | SI of 1986, 1995 & 2010 | 3806, 6328 & 3518 | 1986–2010 |
Fig. 5Location and extent of selected independent LULC change research studies used to validate our change estimates.
Fig. 6The nature of commonly occurring changes (Landsat TM images with false colour composite 4-3-2; codes refer to sub-ecoregions described in Fig. 1). 3b1 & 3b2 = Large-scale afforestation/regeneration of degraded (crop)lands; 3c & 2a = Construction of hydro dams (large-scale changes) and dense burning of woody vegetation for shifting cultivation; 3a3 = Large-scale deforestation of natural forest; 3a4, 3a5, & 3a2 = Small-scale afforestation and deforestation; 5b = Small-scale deforestation for cropland expansion; 4a = Extensive burning of wood/shrub land for shifting cultivation.
The four main types/descriptions of landscape transformation, identified based on consolidation of observed transitions.
| Type of landscape transformation | Code | Description |
|---|---|---|
| Forest degradation | FD | Change transitions characterized by modifications of forest landscape, e.g. from forest to woodland or from woodland to shrubland/bushland. |
| Deforestation | DF | Transitions characterized by conversion of vegetated landscape to non-vegetated landscape. In addition, transitions that induce further land degradation, such as change from cropland to barren lands or conversions of grasslands into wetlands |
| Re-afforestation or Regeneration | RA or RG | Change characterized by forest regrowth due to strict forest protection policies (e.g. in the south), or due to restoration efforts or use changes (e.g. area closures in the north and northeast; afforestation practices on state and communally owned lands in the central region). Also included are transitions linked to wetland restoration. |
| Persistence | P | All pixels that represent no change. |
Fig. 7A four-quadrant conceptual model produced from the change matrix used to describe the type/direction of landscape transformations: forest degradation (FD), deforestation (DF), persistence (P), and afforestation (AF). F=Forest, WL = Woodland, SBL = Shrubland/Bushland, CL = Cropland, GL = Grassland, BL = Bare land, WeL = Wetland, WB = Waterbody, and BU = Built-up
Fig. 8Pixel-based changes summarized for major ecosystems (upper left, forest ecosystems; upper right, cropland ecosystems; lower left, grassland ecosystems; lower right, non-woody vegetation ecosystems).
Summary of overall land change statistics estimated by the present study and selected scientific studies.
| Validation site codes | Representing ecoregion | Overall land change estimates (in %) for each period of analysis | Variation (on overlapping periods) | |||||
|---|---|---|---|---|---|---|---|---|
| Present study estimates | Reference data estimates | |||||||
| Total area (km2) | 1986–2010 | Before 1980s | 1980s–1990s | 1990s–2000s | 1980s–2010 | |||
| 1957–1982 | 1982–1990 | |||||||
| 1 | 1ai | 7 | 23 | 30 | 50 | |||
| 2 | 1aii | 9 | 17 | 25 | 30 | |||
| 1971–1997 | ||||||||
| 3 | 1aiii | 160,000 | 17 | 27 | ||||
| 1957–1980 | 1980–2001 | |||||||
| 4 | 2ai | 1318 | 38 | 48 | - | 43 | -5 | |
| 1974–1986 | 1986–1995 | 1995–2010 | 1986–2010 | |||||
| 5 | 3ai | 4570 | 23 | 22 | 8 | 10 | 18 | +5 |
| 1972–1986 | 1986–1994 | 1994–2009 | ||||||
| 6 | 3aii | 296 | 18 | 38 | 29 | 25 | 54 | -26 |
| 1957–1982 | 1982–1995 | |||||||
| 7 | 3aiii | 271 | 9 | 68 | 14 | |||
| 1957–1986 | 1986–1995 | 1995–2011 | 1986–2011 | |||||
| 8 | 3bi | 9 | 22 | 91 | 10 | 10 | 20 | +2 |
| 1976–1986 | 1986–2008 | |||||||
| 9 | 3bii | 352 | 21 | 43 | - | - | 16 | +14 |
| 1964–1994 | _ | 1994–2006 | ||||||
| 10 | 3ci | 24 | 25 | 28 | _ | 25 | ||
| 11 | 3cii | 17 | 35 | 10 | _ | 38 | ||
| 1985–1995 | 1995–2010 | 1985–2010 | ||||||
| 12 | 4ai | 3806 | 51 | 19 | 26 | 47 | +4 | |
| 13 | 5bi | 6328 | 42 | 13 | 23 | 37 | +5 | |
| 14 | 5ci | 3518 | 33 | 10 | 35 | 45 | +8 | |
| Weighted average percentage of change | 27 | 8 | 31 | 14 | 24 | +3 | ||
The most-frequent transitions occurring in each ecoregion and their class-level proportional area coverage.
| Transition type | Trajectory | Direction | Occurrence and density of transitions in each sub-ecoregion | ||||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
| 1a | 1b | 2a | 3a | 3b | 3c | 4a | 5a | 5b | 5c | RAA | |||
| WL–F | RG | RLT | 21.1 | 0.5 | 6.1 | 1.9 | 2.1 | 0.7 | 1.6 | 0.2 | 6.3 | 4.4 | |
| SBL–F | RG | RLT | 15.1 | 0.2 | 9.5 | 6.2 | 2.1 | 0.2 | 0.1 | 0.1 | 0.8 | 3.8 | |
| CL–F | RG | RLT | 1.8 | 0.2 | 3.9 | 2.2 | 0.1 | 0.04 | 0.4 | 0.9 | |||
| F–WL | FD | FLT | 10.7 | 4.1 | 6.3 | 2.4 | 0.2 | 4.9 | 0.6 | 6.3 | 3.5 | ||
| F–Cl | DF | FLT | 0.5 | 6.4 | |||||||||
| WL–CL | DF | FLT | 9.5 | 3.2 | 31.3 | 13.6 | 5.8 | 0.9 | 8.6 | 1.1 | 3.1 | 10.3 | 9.5 |
| GL–CL | DF | FLT | 7.5 | 5.1 | 12.4 | 28.1 | 6.4 | 2 | 3.6 | 0.6 | 3.8 | 3.6 | 6.4 |
| WL–SBL | FD | FLT | 6.3 | 3.3 | 5.8 | 5.2 | 12.1 | 21.4 | 12.8 | 17.6 | 32.1 | 34.1 | 16.4 |
| WL–GL | DF | FLT | 5.1 | 1.4 | 4 | 1.9 | 2.5 | 1.1 | 9.1 | 7.2 | 7.1 | 6.8 | 5.0 |
| F–SBL | DF | FLT | 4.8 | 0.6 | 1.4 | 1 | 0.8 | 0.2 | 1.3 | 0.7 | 1.4 | 1.3 | |
| GL–WL | RG | RTL | 3.3 | 1.1 | 1 | 7.2 | 2.5 | 0.8 | 1.8 | ||||
| GL–SBL | RG | RTL | 2.3 | 1 | 1.4 | 1.1 | 0.4 | 3.5 | 8.9 | 2.2 | 1.3 | 2.5 | |
| F–GL | DF | FLT | 1.8 | 1.4 | 1.3 | 1 | 0.1 | 0.6 | 0.1 | 0.5 | |||
| GL–F | RG | RTL | 2.8 | 1.8 | 1.5 | 0.1 | 0.03 | 0.1 | 0.7 | ||||
| SBL–F | RG | RTL | 2.6 | 1.8 | 0.6 | 0.1 | 0.8 | 0.7 | |||||
| SBL–WL | RG | RTL | 3.4 | 1 | 2 | 9.1 | 10.9 | 3.6 | 10.1 | 10.3 | 6.7 | 6.3 | |
| SBL–GL | DF | FLT | 1.4 | 3.0 | 1.3 | 6.3 | 4 | 7.4 | 13.8 | 14.8 | 15.2 | 9.1 | 8.5 |
| CL–WL | RG | RTL | 1.1 | 1.1 | 0.6 | 2.1 | 0.5 | ||||||
| SBL–CL | DF | FLT | 1.8 | 3.2 | 3.8 | 7.5 | 8.1 | 8.9 | 26.1 | 0.7 | 6.5 | 4.4 | 7.7 |
| CL–GL | RG | RTL | 1.8 | 2.5 | 4.1 | 0.7 | 8.7 | 3.2 | 0.9 | 2.4 | |||
| WeL–GL | DF | FLT | 1.7 | 2.6 | 0.5 | ||||||||
| GL–F | RG | RTL | 1.0 | 0.1 | |||||||||
| CL–SBL | RG | RTL | 1.0 | 1.6 | 1.19 | 0.9 | 5.8 | 1.2 | 14 | 1.5 | 3.0 | ||
| GL–BL | DF | FLT | 2 | 0.4 | 1.3 | 0.4 | |||||||
| CL–WL | RG | RTL | 1.8 | 0.3 | 1.1 | 0.4 | |||||||
| SBL–BL | DF | FLT | 1.4 | 12.5 | 28.8 | 1.5 | 5.1 | 0.4 | 5.5 | ||||
| F–BL | DF | FLT | 1.2 | 1.2 | 1.6 | 0.1 | 0.3 | 0.5 | |||||
| WL–BL | DF | FLT | 1.1 | 4.2 | 2.9 | 0.2 | 1.7 | 0.5 | 1.2 | ||||
| SBL–F | RG | RTL | 1.1 | 3.7 | 0.7 | 0.6 | |||||||
| CL–BL | DF/FD | FLT | 0.8 | 6.1 | 2.4 | 1.0 | 0.3 | 1.2 | |||||
| Proportion of changed pixels in each sub-ecoregion (in per cent) | 21 | 41 | 24 | 36 | 36 | 59 | 40 | 58 | 43 | 39 | |||
FD = Forest Degradation, D = Deforestation, RG = Regeneration, P = Persistence, FLS = Forward Landscape Transformation, and RLT = Reverse Landscape Transformation
Neighbourhood-level amount of change (number or per cent of change per window) summarized for the nine sub-ecoregions (statistics derived from Fig. 7).
| Code | Change category | Amount of change (in %) for each ecoregion | Average for the RAA | |||||||||
|---|---|---|---|---|---|---|---|---|---|---|---|---|
| 1a | 1b | 2a | 3a | 3b | 3c | 4a | 5a | 5b | 5c | |||
| 1 | Not transformed (<10%) | 59 | 44 | 52 | 56 | 45 | 40 | 13 | 33 | 28 | 30 | 40 |
| 2 | Slightly transformed (10–25%) | 11 | 21 | 21 | 14 | 22 | 25 | 41 | 25 | 26 | 28 | 23 |
| 3 | Moderately transformed (25–50%) | 7 | 15 | 12 | 10 | 15 | 15 | 41 | 22 | 29 | 28 | 19 |
| 4 | Highly transformed (50–75%) | 7 | 11 | 13 | 9 | 13 | 14 | 4 | 13 | 15 | 11 | 11 |
| 5 | Substantially transformed (>75%) | 17 | 9 | 2 | 12 | 5 | 6 | 0 | 6 | 2 | 4 | 6 |
| Total | 100 | 100 | 100 | 100 | 100 | 100 | 100 | 100 | 100 | 100 | 100 | |
Fig. 9Amounts and patterns of landscape transformation, measured based on a merger of multi-scale analyses that proved well suited to depicting changes in different ecoregions.
Fig. 10Maps of the four main types of landscape transformation. The amount of change was obtained by calculating the total number of changed pixels out of maximum possible number of pixels in each respective window.
The amount of change summarized in each type of landscape transformation process occurring at different altitudinal ranges (traditional Agro-Ecological Belts, or AEB).
| Type of landscape transformation | Change category | Lower Kolla | Upper Kolla | Lower Woyna Dega | Upper Woyna Dega | Lower Dega | Higher Dega | Wurch | Average for the RAA |
|---|---|---|---|---|---|---|---|---|---|
| Forest degradation (FD) | 1 (<10%) | 41.2 | 41.5 | 51.5 | 52.4 | 60.6 | 73.4 | 91.3 | 58.8 |
| 2 (10–25%) | 31.7 | 31.5 | 28.1 | 29.6 | 23.5 | 17.1 | 6.1 | 23.9 | |
| 3 (25–50%) | 22.8 | 20.2 | 16.1 | 14.8 | 12.7 | 8.1 | 1.9 | 13.8 | |
| 4 (50–75%) | 4.1 | 5.8 | 3.7 | 2.6 | 2.6 | 1.0 | 0.6 | 2.9 | |
| 5 (>75%) | 0.2 | 1.1 | 0.7 | 0.6 | 0.6 | 0.4 | 0.1 | 0.5 | |
| Deforestation (DF) | 1 (<10%) | 24.9 | 31.8 | 69.4 | 80.5 | 89.0 | 89.5 | 95.5 | 68.7 |
| 2 (10–25%) | 41.7 | 30.9 | 18.8 | 14.7 | 8.8 | 8.7 | 3.4 | 18.1 | |
| 3 (25–50%) | 28.0 | 28.5 | 9.9 | 4.2 | 1.8 | 1.5 | 0.8 | 10.7 | |
| 4 (50–75%) | 5.2 | 8.1 | 1.8 | 0.5 | 0.3 | 0.2 | 0.2 | 2.3 | |
| 5 (>75%) | 0.3 | 0.7 | 0.1 | 0.1 | 0.1 | 0.1 | 0.1 | 0.2 | |
| Persistence (P) | 1 (<10%) | 0.5 | 1.6 | 1.6 | 1.3 | 1.1 | 1.7 | 0.9 | 1.2 |
| 2 (10–25%) | 7.6 | 8.7 | 5.1 | 3.4 | 2.9 | 4.5 | 2.4 | 4.9 | |
| 3 (25–50%) | 40.8 | 43.4 | 24.5 | 17.7 | 18.2 | 18.4 | 8.1 | 24.4 | |
| 4 (50–75%) | 34.3 | 33.0 | 38.5 | 39.3 | 37.8 | 37.8 | 19.6 | 34.3 | |
| 5 (>75%) | 16.9 | 13.3 | 30.2 | 38.3 | 40.1 | 37.5 | 69.1 | 35.1 | |
| Regeneration (RG) or Revegetation (RV) | 1 (<10%) | 53.1 | 60.5 | 58.6 | 61.9 | 60.8 | 54.3 | 63.3 | 58.9 |
| 2 (10–25%) | 28.6 | 26.6 | 30.3 | 28.0 | 28.6 | 27.2 | 21.0 | 27.2 | |
| 3 (25–50%) | 15.6 | 11.0 | 9.8 | 9.0 | 9.3 | 13.9 | 11.2 | 11.4 | |
| 4 (50–75%) | 2.4 | 1.5 | 1.1 | 0.9 | 1.0 | 3.5 | 3.5 | 2.0 | |
| 5 (>75%) | 0.2 | 0.4 | 0.2 | 0.2 | 0.2 | 1.0 | 1.0 | 0.5 |