Ross N Cuthbert1,2, Amanda Callaghan2, Jaimie T A Dick1. 1. Institute for Global Food Security, School of Biological Sciences, Queen's University Belfast, Medical Biology Centre, Belfast, BT9 7BL, Northern Ireland. 2. Ecology and Evolutionary Biology, School of Biological Sciences, University of Reading, Harborne Building, Reading, RG6 6AS, England.