Qiaohong Duan1, Alice Y Cheung2,3,4. 1. State Key Laboratory of Crop Biology, College of Horticulture Science and Engineering, Shandong Agricultural University, Tai-An, China. 2. Department of Biochemistry and Molecular Biology, University of Massachusetts, Amherst, MA, USA. 3. Molecular and Cellular Biology Program, University of Massachusetts, Amherst, MA, USA. 4. Plant Biology Graduate Program, University of Massachusetts, Amherst, MA, USA.