Bert W Hoeksema1,2, Jessica Bouwmeester3, Pedro Range3, Radhouan Ben-Hamadou3. 1. Taxonomy and Systematics Group, Naturalis Biodiversity Center, PO Box 9517, 2300 RA, Leiden, The Netherlands. 2. Institute of Biology Leiden, Leiden University, PO Box 9505, 2300 RA, Leiden, The Netherlands. 3. Department of Biological and Environmental Sciences, College of Arts and Sciences, Qatar University, PO Box 2713, Doha, Qatar.