Jia-Wei Wang1,2, Yijun Qi3,4. 1. National Key Laboratory of Plant Molecular Genetics (NKLPMG), CAS Center for Excellence in Molecular Plant Sciences, Institute of Plant Physiology and Ecology (SIPPE), Shanghai Institutes for Biological Sciences (SIBS), Shanghai, 200032, China. jwwang@sibs.ac.cn. 2. ShanghaiTech University, Shanghai, 200031, China. jwwang@sibs.ac.cn. 3. Center for Plant Biology, School of Life Sciences, Tsinghua University, Beijing, 100084, China. qiyijun@tsinghua.edu.cn. 4. Tsinghua-Peking Center for Life Sciences, Beijing, 100084, China. qiyijun@tsinghua.edu.cn.