Structure and dynamics of a nearly steady-state subalpine forest in south-central British Columbia, Canada.

We examined the size, age, and spatial structure of trees in an old Engelmann spruce (Picea engelmannii)-subalpine fir (Abies lasiocarpa) forest based on four stem-mapped, 0.25 ha plots. Dendrochronological techniques were used on basal discs of 1,190 trees to reconstruct age and growth pattern, including dates of rapid growth increases. There were no obvious age cohorts or other evidence of past major disturbance. The abundance of both subalpine fir and spruce decreased rapidly with age, especially beyond the ages of 150 years. Very old trees were present, but rare. The best evidence from tree-ring width patterns for past disturbance was a period of release 100 years ago. However, few of the released trees grew into the canopy, which suggests a disturbance of low intensity. Patch dynamics and gap processes were not pronounced in the stand. Clumping was generally weak and only present at small spatial scales (<5 m) for live trees, and largely non-existent for dead trees; mortality was spatially random in this forest. Although spruce were sparse (5.1% of trees) in the forest relative to fir, which is consistent with predictions that fir will ultimately replace spruce in the absence of disturbance, coexistence seems more likely judging from the age structure and numbers of dead trees. In contrast to almost all spruce-fir forests studied previously, the stand we examined showed no record of major disturbances. Thus this stand falls at the limit of the range of dynamics - from disturbance-structured to near steady-state - encompassed in current thinking about forest ecosystems.