Ian G Morgan1, Regan S Ashby2. 1. Research School of Biology, Australian National University, Canberra, Australia; ian.morgan@anu.edu.au2State Key Laboratory of Ophthalmology, Zhongshan Ophthalmic Center, Sun Yatsen University, Guangzhou, China. 2. Centre for Research in Therapeutic Solutions, Health Research Institute, Faculty of Education, Science, Technology and Mathematics, University of Canberra, Canberra, Australia.