Marine biodiversity baseline for Área de Conservación Guanacaste, Costa Rica: published records.

The diversity of tropical marine organisms has not been studied as intensively as the terrestrial biota worldwide. Additionally, marine biodiversity research in the tropics lags behind other regions. The 43,000 ha Sector Marino of Área de Conservación Guanacaste (ACG, Marine Sector of Guanacaste Conservation Area), on the North Pacific coast of Costa Rica is no exception. For more than four decades, the terrestrial flora and fauna has been studied continuously. The ACG marine biodiversity was studied in the 1930's by expeditions that passed through the area, but not much until the 1990's, except for the marine turtles. In the mid 1990's the Center for Research in Marine Science and Limnology (CIMAR) of the Universidad de Costa Rica (UCR) initiated the exploration of the marine environments and organisms of ACG. In 2015, ACG, in collaboration with CIMAR, started the BioMar project whose goal is to inventory the species of the marine sector of ACG (BioMar ACG project). As a baseline, here I have compiled the published records of marine ACG species, and found that 594 marine species have been reported, representing 15.5% of the known species of the Pacific coast of Costa Rica. The most diverse groups were the crustaceans, mollusks and cnidarians comprising 71.7% of the ACG species. Some taxa, such as mangroves and fish parasites are well represented in ACG when compared to the rest of the Costa Rican coast but others appear to be greatly underrepresented, for example, red algae, polychaetes, copepods, equinoderms, and marine fishes and birds, which could be due to sampling bias. Thirty species have been originally described with specimens from ACG, and 89 species are not known from other localities on the Pacific coast of Costa Rica except ACG. Most of the sampling has been concentrated in a few localities in Sector Marino, Playa Blanca and Islas Murciélago, and in the nearby waters of Bahía Santa Elena. In an effort to fill this gap, CIMAR is collaborating with ACG and a private foundation to start an inventory of the marine organisms of the conservation area. The project will be assisted by two marine parataxonomists, and all samples will be catalogued, photographed, bar coded and voucher specimens deposited at the Museo de Zoología, UCR. All the information will be available through Internet. It is anticipated that the BioMar project will fill many of the knowledge gaps and significantly more marine species will be encountered. This project could become a viable model for marine biodiversity inventories in other Costa Rican Conservation Areas (Áreas de Conservación) and in other countries.

Introduction

Marine biodiversity studies have lagged behind terrestrial research, especially in the tropics, with a few exceptions such as Australia (Chapman 2009). Some studies in the Neotropics regarding marine biodiversity have been published, most focused on coral reef areas (Cortés et al. 2017). Several taxonomic groups are fairly well known, such as mollusks and fishes, with monographs, many papers and guides, while others are poorly known, to mention a few, microorganisms and smaller phyla. The same occurs geographically: some countries in the tropics have been relative well studied, for example, Costa Rica (Wehrtmann and Cortés 2009), while in other countries (such as Nicaragua) research and publications on marine biodiversity are scarce.

Costa Rica comprises 11 Conservation Areas (Áreas de Conservación), one of which is Área de Conservación Guanacaste (ACG) on the northwest Pacific coast of Costa Rica (Fig. 1). The ACG contains much of the last remnants of Costa Rican tropical dry forest and its terrestrial biodiversity has been and still is the subject of intensive research and restoration (Janzen and Hallwachs 2016). The ACG covers an area of 163000 hectares, 43000 of them marine, and 150 km of protected coastline (http://www.acguanacaste.ac.cr/acg/que-es-el-acg). It was declared a UNESCO World Heritage Natural Site in 1999. Compared to the terrestrial area, the marine sector (officially Sector Marino) has not been studied intensively. A new initiative, BioMar ACG (Marine Biodiversity of ACG), was started in 2015 to inventory the marine organisms of the area, and then make all the information publicly available, mainly through the Internet, but also with scientific and popular publications. This project is a 5-year collaboration between the conservation area, a private foundation and academia; all samples are being catalogued, photographed, bar coded, and vouchers deposited at the Museo de Zoología (Museum of Zoology) at the Universidad de Costa Rica (UCR).

Figure 1.

Map of the Área de Conservación Guanacaste (ACG) in the northern Pacific coast of Costa Rica with indication of the sites mentioned in the text. See Table 2 for the codes of the sites. Stars = beaches, triangle = mangrove forests, circle = bays; green = protected area; blue circles = shoals.

Table 2.

Localities of the samples reported in the Appendix 1. # spp. = number of species reported from that site. a = Protected area, b = area in the process of being officially protected, c = marine area not protected, and d = private reserve (protected area).

Code	Locality / area	Notes	# spp.
ACG a	Área de Conservación Guanacaste	Entire Conservation Area	13
BEHc	Bahía El Hachal	Bay	6
BJuc	Bahía Junquillal	Bay	5
BPGa	Bahía Potrero Grande	Bay	18
BRoc	Bajo Rojo	Shoal	2
BSEb	Bahía Santa Elena	Bay	371
BVic	Bajo Viejón	Shoal	5
CSEa	Cabo Santa Elena	Tip of PSE	23
Cuac	Cuajiniquil	Off Cuajiniquil	6
IBoc	Isla Bolaños	Island	1
IDac	Isla David	Island	7
ILCb	Isla Los Cabros	Island	1
IMua	Islas Murciélago	Archipelago	103
Juna	Junquillal	Off Junquillal	21
LDac	La Danta	Shoal	1
MCaa	Manglar de Cuajiniquil	Mangrove forest	14
MJua	Manglar de Junquillal	Mangrove forest	6
MPGa	Manglar de Potrero Grande	Mangrove forest	14
MPNa	Manglar de Playa Naranjo	Mangrove forest	19
MSaa	Manglar Salinita	Mangrove forest	14
PBla	Playa Blanca	Beach	104
PBra	Peña Bruja	Islet off PNa	2
PPGa	Playa de Potrero Grande	Beach	4
PJua	Playa Junquillal	Beach	2
PMod	Playa Mostrencal	Beach	3
PNaa	Playa Naranjo	Beach	10
PNca	Playa Nancite	Beach	16
PSEa	Península de Santa Elena	Peninsula	12
SMaa	Sector Marino ACG	Marine Sector of ACG	4

The marine sector of ACG has a high diversity of habitats, with high species richness worthy of more study (Beebe 1942, Cortés 1996–1997b). There is a well-represented suite of coastal and marine ecosystems, such as mangrove forest of variable sizes, beaches of different composition and size, bays and coves, rocky intertidal zones with several wave regimens, mud flats, rocky subtidal sites, coral reefs, rhodolith beds and deep areas – more than 50 m, plus an archipelago (Islas Murciélago), shoals, and several more isolated islands (Cortés and Wehrtmann 2009, Cortés 2016). The main PageBreaknesting site in the country of the frigate bird, , is on one of the nearby islands, Isla Bolaños, in Bahía Salinas (Alvarado-Quesada 2006). An outstanding oceanographic feature of the region is the seasonal upwelling (the Papagayo UpPageBreakPageBreakwelling) that brings deep cold, nutrient-rich and CO2-rich waters to the surface during the trade winds season (December to April-May) (McCreary et al. 1989, Alfaro and Cortés 2012, Rixen et al. 2012). Micro- and macroalgal growth increases significantly as a consequence of the upwelling (Cortés et al. 2014).

What is now ACG’s Sector Marino (Fig. 1) was first explored, samples collected, and papers published by several marine expeditions from the United States starting in the 1930’s (Cortés 2009a, Table 1). The first expedition was the Templeton Crocker Expedition of the California Academy of Sciences in 1932 aboard the SY Zaca, when they visited Bahía Murciélago and Bahía de Santa Elena (previously known as Port Parker) (Crocker 1933). In 1935, as part of the Allan Hancock Pacific Expeditions aboard the MY Velero III, biologists visited Bahía Santa Elena and Bahía Salinas (Fraser 1943a, b). The SY Zaca was again in the region in 1937–1938, but this time in an expedition of the New York Zoological Society; they collected in Bahía Santa Elena, around Islas Murciélago and around Playa Potrero Grande (Beebe 1938, 1942). These three expeditions generated a significant number of publications on ACG marine organisms (Table 1). There were no additional expeditions until 1959, when the MY Stella Polaris visited the country (Dawson and Beaudette 1959). In 1972, the RV Searcher collected samples in the region and new species of fish were described (Bussing and Lavenberg 2003). The next expedition that visited the area was the Eastern Pacific RV Alpha Helix Expedition, in 1978 organized by the Scripps Institution of Oceanography (SIO). They collected samples that are deposited at SIO, but few papers were published (Luke 1995). Chan et al. (2016) recently published on some of the barnacles collected during that expedition. The most recent expedition was the Smithsonian Tropical Research Institute RV Urracá to the northern and central Pacific coast of Costa Rica in 2005 (Vargas-Castillo 2008).

Table 1.

Historical account of marine studies at the Área de Conservación Guanacaste, Pacific coast of Costa Rica.

Years	Expedition/Project/Institutions/Individual	Taxon/System	References
1932	The Templeton Crocker Expedition of the California Academy of Sciences, aboard the SY Zaca	Algae and mollusks	93, 104, 184
1935	The Allan Hancock Pacific Expeditions, aboard the MY Velero III	Foraminifera, corals, hydroids, mollusks, crustaceans and echinoderms	5, 28, 50, 51, 52, 53, 54, 62, 63, 68, 76, 77, 78, 79, 90, 94, 95, 96, 119, 127, 148, 152, 156, 157, 158, 180, 186, 188, 203
1937–1938	Eastern Pacific Expeditions of the New York Zoological Society, aboard the SY Zaca	Mollusks, crustaceans and echinoderms	44, 45, 46, 47, 61, 80, 81, 82, 91, 105, 106, 107, 108, 109, 110, 111, 112, 113, 114, 136, 189
1959	Eastern Pacific cruise, aboard the MY Stella Polaris	Algae	56, 57, 58
1970 -present	Many individuals, for example, SE Cornelius, LG Fonseca, DA Hughes, JD Richard, DC Robinson, JR Spotila and RA Valverde	Turtle studies	1, 10, 25, 33, 34, 35, 36, 37, 48, 49, 65, 66, 67, 70, 71, 74, 84, 89, 115, 120, 121, 122, 123, 147, 150, 153, 155, 164, 165, 166, 167, 168, 169, 175, 183, 190, 192, 193, 202
1972	Central American Expedition/Janss Foundation, aboard the RV Searcher	Crustacean and fish	21
1973 -present	Several individuals and groups, e.g. DJ Pool, FE Putz and CIMAR, UCR	Mangroves	128, 170, 172, 208
1978	Caribbean-Pacific Expedition Phase VI/ Scripps Institution of Oceanography, aboard the RV Alpha Helix	Mollusks and crustaceans	27, 129, 130
1984 -present	CIMAR, UCR	Coral reefs	7, 39, 42, 124
1984 -present	CIMAR, UCR	Octocorals, corals, anemones, crustaceans, fishes, marine mammals,	11, 12, 13, 14, 15, 16, 17, 18, 19, 20, 38, 41, 59, 60, 64, 72, 141, 142, 143, 144, 145, 151, 160, 187
1991, 2013, present	Museo de Zoología, UCR	Crustaceans	102, 194, 195, 196, 197, 199, 201
1996, 1998	Fish parasite studies	Platyhelmiths and acathocephalans	24, 138, 149, 159
1996, 2002	Instituto Nacional de Biodiversidad	Mollusks	22, 23, 133, 191
2005	Benthic survey of northern and central Costa Rica/Smithsonian Tropical Research Institute, aboard the RV Urracá	Crustaceans	198
2005	Museo Nacional de Costa Rica	Birds	2
2006, 2011	Universidad Nacional, Heredia	Ascidians and cetaceans	139, 154
2014 -present	CIMAR, UCR	Beaches and rocky shores	185

Many individuals, groups of researchers or institutions have contributed to the knowledge of ACG marine biodiversity (Table 1). Elmer Y. Dawson published several papers on macroalgae of Costa Rica, including the ACG (Dawson 1960, 1961). Richard and Hughes (1972) and Cornelius (1975) published on marine turtles of the ACG, with the first observations in 1970–1971. In 1996, Marques et al. (1997) and Monks et al. (1997) collected and later described several fish parasites. Between 1996 and 2002, the Instituo Nacional de Biodiversidad collected mollusks in the ACG, and generated several papers on the opistobranchs (Valdés and Camacho-García 2004, Camacho-García et al. 2005, Camacho-García and Gosliner 2008). The CIMAR of the UCR has published papers on marine organisms and environments of Costa Rica that include the ACG: e.g., Cutler et al. (1992) on sipunculids, Moran and Dittel (1993) - crustaceans, Cortés and Guzmán (1998) - corals, Dean (2001, 2004) - polychaetes, Suárez-Morales and Morales-Ramírez (2001) - copepods, and Heard et al. (2009) - tanaidaceans. Also, new species have been described from the ACG: a crustacean (Vargas 2000), two octocorals (Breedy and Guzman 2003) and a fish (Del Moral-Flores et al. 2015). Cortés and Jiménez (2003) provided a description of the coral reefs of the ACG, while Loría-Naranjo et al. (2014) evaluated the main mangrove forests and Sibaja-Cordero et al. (2014) the beach fauna. Even so, our knowledge about the species diversity of the ACG is far from complete.

The objective of this contribution is to generate a baseline of the marine biodiversity of ACG’s Sector Marino and adjacent unprotected areas, some of which are in the process of being officially protected. This will serve as a starting point for the recently initiated BioMar ACG project (Marine Biodiversity of the Guanacaste Conservation Area). This five-year project (2015–2019), funded by the Guanacaste Dry Forest Conservation Fund, and with support from the Ministry of the Environment and Energy of the Costa Rican government and the UCR, will collect, identify and provide publicly accessible information about most of ACG’s species of marine macroorganisms and as many of the microorganisms as feasible.

Materials and methods

The study area is Sector Marino of the ACG and adjacent areas, located on the North Pacific of Costa Rica (Fig. 1, Table 2). Publications about ACG marine organisms were compiled and analyzed. A list of recorded species was created based on those publications. Later all scientific names were updated using WoRMS (World Register of Marine Species, http://www.marinespecies.org/), AlgaeBase, http://www.algaebase.org (Guiry and Guiry 2016), Encyclopedia of Life (http://eol.org/), Bryozone (http://bryozone.myspecies.info/), Integrated Digitized Biocollections (https://www.idigbio.org/), Worldwide Mollusc Species Data Base (http://www.bagniliggia.it/WMSD/Lindex_aaa.htm), SeaLifeBase (http://www.sealifebase.org/) and ZipcodeZoo (http://zipcodezoo.com/index.php/Main_Page).

The resulting list of species was compared to the remainder of the Pacific coast of Costa Rica and to available species lists from other countries in the Eastern Tropical Pacific. Knowledge gaps were identified and potential areas of future research suggested.

Results

Five hundred ninety four marine species have been reported so far for the ACG (Table 3, Appendix 1), which represents 15.5% of the known species of the Pacific coast of Costa Rica. The most diverse groups were crustaceans (193 spp.), mollusks (187 spp.) and cnidarians (46 spp.), comprising together 71.7% of the ACG’s marine species. These three groups represent 23.9%, 18.2% and 26.7%, respectively of the known species of the Pacific coast of the country (Table 3). Some groups are well represented in the ACG when compared to the rest of the coast (e.g., species of mangroves and fish parasites), while others are greatly underrepresented. For example, red algae, polychaetes, copepods, equinoderms, and marine fishes and birds are poorly represented in the published reports (Table 3). Other groups of organisms have been observed and identified (e.g., various species of sponges, flat worms, ophiuroids, and ascidian) but there are no published records of these species (Table 4). Other taxa (such as diatoms, nemerteans and appendicularians) undoubtedly inhabit the study area but have not been observed or collected yet (Table 4).

Table 3.

Number of marine species reported from Área de Conservación Guanacaste (complete list of species in the Appendix 1), Pacific coast of Costa Rica (see Cortés 2012, plus references indicated as superindex) (species reported only for Isla del Coco were excluded); percentage of the species of the Pacific reported form ACG, and species only found in ACG. n.k. = not known.

TAXA	Species from ACG	Species from Pacific Costa Rica	% of species of the Pacific	Species only at ACG
Bacteria	15	>17 [103, 183]	88.2	15
Cyanobacteria	4	28	14.3	2
Chlorophyta	4	44 [73]	9.1	2
Phaeophyceae	6	26 [73]	23.1	1
Rhodophyta	15	146 [73]	10.3	9
Mangroves	7	8	87.5	0
Foraminifera	24	76	31.6	12
Cnidaria	46	172	26.7	2
Anthozoa	35	59	59.3	2
Hydrozoa	11	108	10.2	0
Platyhelminthes	7	38 [40, 178]	18.4	7
Trematoda	4	20 [40178, 182]	20.0	4
Cestoda	3	12 [40, 178]	25.0	3
Acanthocephala	1	1 [149]	100	0
Mollusca	187	1025	18.2	0
Gastropoda	85	631	13.5	0
Bivalvia	102	362	28.2	0
Sipuncula	3	15	20.0	0
Annelida	24	313	7.7	11
Nemertea	1	Several species	n.k.	n.k.
Crustacea	193	807	23.9	13
Amphipoda	13	106	12.3	8
Cumacea	1	19 [161]	5.3	1
Decapoda	162	409	39.6	1
Mysida	1	5	20.0	0
Stomatopoda	10	27	37.0	0
Tanaidacea	1	5	20.0	1
Copepoda	1	163	0.61	1
Cirripedia	4	36	11.1	1
Bryozoa	9	39	23.1	8
Echinodermata	15	105	14.3	7
Asteroidea	1	12	8.3	0
Echinoidea	1	28	3.6	0
Holothuroidea	13	28	46.4	7
Chordata	33	961	3.4	0
Ascidiacea	5	14	35.7	0
Cephalochordata	1	2	50	0
Elasmobranchii	3	68	4.4	0
Actinopterygii	11	774	1.4	0
Reptilia	4	5	80.0	0
Aves	2	76	2.6	0
Mammalia	7	22	31.8	0
TOTAL	594	3821+	15.5	89

Table 4.

Taxa reported from other sites of Pacific Costa Rica (see Cortés 2012, plus references indicated as superindex), but not from Área de Conservación Guanacaste. n.k. = not known; Present = have been observed or collected but there are no publications; Probably = there is a high probability that they are present but have not been observed yet.

Taxonomic group	Number of species reported	ACG
Diatoms	174 [131,132,200]	Present
Dinoflagellates	102	Present
Marine fungi	5 genera	n.k.
Seagrasses	2	n.k.
Porifera	62	Present
Pennatulaceans	4	Present
Scyphozoans	10	Present
Polyplacophorans	24	Present
Cephalopods	20	Present
Echiurians	1	Present
Monogeneans	10 [40]	Probably
Nemerteans	Several species	Probably
Kinorhynchans	2	n.k.
Euphausiids	20	Present
Isopods	37	Present
Branchiopods	1	n.k.
Ostracods	2	Probably
Pycnogonids	10	Probably
Marine insects	9	Probably
Chaetognaths	27	Present
Brachiopods	8	n.k.
Phoronids	1	n.k.
Crinoids	2	n.k.
Ophiuroids	54	Present
Appendicularians	10	Probably
Thaliaceans	4	Probably
Turtle parasites	34	Present

Over 85% of the species reported are also found in other areas of the coast of Costa Rica and in the Eastern Tropical Pacific; however, most areas, including the ACG, have not been intensively collected, and the same common species are found repeatedly by collecting expeditions. Thirty new species have been described from specimens collected in the ACG: one foraminiferan, one echinoderm, two octocorals, three parasitic flatworms, four fishes, eight crustaceans and 11 mollusks (Appendix 1). Eighty-nine species are currently known only from the ACG along the Pacific coast of Costa Rica (Table 3, Appendix 1).

Most of the sampling has been concentrated in a few localities of the marine area of the ACG and those sites therefore have the highest number of reported species. For example, Bahía Santa Elena (371 spp.), Playa Blanca (104 spp.) and in some of the PageBreakPageBreakIslas Murciélago (103 spp.) seem very species-rich (Table 2, Appendix 1). Other areas within ACG have not been sampled at all, for example the northern shore of the Santa Elena Peninsula or some of the Islas Murciélago. The soft bottom substrate has not been sampled thoroughly nor most of the rocky intertidal zones.

Discussion

Compared to other areas on the Pacific of Costa Rica, the ACG has fewer known marine species (594 spp.) than does Golfo Dulce (1028 spp.: Morales-Ramírez 2011) or Isla del Coco (1688 spp.: Cortés 2012), but about the same as what is currently known for Bahía Culebra (577 spp: Cortés et al. 2012). But that number will defiPageBreaknitely increase as more taxa, other sites and environments within the ACG are inventoried.

Cortés et al. (2017) synthesized the knowledge of marine biodiversity of the Eastern Tropical Pacific, mainly from coral reefs, where most studies have been done. For example, 857 marine species have been reported for Clipperton Atoll, France, (Charpy 2009, Payri et al. 2009, Fourriére et al. 2014), 968 spp. for El Salvador (Barraza 2000, 2014a, b), 2157 spp. for the coast of Oaxaca, México (Bastida-Zavala et al. 2013), 3536 spp. for the Galápagos Islands (Bustamante et al. 2002, Hickman 2009), 3838 spp. for the Pacific coast of Costa Rica (Table 3, this paper), and 5740 spp. for the entire Gulf of California, México (Aburto-Oropeza and Balart 2001, Reyes-Bonilla et al. 2012). In other countries, for example, Panamá and Colombia, there are detailed inventories of some higher taxa, but not a compilation of all macrotaxa (Cortés et al. 2017). None of these inventories attempted to include the microorganisms.

There are large differences in the numbers of species among different sites in the Eastern Tropical Pacific and these differences could be due to several causes. First, the number, diversity and depth of research efforts influence the extent of the knowledge of the marine biodiversity of a region. Second, the extent of each region will also have an effect on species diversity, because larger areas will probably include more habitats and environments, and thus species. The ACG marine area comprises 430 km2, while the Gulf of California has about 160000 km2. Third, some sites may differ in species richness and diversity because of differences in geomorphology, oceanography, geological history and biogeography. Fourth, natural disturbances such as warming or cooling events can have a long-term impact on local biodiversity.

Knowing and documenting which species occurs where is a critical first step in understanding and conserving the biodiversity of a particular area. As outlined in Tables 3 and 4, there are important gaps in our knowledge in taxonomy and geographic distribution of marine organisms in the ACG. Much more work is needed to have an even approximately complete inventory, understand the ecological role of the species, their habitats, population structure, and distribution. Researchers of the BioMar-ACG project will fill many of these gaps, and together with other researchers from Costa Rica and elsewhere, the understanding of the marine biodiversity of the ACG will increase greatly. The BioMar project incorporates several innovative aspects, including marine parataxonomists, DNA barcoding of all organims and fast accessibility of the information. This project could serve as a viable model for marine biodiversity inventory in other Costa Rican conservation areas and in other countries.

Marine species reported from Área de Conservación Guanacaste (ACG). Species in bold type reported only for the ACG in Costa Rica (in the case of some have been reported in people but not in marine organisms). Localities as in Figure 1 and Table 1. Localities in bold type = a) Type locality, b) Paratype locality and c) Neotype specimen. References numbered as in the reference list.

	Species	Locality	References
Phylum ACTINOBACTERIA, Class Actinobacteria, Order Actinomycetales, Family Corynebacteriaceae	Corynebacterium spp.	PNc	183
Phylum FIRMICUTES, Class Bacilli, Order Bacillales, Family Bacillaceae	Bacillus spp.	PNc	183
Order Lactobacillales, Family Enterococcaceae	Enterococcus faecalis (Orla-Jensen, 1919) Schleifer & Kilpper-Bälz, 1984	PNc	183
Order Lactobacillales, Family Lactobacillaceae	Lactobacillus spp.	PNc	183
Order Bacillales, Family Staphylococcaceae	Staphylococcus aureus Rosenbach, 1884	PNc	183
Phylum PROTEOBACTERIA, Class Betaproteobacyeria, Order Burkholderiales, Familia Alcaligenaceae	Alcaligenes faecalis Castellani & Chalmers, 1919	PNc	183
Class Gammaproteobacteria, Order Aeromonadales, Family Aeromonadaceae	Aeromonas spp.	PNc	183
Order Enterobacteriales, Family Enterobacteriaceae	Citrobacter freundi (Braak 1928) Werkman & Gillen, 1932	PNc	183
	Pantoea agglomerans (Ewing & Fife, 1972) as Enterobacter agglomerans	PNc	183
	Escherichia coli Castellani & Chalmers, 1919	PNc	183
	Proteus mirabilis Hauser, 1885	PNc	183
	Salmonella spp.	PNc	183
Order Pseudomonadales, Family Moraxellaceae	Acinetobacter spp.	PNc	183
Order Pseudomonadales, Family Pseudomondaceae	Pseudomonas aeruginosa (Schroeter, 1872) Migula, 1900	PNc	183
	Pseudomonas spp.	PNc	183
Phylum CYANOBACTERIA, Class Cyanophyceae, Order Chroococcales, Family Dermocarpellaceae	Cyanocystis violacea (P.L. Crouan & H.M. Crouan) Komárek & Anagnostidis, 1986 as Dermocarpa violacea	BSE	188
Order Chroococcales, Family Entophysalidaceae	Entophysalis granulosa Kützing, 1843	BSE	188
Order Oscillatoriales, Family Oscillatoriaceae	Lyngbya semiplena J. Agardh ex Gomont, 1892	BSE	188
Class Cyanophyceae	One species	BEH	185
Phylum CHLOROPHYTA, Class Ulvophyceae, Order Cladophorales, Family Cladophoraceae	Cladophora lehmanniana (Lindenberg) Kützing, 1843 as Cladophora utriculosa	BPG	56, 57
	Cladophora sp.	BSE	128
Order Ulvales, Family Ulvaceae	Ulva flexuosa Wulfen, 1803 as Enteromorpha flexuosa and as Enteromorpha lingulata	BSE	57, 188
	Ulva lactuca Linnaeus, 1753	BSE	188
	Ulva prolifera O.F. Müller 1778 as Enteromorpha prolifera var. flexuosa	BSE	57, 128
	Ulva sp.	Jun	185
Phylum OCHROPHYTA, Class Phaeophyceae Order Dictyotales, Family Dictyotaceae	Padina sp.	BPG	58
Order Ectocarpales, Family Scytosiphonaceae	Colpomenia durvillei (Bory de Saint-Vincent) M.E. Ramírez, 1991 as Colpomenia phaeodactyla	BSE	207
	Colpomenia ramosa W.R. Taylor, 1945	BSE	57, 188
	Colpomenia sinuosa (Mertens ex Roth) Derbès & Solier, 1851	BSE	57
	Rosenvingea orientalis (J. Agardh) Børgesen, 1914	BPG	56, 57
Order Fucales, Family Sargassaceae	Sargassum liebmannii J. Agardh 1847	BSE	184
	Sargassum sp.	BPG	58
Phylum RHODOPHYTA, Class Bangiophyceae, Order Bangiales, Family Bangiaceae	Bangia fuscopurpurea (Dillwyn) Lyngbye, 1819	BSE	188
	Pyropia thuretii (Setchell & E.Y.Dawson) J.E. Sutherland, L.E. Aguilar Rosas & R. Aguilar Rosas, 2011	BSE	57
Class Compsopogonophyceae, Order Erythropeltidales, Family Erythrotrichiaceaea	Smithora naiadum (C.L. Anderson) Hollenberg, 1959 as Porphyra naiadum	BSE	188
Class Florideophyceae, Order Acrochaetiaceae, Family Acrochaetiaceae	Acrochaetium arcuatum (K.M. Drew) C.K. Tseng, 1945 as Acrochaetium penetrale	BSE	57, 188
Order Ceramiales, Family Rhodomelaceae	Bostrychia sp.	Jun	185
	Chondria dangeardii E.Y. Dawson, 1954 as Chondria platyclada	BPG	58
Order Ceramiales, Family Ceramiaceae	Ceramium avalonae E.Y. Dawson, 1949	BPG	57
	Ceramium personatum Setchell & N.L. Gardner, 1930	BSE	57, 188
Order Ceramiales, Family Rhodomelaceae	Neosiphonia beaudettei (Hollenberg) M.-S. Kim & I.A. Abbott, 2006 as Polysiphonia beaudettei	BPG	57
	Polysiphonia bifurcata Hollenberg in W.R. Taylor, 1945	PBl	57
Order Gigartinales, Family Dicranemataceae	Dicranema rosaliae Setchell & Gardner, 1924	BSE	57
Order Gracilariales, Family Gracilariaceae	Gracilaria symmetrica Dawson, 1949	BPG	58
	Gracilaria sp.	BSE	57
	Gracilariopsis sp.	BPG	58
Order Hildebrandiales, Family Hildebrandiaceae	Hildenbrandia rubra (Sommerfelt) Meneghini, 1841 as Hildenbrandia prototypus	BSE	188
Order Rhodymeniales, Family Rhodymeniaceae	Botryocladia beaudettei E.Y. Dawson, 1960	BPG	57
Division MAGNOLIOPHYTA, Class Magnoliopsida, Order Ericales, Family Tetrameristaceae	Pelliciera rhizophoreae Triana & Planchon, 1862	BSE	128
	Pelliciera rhizophoreae Triana & Planchon, 1862	MPG	32, 128, 208
	Pelliciera rhizophoreae Triana & Planchon, 1862	MPN	176
Order Lamiales, Family Acanthaceae	Avicennia bicolor Standley, 1923	MJu	32
	Avicennia bicolor Standley, 1923	MCa	32
	Avicennia bicolor Standley, 1923	MSa	32
	Avicennia bicolor Standley, 1923	BSE	128
	Avicennia bicolor Standley, 1923	MPG	32, 208
	Avicennia bicolor Standley, 1923	MPN	32, 208
	Avicennia germinans Linnaeus, 1764	MJu	32
	Avicennia germinans Linnaeus, 1764	MCa	32
	Avicennia germinans Linnaeus, 1764	MSa	32
	Avicennia germinans Linnaeus, 1764	BSE	128
	Avicennia germinans Linnaeus, 1764	MPG	32, 128, 208
	Avicennia germinans Linnaeus, 1764 as Avicennia tonduzii in reference 172	MPN	32, 116, 172, 206
	Avicennia spp.	MPN	176
Order Myrtales, Family Combretaceae	Conocarpus erectus Linnaeus, 1753	MJu	32
	Conocarpus erectus Linnaeus, 1753	MCa	32
	Conocarpus erectus Linnaeus, 1753	MSa	32
	Conocarpus erectus Linnaeus, 1753 as Conocarpus erecta in reference 176	MPN	172, 176, 206
	Laguncularia racemosa (L.) Gärtner, 1807	MJu	32
	Laguncularia racemosa (L.) Gärtner, 1807	MSa	32
	Laguncularia racemosa (L.) Gärtner, 1807	BSE	128
	Laguncularia racemosa (L.) Gärtner, 1807	MPG	128, 208
	Laguncularia racemosa (L.) Gärtner, 1807	MPN	172, 176, 206
Order Rhizophorales, Family Rhizophoraceae	Rhizophora mangle Linnaeus, 1753	MJu	32
	Rhizophora mangle Linnaeus, 1753	MCa	32
	Rhizophora mangle Linnaeus, 1753	MSa	32
	Rhizophora mangle Linnaeus, 1753	BSE	128
	Rhizophora mangle Linnaeus, 1753	MPG	32, 128
	Rhizophora mangle Linnaeus, 1753	MPN	32, 170, 208
	Rhizophora racemosa Meyer, 1818	MJu	32
	Rhizophora racemosa Meyer, 1818	MCa	32
	Rhizophora racemosa Meyer, 1818	MSa	32
	Rhizophora racemosa Meyer, 1818	BSE	128
	Rhizophora racemosa Meyer, 1818	MPG	32, 128, 208
	Rhizophora racemosa Meyer, 1818	MPN	32, 206, 208
	Rhizophora spp.	MPN	176
Phylum FORAMINIFERA, Class Globothalamea, Order Lituolida, Family Discamminidae	Ammoscalaria compressa (Cushman & McCulloch, 1939) as Ammofrondicularia compressa	PBl	50
Order Lituolida, Family Haplophragmoididae	Haplophragmoides planissima Cushman, 1927 as Haplophragmoides planissimum	BSE	50
	Labrospira columbiensis (Cushman, 1925) as Haplophragmoides columbiense	PBl	50
	Labrospira columbiensis (Cushman, 1925) as Haplophragmoides columbiense	BSE	50
Order Lituolida, Family Lituolidae	Eratidus foliaceus (Brady, 1881) as Ammobaculites foliaceus	PBl	50
Order Lituolida, Family Reophacidae	Reophax curtus Cushman, 1920	PBl	50
	Reophax scorpiurus de Montfort, 1808	BSE	50
Order Lituolida, Family Nouriidae	Nouria polymorphinoides Heron-Allen & Earland, 1914	BSE	50
	Nouria polymorphinoides Heron-Allen & Earland, 1914	PBl	50
Order Lituolida, Family Remaneicidae	Remaneica kellettae (Thalmann, 1932) as Trochammina kellettae	BSE	50
Order Lituolida, Family Trochamminidae	Portatrochammina pacifica (Cushman, 1925) as Trochammina pacifica	BSE	50
	Portatrochammina pacifica (Cushman, 1925) as Trochammina pacifica	PBl	50
Order Rotaliida, Family Bolivinitidae	Bolivina pygmaea (Brady, 1881)	PBl	52
	Loxostomina limbata (Brady, 1881) as Loxostoma limbatum	BSE	52
Order Rotaliida, Family Buliminellidae	Buliminella elegantissima (d’Orbigny, 1839)	PBl	53
Order Rotaliida, Family Elphidiidae	Elphidium seymourense McCulloch, 1977 as Elphidium crispum var. subcrispum	PBL	51
Order Rotaliida, Family Heterohelicidae	Bifarina pacifica Cushman & McCulloch, 1942	BSEa	52
Order Textulariida, Family Textulariidae	Sahulia conica (d’Orbigny, 1839) as Textularia conica	BSE	127
	Textularia calva Lalicker, 1940	BSE	127
	Textularia candeiana d’Orbigny, 1839	BSE	127
	Textularia candeiana d’Orbigny, 1839	PBl	127
	Textularia corrugata Herron-Allen & Earland, 1915	PBl	127
	Textularia foliacea Heron-Allen & Earland, 1915	PBl	127
	Textularia panamensis Cushman, 1918	PBl	127
	Textularia secasensis Lalicker & McCulloch, 1940	BSE	127
	Textularia secasensis Lalicker & McCulloch, 1940	PBl	127
Class Tubothalamea, Order Spirillinida, Family Ammodiscidae	Glomospira gordialis (Jones & Parker, 1860)	BSE	50
Class incerta sedis, Order Lagenida, Family Lagenidae	Lagena amphora Reuss, 1863	PBl	54
	Lagena striata (d’Orbigny, 1839)	PBl	54
Phylum CNIDARIA, Class Anthozoa, Orden Antipatharia, Family Antipathidae	Antipathes sp.	IMu	7
Order Alcyonacea, Family Gorgoniidae	Eugorgia aurantiaca (Horn, 1860)	IMu	38
	Eugorgia daniana Verrill, 1868	BSE	11
	Eugorgia rubens Verrill, 1868	IMu	11, 18
	Leptogorgia alba (Duchassaing & Michelotti, 1864)	IBo	11, 15, 38
	Leptogorgia alba (Duchassaing & Michelotti, 1864)	BSE	11, 15, 38
	Leptogorgia alba (Duchassaing & Michelotti, 1864)	PBl	7
	Leptogorgia alba (Duchassaing & Michelotti, 1864)	IMu	7, 11, 15, 38
	Leptogorgia cofrini Breedy & Guzman, 2005	IMub	14
Order Alcyonacea, Family Gorgoniidae	Leptogorgia cuspidata Verrill, 1865	IMu	15
	Leptogorgia regis Hickson, 1928	BSE	11, 15
	Leptogorgia regis Hickson, 1928	IMu	11, 15
	Pacifigorgia firma Breedy & Guzman, 2003	BSE	11
	Pacifigorgia irene Bayer, 1951	IMu	7, 11, 12, 13
	Pacifigorgia rubicunda Breedy & Guzman, 2003	IMub	11, 13
	Pacifigorgia senta Breedy & Guzman, 2003	Cuaa	11, 13
	Pacifigorgia senta Breedy & Guzman, 2003	PBl	7
	Pacifigorgia senta Breedy & Guzman, 2003	IMub	7, 11, 13
	Pacifigorgia stenobrochis (Valenciennes, 1846)	IMu	11, 13
	Pacifigorgia tupperi Breedy & Guzman, 2003	IMua	11, 13
Order Alcyonacea, Family Plexauridae	Muricea austera Verrill, 1869	PBr	17
	Muricea plantaginea (Valenciennes, 1846)	BSE	17
	Muricea squarrosa Verrill, 1869	LDa	16
	Muricea sp.	ACG	38
Order Actiniaria, Family Nemanthidae	Nemanthus californicus Carlgren, 1940	IMu	72
Order Scleractinia, Family Agariciidae	Gardineroseris planulata (Dana, 1846)	IMu	7, 42, 124
	Pavona clavus (Dana, 1846)	PSE	38, 39
	Pavona clavus (Dana, 1846)	PBl	7
	Pavona clavus (Dana, 1846)	IMu	7, 41, 42, 124
	Pavona clavus (Dana, 1846)	ACG	39, 42
	Pavona gigantea Verrill, 1869	PSE	38, 39, 43
	Pavona gigantea Verrill, 1869	PBl	7
	Pavona gigantea Verrill, 1869	IMu	7, 42, 124
	Pavona gigantea Verrill, 1869	ACG	39, 41, 42
	Pavona maldivensis (Gardiner, 1905)	ACG	41
	Pavona varians Verrill, 1864	IMu	7, 41
	Pavona varians Verrill, 1864	ACG	41
Order Scleractinia, Family Dendrophylliidae	Cladopsammia eguchii (Wells, 1982)	IMu	38
	Tubastraea coccinea Lesson, 1829 as Tubastrea tenuilamellosa	PBl	68
	Tubastraea coccinea Lesson, 1829	IDa	154
	Tubastraea coccinea Lesson, 1829	BVi	154
	Tubastraea coccinea Lesson, 1829 as Tubastrea coccinea	PSE	38
	Tubastraea coccinea Lesson, 1829 as Tubastrea coccinea	IMu	7, 42, 124
	Tubastraea coccinea Lesson, 1829 as Tubastrea coccinea	ACG	39, 42
Order Scleractinia, Family Pocilloporidae	Pocillopora damicornis (Linnaeus, 1758)	PSE	43
	Pocillopora damicornis (Linnaeus, 1758)	IMu	7, 41, 42, 124
	Pocillopora damicornis (Linnaeus, 1758)	ACG	39, 41
	Pocillopora elegans Dana, 1846	BSE	7
	Pocillopora elegans Dana, 1846	PBl	7
	Pocillopora elegans Dana, 1846	IMu	41, 42, 124
	Pocillopora elegans Dana, 1846	ACG	39
Order Scleractinia, Family Pocilloporidae	Pocillopora eydouxi Milne Edwards & Haime, 1860	ILC	7, 39
	Pocillopora eydouxi Milne Edwards & Haime, 1860	PSE	41, 43
	Pocillopora eydouxi Milne Edwards & Haime, 1860	IMu	7, 41, 42, 124
	Pocillopora eydouxi Milne Edwards & Haime, 1860	ACG	39
	Pocillopora inflata Glynn, 1999, but see Paz-García et al. 2015	BSE	7
	Pocillopora inflata Glynn, 1999	IMu	42, 86
	Pocillopora meandrina Dana, 1846	BSE	38
	Pocillopora meandrina Dana, 1846	PSE	41
	Pocillopora meandrina Dana, 1846	IMu	7, 41
	Pocillopora meandrina Dana, 1846	ACG	39
Order Scleractinia, Family Poritidae	Porites lobata Dana, 1846	PBl	7
	Porites lobata Dana, 1846	IMu	7, 42, 124
	Porites lobata Dana, 1846	ACG	39
	Porites panamensis Verrill, 1866	IMu	42, 124
	Porites panamensis Verrill, 1866	ACG	39, 41, 42
Order Scleractinia, Family Rhizangiidae	Oulangia bradleyi Verrill, 1866	ACG	39
Order Scleractinia, Family Siderastreidae	Psammocora stellata (Verrill, 1866)	BSE	7
	Psammocora stellata (Verrill, 1866)	PSE	43
	Psammocora stellata (Verrill, 1866)	IMu	7, 41, 42
	Psammocora profundacella Gardiner, 1898 as Psammocora superficialis	BSE	7
Class Hydrozoa, Order Anthoathecata Family Bougainvilliidae	Garveia gracilis (Clark, 1876) as Bimeria gracilis	BSE	38, 76, 77
Order Leptothecata, Family Aglaopheniidae	Aglaophenia trifida Agassiz, 1862 as Aglaophenia rigida	PBl	38, 76
Order Leptothecata, Family Campanulariidae	Clytia fascicularis Fraser, 1938	PBl	38, 77
	Clytia gracilis (Sars, 1850) as Clytia cylindrica and as Gonothyraea gracilis	BSE	38, 77
	Clytia universitatis Torrey, 1904	BSE	38, 77
Order Leptothecata, Family Haleciidae	Halecium washingtoni Nutting, 1901	BSE	38, 77
Order Leptothecata, Family Plumularidae	Plumularia micronema Fraser, 1938 as Plummularia micronema	BSE	38, 76
	Plumularia micronema Fraser, 1938 as Plummularia micronema	PBl	38, 76
Order Leptothecata, Family Sertulariidae	Amphisbetia furcata (Trask, 1857) as Sertularia furcata	CSE	76
	Dynamena crisioides Lamouroux, 1824 as Thuiaria tubuliformis	BSE	38, 76, 77
	Dynamena crisioides Lamouroux, 1824 as Thuiaria tubuliformis	PBl	38, 77
	Sertularia distans (Lamouroux, 1816) as Sertularia stookeyi	CSE	76
Orden Siphonophorae, Familia Physaliidae	Physalia physalis (Linnaeus, 1758) as Physalia physalia	IMu	38
Phylum PLATYHELMINTHES, Class Trematoda Order Plagiorchiida, Family Acanthocolpidae	Stephanostomum casum (Linton, 1910)	Jun	40, 159
Order Plagiorchiida, Family Lecithasteridae	Trifoliovarium sp. Yamaguti, 1940 as Pseudolecithaster	Cua	159, 178
Order Plagiorchiida, Family Hemiuridae	Theletrum lamothei Pérez-Ponce de León, León-Règagnon & Monks, 1998	Cuaa	159, 178
Order Plagiorchiida, Family Lepocreadiidae	Hypocreadium myohelicatum Bravo-Hollis & Manter, 1957	Jun	159
Class Cestoda, Order Onchoproteocephalidea, Family Onchobothriidae	Acanthobothrium franus Marques, Centritto & Stewart, 1997	Cuaa	138, 178
	Acanthobothrium inbiorium Marques, Centritto & Stewart, 1997	Juna	138, 178
Order Trypanorhyncha, Family Pterobothriidae	Pterobothrioides carvajali Campbell & Beveridge, 1997	Cuab	24, 178
Phylum ACANTHOCEPHALA, Class Palaeacanthocephala, Order Echinorhynchida Family Illiosentidae	Koronacatha pectinaria (Van Cleave, 1940)	Jun	149, 178
Phylum MOLLUSCA, Class Gastropoda, Subclass Heterobranchia, Order Unassigned, Family Acteonidae	Acteon traskii Stearns, 1897	BSE	191
Infraclass Opisthobranchia, Order Cephalaspidea, Family Acteocinidae	Acteocina carinata (Carpenter, 1857)	IDa	191
	Acteocina carinata (Carpenter, 1857)	IDa	191
	Acteocina carinata (Carpenter, 1857)	BSE	191
	Acteocina carinata (Carpenter, 1857)	PBl	191
	Acteocina carinata (Carpenter, 1857)	BPG	191
	Acteocina infrequens (C. B. Adams, 1852)	BJu	191
	Acteocina infrequens (C. B. Adams, 1852)	IDa	191
	Acteocina infrequens (C. B. Adams, 1852)	BSE	191
	Acteocina infrequens (C. B. Adams, 1852)	BPG	191
	Acteocina sp.	BEH	191
Order Cephalaspidea, Family Aglajidae	Navanax aenigmaticus (Bergh, 1894)	BJu	191
Order Cephalaspidea, Family Bullidae	Bulla punctulata A. Adams in Sowerby, 1850	Jun	134
Order Cephalaspidea, Family Cylichnidae	Cylichna atahualpa (Dall, 1908)	BSE	191
	Cylichnella tabogaensis (Strong & Hertlein, 1939)	BJu	191
	Cylichnella tabogaensis (Strong & Hertlein, 1939)	IDa	191
	Cylichnella tabogaensis (Strong & Hertlein, 1939)	BSE	191
	Cylichnella tabogaensis (Strong & Hertlein, 1939)	PBl	191
Order Cephalaspidea, Family Haminoeidae	Atys defuncta (Baker & Hanna, 1927)	IDa	191
	Atys defuncta (Baker & Hanna, 1927)	BSE	191
	Atys defuncta (Baker & Hanna, 1927)	IMu	191
	Atys defuncta (Baker & Hanna, 1927)	CSE	191
	Atys defuncta (Baker & Hanna, 1927)	BPG	191
	Atys exaratus (Carpenter, 1857) as Atys exarata	BJu	191
	Atys exaratus (Carpenter, 1857) as Atys exarata	IDA	191
	Atys exaratus (Carpenter, 1857) as Atys exarata	PBl	191
	Atys exaratus (Carpenter, 1857) as Atys exarata	IMu	191
Order Cephalaspidea, Family Retusidae	Retusa paziana Dall, 1919	IDa	191
	Retusa paziana Dall, 1919	BSE	191
	Retusa sp.	PNa	191
Order Cephalaspidea, Family Rhizoridae	Volvulella cylindrica (Carpenter, 1864)	PJu	191
	Volvulella cylindrica (Carpenter, 1864)	BSE	191
	Volvulella cylindrica (Carpenter, 1864)	PBl	191
	Volvulella cylindrica (Carpenter, 1864)	IMu	191
	Volvulella cylindrica (Carpenter, 1864)	BPG	191
	Volvulella cylindrica (Carpenter, 1864)	PNa	191
Order Nudibranchia, Family Discodorididae	Atagema notacristata Camacho-García & Gosliner 2008	IMu	22
Orden Nudibranchia, Family Fionidae	Fiona pinnata (Eschscholtz, 1831)	BSE	130
Orden Nudibranchia, Family Polyceridae	Limacia janssi (Bertsch & Ferreira, 1974) as Laila janssi	BSEa, b	8
Order Sacoglossa, Family Plakobranchiadae	Elysia sp.	IMu	23
Infraclass Pulmonata, Order Unassigned, Family Siphonariidae	Siphonaria gigas Sowerby, 1825	CSE	130
	Siphonaria gigas Sowerby, 1825	Jun	185
Infraclass Unassigned, Family Pyramidellidae	Eulimastoma dotella (Dall & Bartsch, 1909) as Odostomia (Telloda) subdotella	BSE	114
	Odostomia costaricensis Hertlein & Strong, 1951 as Odostomia (Chrysallida) costaricensis	BSE	114
	Odostomia nicoyana Hertlein & Strong, 1951 as Odostomia (Menestho) nicoyana	BSE	114
	Odostomia woodhridgei Hertlein & Strong, 1951 as Odostomia (Chrysallida) woodbridgei	BSE	114
	Odostomia (Besla) caneloensis Hertlein & Strong, 1951	BSEa	114
	Turbonilla amiriana Hertlein & Strong, 1951 as Turbonilla (Pyrgiscus) amiriana	BSE	114
	Turbonilla ayamana Hertlein & Strong, 1951 as Turbonilla (Pyrgiscus) ayamana	BSE	114
	Turbonilla biolleyi Hertlein & Strong, 1951 as Turbonilla (Pyrgiscus) biolleyi	BSE	114
	Turbonilla ekidana Hertlein & Strong, 1951 as Turbonilla (Pyrgiscus) ekidana	BSE	114
	Turbonilla guanacastensis Hertlein & Strong, 1951 as Turbonilla (Pyrgiscus) guanacastensis	BSE	114
	Turbonilla nicoyana Hertlein & Strong, 1951 as Turbonilla (Pyrgiscus) nicoyana	BSE	114
	Turbonilla portoparkerensis Hertlein & Strong, 1951 as Turbonilla (Ptycheulimella) portoparkensis	BSE	114
	Turbonilla sulacana Hertlein & Strong, 1951 as Turbonilla (Pyrgiscus) sulacana	BSE	114
	Turbonilla templetonis Hertlein & Strong, 1951 as Turbonilla (Pyrgiscus) templetonis	BSE	114
	Turbonilla utuana Hertlein & Strong, 1951 as Turbonilla (Pyrgisculus) utuana	BSE	114
	Turbonilla zacae Hertlein & Strong, 1951 as Turbonilla (Pyrgiscus) zacae	BSE	114
Subclass Caenogastropoda, Order Littorinimorpha, Family Cypraeidae	Pseudozonaria arabicula (Lamarck, 1810) as Zonaria (Zonaria) arabicula	IMu	26
Order Littorinimorpha, Family Ficidae	Ficus ventricosa (G. B. Sowerby I, 1825)	IMu	130
Order Littorinimorpha, Family Littorinidae	Echinolittorina atrata (C. B. Adams, 1852) as Nodilittorina atrata	PSE	174
	Echinolittorina fuscolineata (Reid, 2002) as Nodilittorina fuscolineata	IMu	174
	Echinolittorina modesta (Philippi, 1846) as Littorina modesta	IMu	130
	Echinolittorina peruviana (Lamarck, 1822) as Littoraria zebra	MCa	32
	Echinolittorina peruviana (Lamarck, 1822) as Littoraria zebra	MSa	32
	Echinolittorina peruviana (Lamarck, 1822) as Littoraria (Littoraria) zebra	BSE	173
	Echinolittorina peruviana (Lamarck, 1822) as Littoraria zebra	PPG	32, 208
	Echinolittorina peruviana (Lamarck, 1822) as Littoraria zebra	MPN	32, 208
Order Littorinimorpha, Family Personidae	Distorsio decussata (Valenciennes, 1832)	IMu	130
Order Littorinimorpha, Family Rissoinidae	Zebinella alarconi (Hertlein & Strong, 1951) as Rissoina alarconi	BSE	114
Order Littorinimorpha, Family Strombidae	Persististrombus granulatus (Swainson, 1822) as Strombus granulatus Swainson, 1822	IMu	130
Order Littorinimorpha, Family Tornidae	Anticlimax willetti Hertlein & Strong, 1951 as Anticlimax (Subclimax) willetti	BSEa	114
	Aorotrema humboldti (Hertlein & Strong, 1951) as Cyclostremiscus humboldti	BSE	114
	Teinostoma herbertianum Hertlein & Strong, 1951 as Teinostoma herbertiana	BSE	114
	Teinostoma zacae Hertlein & Strong, 1951	BSE	114
Order Neogastropoda, Family Columbellidae	Anachis fluctuata (G. B. Sowerby I, 1832) as Anachis (Parvanachis) fluctuata	IMu	130
	Anachis pardalis (Hinds, 1843) as Anachis (Parvanachis) carmen	IMu	130
	Clavistrombina clavulus (G. B. Sowerby I, 1834)	BSE	125
	Cosmioconcha rehderi (Hertlein & Strong, 1951) as Anachis rehderi	BSEa	114
	Cotonopsis hirundo (Gaskoin, 1852) as Cotonopsis (Turrina) hirundo	BSE	125
	Mazatlania fulgurata (Philippi, 1846) as Terebra moolenbeeki	PNaa	4, 9
	Sincola dorsata (G. B. Sowerby I, 1832) as Sincola (Dorsina) dorsata	BSE	125
	Strombina elegans (G. B. Sowerby I, 1832) as Strombina (Spiralta) elegans	BSE	125
	Sincola gibberula (G. B. Sowerby I, 1832) as Sincola (Dorsina) gibberula	BSE	125
	Strombina elegans (G. B. Sowerby I, 1832) as Strombina (Spiralta) elegans	PBl	125
	Strombina elegans (G. B. Sowerby I, 1832) as Strombina (Spiralta) elegans	IMu	125
	Strombina maculosa (G. B. Sowerby I, 1832) as Strombina (Spiralta) maculosa	BSE	125
	Strombina maculosa (G. B. Sowerby I, 1832) as Strombina (Spiralta) maculosa	IMu	125
Order Neogastropoda, Family Columbellidae	Strombina pulcherrima (G. B. Sowerby I, 1832) as Strombina (Lirastrombina) pulcherrima	BSE	125
	Strombina pulcherrima (G. B. Sowerby I, 1832) as Strombina (Lirastrombina) pulcherrima	IMu	125
	Strombina recurva (G. B. Sowerby I, 1832) as Strombina (Recurvina) recurva	BSE	125
	Strombina recurva (G. B. Sowerby I, 1832) as Strombina (Recurvina) recurva	PBl	125
	Strombina recurva (G. B. Sowerby I, 1832) as Strombina (Recurvina) recurva	BPG	125
	Strombina solidula (Reeve, 1859) as Strombina (Lirastrombina) solidula – doubtful record	BSE	125
Order Neogastropoda, Family Conidae	Conasprella lucida (W. Wood, 1828) as Conus lucidus Wood, 1828	BSE	93
	Conasprella perplexa (G. B. Sowerby II, 1857) as Conus perplexus	BSE	93
	Conasprella tornata (G. B. Sowerby I, 1833) as Conus tornatus	BSE	93
	Conus brunneus Wood, 1828	BSE	93
	Conus brunneus Wood, 1828	IMu	93
	Conus dalli Stearns, 1873	BSE	93
	Conus fergusoni G. B. Sowerby II, 1873	BSE	93
	Conus gladiator Broderip, 1833	BSE	93
	Conus recurvus Broderip, 1833	BSE	93
	Conus vittatus Hwass in Bruguière, 1792	BSE	93
Order Neogastropoda, Family Fasciolariidae	Fusinus dupetitthouarsi (Kiener, 1840)	IMu	130
	Opeatostoma pseudodon (Burrow, 1815)	IMu	130
	Pustulatirus hemphilli (Hertlein & Strong, 1951) as Latirus hemphilli	BSEa	114
	Pustulatirus mediamericanus (Hertlein & Strong, 1951) as Latirus mediamericanus	BSE	114
Order Neogastropoda, Family Marginellidae	Dentimargo zetetes Roth, 1978	BSEa	181
	Prunum aletes Roth, 1978 as Prunum (microspira) aletes	BSE	181
	Prunum aletes Roth, 1978 as Prunum (microspira) aletes	CSE	181
	Prunum aletes Roth, 1978 as Prunum (microspira) aletes	IMua	181
	Prunum lizanoi Magaña, Espinosa & Ortea, 2003	BJua	133, 199
Order Neogastropoda, Family Muricidae	Acanthina sp.	CSE	130
	Murexsul zeteki (Hertlein & Strong, 1951) as Muricopsis zeteki	BSE	114
	Plicopurpura columellaris (Lamarck, 1816) as Purpura pansa	CSE	130
	Vasula melones (Duclos, 1832)	BEH	185
Order Neogastropoda, Family Pseudomelatomidae	Crassispira xanti Hertlein & Strong, 1951	BSE	114
Order Unassigned, Family Cerithiopsidae	Cerithiopsis guanacastensis Hertlein & Strong, 1951	BSEa	114
	Seila kanoni (de Folin, 1867)	BSE	69
	Seila montereyensis Bartsch, 1907	CSE	69
Order Unassigned, Family Potamididae	Cerithideopsis californica (Haldeman, 1840) as Cerithidea valida	MCa	32
Order Unassigned, Family Potamididae	Cerithideopsis californica (Haldeman, 1840) as Cerithidea valida	MSa	32
	Cerithideopsis californica (Haldeman, 1840) as Cerithidea valida	PPG	32, 203
	Cerithideopsis californica (Haldeman, 1840) as Cerithidea valida	MPN	30, 208
Subclass Neritimorpha, Order Cycloneritimorpha, Family Neritidae	Nerita costata Gmelin, 1791 as Nerita scabricosta	BSE	128
	Nerita costata Gmelin, 1791 as Nerita scabricosta	IMu	130
Subclass Vetigastropoda, Order Unassigned, Family Scissurellidae	Scissurella kaiserae Geiger, 2006	CSE	85
Class Bivalvia, Subclass Heterodonta, Order Cardiida, Family Cardiidae	Americardia biangulata (Broderip & G. B. Sowerby I, 1829) as Cardium biangulatum	BSE	109
	Laevicardium substriatum (Conrad, 1837) as Cardium alenense	BSE	109
	Lophocardium annettae (Dall, 1889) as Cardium annettae	BSE	109
	Microcardium pazianum (Dall, 1916) as Cardium pazianum	BSE	109
	Trachycardium consors (G. B. Sowerby I, 1833) as Cardium consors	BSE	109
	Trachycardium procerum (G. B. Sowerby I, 1833) as Cardium procerum	BSE	109
	Trigoniocardia granifera (Broderip & G. B. Sowerby I, 1829) as Cardium graniferum	BSE	109
Order Cardiida, Family Psammobilidae	Heterodonax bimaculatus (Linnaeus, 1758) as Heterodonax bimaculata	BSE	113
Order Cardiida, Family Semelidae	Cumingia lamellosa G. B. Sowerby I, 1833	BSE	112
	Semele jovis (Reeve, 1853)	BSE	112
	Semele pallida (G. B. Sowerby I, 1833) as Semele simplicissima	BSE	112
	Semele verrucosa Mörch, 1860 as Semele pacifica	BSE	112
Order Cardiida, Family Solecurtidae	Tagelus affinis (C. B. Adams, 1852) as Tagelus (Tagelus) affinis	BSE	113
	Tagelus politus (Carpenter, 1857) as Tagelus (Mesopleura) politus	BSE	113
Order Cardiida, Family Tellinidae	Cymatoica undulata (Hanley, 1844) as Macoma (Cymatoica) undulata	BSE	111
	Macoma panamensis Dall, 1900 as Macoma (Psammacoma) panamensis	BSE	111
	Tellina amianta Dall, 1900 as Tellina (Moerella) amianta	BSE	111
	Tellina inaequistriata Donovan, 1802 as Tellina (Eurytellina) inaequistriata	BSE	111
	Tellina martinicensis d’Orbigny, 1853 as Tellina (Merisca) proclivis	BSE	111
	Tellina pristiphora Dall, 1900 as Tellina (Phyllodina) pristiphora	BSE	111
	Tellina prora Hanley, 1844 as Tellina (Eurytellina) prora	BSE	111
	Tellina rubescens Hanley, 1844 as Tellina (Eurytellina) rubescens	BSE	111
	Tellina tabogensis Salisbury, 1934 as Tellina (Moerella) recurvata	BSE	111
Order Carditida, Family Carditidae	Carditamera affinis (G. B. Sowerby I, 1833) as Cardita (Carditamera) affinis	BSE	108
	Carditamera radiata (G. B. Sowerby I, 1833) as Cardita (Carditamera) radiate	BSE	108
	Cardites laticostatus (G. B. Sowerby I, 1833) as Cardita tricolor	BSE	108
	Strophocardia megastropha (J.E. Gray, 1825) as Cardita megastropha	BSE	108
Order Carditida, Family Condylocardiidae	Condylocardia sparsa Coan, 2003	CSE	29
Order Carditida, Family Crassatellidae	Crassinella pacifica (C. B. Adams, 1852)	BSE	108
	Eucrassatella antillarum (Reeve, 1842) as Crassatellites (Hybolophus) digueti	BSE	108
	Eucrassatella gibbosa (G. B. Sowerby I, 1832) as Crassatellites (Hybolophus) gibbosus	BSE	108
Order Lucinida, Family Lucinidae	Codakia distinguenda (Tryon, 1872)	BSE	108
	Ctena mexicana (Dall, 1901)	BSE	108
	Liralucina approximata (Dall, 1901) as Lucina (Parvillucina) approximata	BSE	108
	Radiolucina cancellaris (Philippi, 1846) as Lucina (Bellucina) cancellaris	BSE	108
Order Myida, Family Corbulidae	Caryocorbula biradiata (G. B. Sowerby I, 1833) as Aloidis (Caryocorbula) biradiata	BSE	113
	Caryocorbula marmorata (Hinds, 1843) as Aloidis (Caryocorbula) marmorata	BSE	113
	Caryocorbula nasuta (G. B. Sowerby I, 1833) as Aloidis (Caryocorbula) nasuta	BSE	113
Order Myida, Family Pholadidae	Jouannetia pectinata (Conrad, 1849) as Jouannetia (Triomphalia) pectinata	BSE	113
Order Venerida, Family Veneridae	Agriopoma catharium (Dall, 1902) as Pitar (Pitarella) mexicanus	BSE	110
	Anomalocardia subimbricata (Sowerby, 1835)	BSE	110
	Chione compta (Broderip, 1835) as Chione (Chione) compta	BSE	110
	Cyclinella subquadrata (Hanley, 1844)	BSE	110
	Dosinia dunkeri (Philippi, 1844) as Dosinia (Dosinidia) dunkeri	BSE	110
	Dosinia ponderosa (Gray, 1838) as Dosinia (Dosinidia) ponderosa	BSE	110
	Gouldia californica Dall, 1917	BSE	110
	Iliochione subrugosa (W. Wood, 1828) as Anomalocardia subrugosa	BSE	110
	Leukoma asperrima (G. B. Sowerby I, 1835) as Chione (Nioche) asperrima	BSE	110
	Lirophora mariae (d’Orbigny, 1846) as Chione (Lirophora) mariae	BSE	110
	Megapitaria aurantiaca (G. B. Sowerby I, 1831)	BSE	110
	Megapitaria squalida (G. B. Sowerby I, 1835)	BSE	110
	Periglypta multicostata (G. B. Sowerby I, 1835) as Antigona (Periglypta) multicostata	BSE	110
	Pitar consanguineus (C. B. Adams, 1852) as Pitar (Pitar) consanguineous	BSE	110
	Pitar unicolor (Sowerby, 1835)	IMu	130
	Protothaca grata (Say, 1830)	MCa	32
Order Venerida, Family Veneridae	Protothaca grata (Say, 1830)	MSa	32
	Protothaca grata (Say, 1830) as Protothaca (Callithaca) grata	BSE	110
	Protothaca grata (Say, 1830)	MPG	32
	Protothaca grata (Say, 1830)	MPN	32
Order Unassigned, Family Chamidae	Arcinella californica (Dall, 1903) as Echinochama californica	BSE	108
	Chama echinata Broderip, 1835	Jun	185
Order Unassigned, Family Galeommatidae	Bellascintilla parmaleeana Coney, 1990	PNa	31
Order Unassigned, Family Ungulinidae	Diplodonta semirugosa Dall, 1899 as Taras semirugosus	BSE	109
	Diplodonta subquadrata Carpenter, 1856 as Taras subquadratus	BSE	109
Subclass Protobranchia, Order Nuculanida, Family Nuculanidae	Saccella elenensis (G. B. Sowerby I, 1833) as Nuculana (Saccella) elenensis	BSE	105
	Saccella impar (Pilsbry & Lowe, 1932) as Nuculana (Saccella) impar	BSE	105
	Saccella laeviradius (Pilsbry & Lowe, 1932) as Nuculana (Saccella) laeviradius	BSE	105
Subclass Pteriomorphia, Order Arcida, Family Arcidae	Acar gradata (Broderip & Sowerby, 1829) as Acar (Arca) gradata	BSE	105
	Acar gradata (Broderip & Sowerby, 1829)	PBl	180
	Anadara biangulata (G. B. Sowerby I, 1833) as Acar (Anadara) biangulata	BSE	106
	Anadara nux (G. B. Sowerby I, 1833) as Arca (Cunearca) nux	BSE	106, 180
	Anadara perlabiata (Grant & Gale, 1931) as Arca (Cunearca) perlabiata	BSE	106
	Anadara reinharti (Lowe, 1935) as Arca (Anadara) reinharti	BSE	106
	Anadara reinharti (Lowe, 1935) as Arca (Scapharca) reinharti	BSE	180
	Anadara tuberculosa (G. B. Sowerby I, 1833)	MCa	32
	Anadara tuberculosa (G. B. Sowerby I, 1833)	MSa	32
	Anadara tuberculosa (G. B. Sowerby I, 1833)	PPG	32, 208
	Anadara tuberculosa (G. B. Sowerby I, 1833)	MPN	32
	Arca mutabilis (G. B. Sowerby I, 1833) as Arca (Arca) mutabilis	BSE	106, 180
	Arca mutabilis (G. B. Sowerby I, 1833) as Arca (Arca) mutabilis	PBl	180
	Barbatia illota (G. B. Sowerby I, 1833) Barbatia (Fugleria) illota	PBl	180
	Barbatia reeveana (d’Orbigny, 1846) as Barbatia (Cucullaearca) reeveana	PBl	180
	Calloarca alternata (G. B. Sowerby I, 1833) as Arca (Calloarca) alternata	BSE	106
	Larkinia grandis (Broderip & G. B. Sowerby I, 1829) as Grandiarca grandis	MCa	32
	Larkinia grandis (Broderip & G. B. Sowerby I, 1829) as Arca (Lakinia) grandis	BSE	106
	Larkinia grandis (Broderip & G. B. Sowerby I, 1829) as Grandiarca grandis	MPG	32, 208
	Larkinia grandis (Broderip & G. B. Sowerby I, 1829) as Grandiarca grandis	MPN	32
Subclass Pteriomorphia, Order Arcida, Family Arcidae	Larkinia multicostata (G. B. Sowerby I, 1833) as Anadara multicostata	MCa	32
	Larkinia multicostata (G. B. Sowerby I, 1833) as Anadara multicostata	MSa	32
	Larkinia multicostata (G. B. Sowerby I, 1833) as Arca (Larkinia) multicostata	BSE	106
	Larkinia multicostata (G. B. Sowerby I, 1833) as Anadara multicostata	MPG	32, 204
	Larkinia multicostata (G. B. Sowerby I, 1833) as Anadara multicostata	MPN	32
Order Arcida, Family Glycymerididae	Tucetona strigilata (G. B. Sowerby I, 1833) as Glycymeris (Tuceta) tessellata strigilata and as Glycymeris (Tuceta) tessellata	BSE	106
Order Arcida, Family Noetiidae	Arcopsis solida (G. B. Sowerby I, 1833) as Arca (Arcopsis) solida	BSE	106
	Arcopsis solida (G. B. Sowerby I, 1833)	PBl	180
Order Arcida, Family Philobryidae	Philobrya setosa (Carpenter, 1864)	CSE	30
Order Limida, Family Limidae	Limaria orbignyi (Lamy, 1930) as Lima (Limaria) orbignyi	BSE	107
Order Mytilida, Family Mytilidae	Amygdalum americanum Soot-Ryen, 1955	PBl	186
	Brachidontes adamsianus (Dunker, 1856) as Hormomya adamsiana	BSE	186
	Brachidontes puntarenensis (Pilsbry & Lowe, 1932)	BSE	186
	Brachidontes sp.	Jun	185
	Crenella divaricata (Orbigny, 1853)	BSE	107, 186
	Leiosolenus aristatus (Dillwyn, 1817) as Lithophaga (Myoforceps) aristata	BSE	107, 186
	Leiosolenus aristatus (Dillwyn, 1817) as Lithophaga (Myoforceps) aristata	PBl	186
	Leiosolenus attenuatus (Deshayes, 1836) as Lithophaga (Labis) attenuata	BSE	107, 186
	Leiosolenus attenuatus (Deshayes, 1836) as Lithophaga (Labis) attenuata	PBl	186
	Leiosolenus plumula (Hanley, 1843) as Lithophaga (Diberus) plumula	BSE	107, 186
	Leiosolenus plumula (Hanley, 1843) as Lithophaga (Diberus) plumula	PBl	186
	Modiolus capax (Conrad, 1837) as Volsella (Volsella) capax in reference 107	BSE	107, 186
	Modiolus capax (Conrad, 1837)	PBl	186
	Mytilus sp.	MCa	32
	Mytilus sp.	MSa	32
	Mytilus sp.	MPG	32
	Mytilus sp.	MPN	32
	Septifer zeteki Hertlein & Strong, 1946	BSE	186
Order Ostreida, Family Ostreidae	Crassostrea corteziensis (Hertlein, 1951)	MPG	208
	Saccostrea palmula (Carpenter, 1857)	Jun	185
	Saccostrea palmula (Carpenter, 1857)	BSE	107
Order Ostreida, Family Pinnidae	Pinna rugosa G. B. Sowerby I, 1835	BSE	106
Order Ostreida, Family Pteriidae	Pinctada mazatlanica (Hanley, 1856)	BSE	106
	Pinctada mazatlanica (Hanley, 1856)	IMu	7
	Pteria sterna (Gould, 1851)	IMu	7
Order Pectinida, Family Anomiidae	Placunanomia cumingii Broderip, 1832	BSE	107
Order Pectinida, Family Pectinidae	Argopecten irradians concentricus (Say, 1822) as Pecten (Plagioctenium) circularis	BSE	107
	Euvola vogdesi (Arnold, 1906) as Pecten (Pecten) vogdesi	BSE	107
	Leopecten sericeus (Hinds, 1845) as Pecten (Pecten) sericeus	BSE	107
	Leptopecten biolleyi (Hertlein & Strong, 1946) as Pecten (Leptopecten) velero biolleyi	BSEa	107
	Nodipecten subnodosus (G. B. Sowerby I, 1835) as Pecten (Lyropecten) subnodosus	BSE	104, 107
Order Pectinida, Family Propeamussiidae	Cyclopecten pernomus (Hertlein, 1935)	BSE	107
Order Pectinida, Family Spondylidae	Spondylus sp.	IMu	7
Phylum SIPUNCULA, Class Phascolosomatidea, Order Phascolosomatida, Family Phascolosomatidae	Antillesoma antillarum (Grube & Oersted 1858)	Jun	185
	Antillesoma antillarum (Grube & Oersted 1858)	IMu	55, 59
	Phascolosoma (Phascolosoma) perlucens Baird, 1868	IMu	59
	Phascolosoma sp.	Jun	185
	Sipunculus (Sipunculus) nudus Linnaeus, 1766	Jun	185
Phylum ANNELIDA, Class Polychaeta, Subclass Errantia, Order Amphinomida, Family Amphinomidae	Eurythoe complanata (Pallas, 1776) as Eurythoe complanata in reference 95	BSE	60, 95, 189
	Eurythoe complanata (Pallas, 1776) as Eurythoe complanata in reference 95	PBl	60, 95
	Hermodice carunculata (Pallas, 1766) doubtful record	BSE	60, 189
	Notopygos ornata Grube, 1856	BSE	189
	Notopygos ornata Grube, 1856	PBl	60, 95
Order Eunicida, Family Eunicidae	Nicidion mutilata (Webster, 1884) as Eunice mutilata	BSE	60, 96
	Palola siciliensis (Grube, 1840)	BSE	60, 96
Order Eunicida, Family Lumbrineridae	One species	PJu	185
	Scoletoma tetraura (Schmarda, 1861) as Lumbrineris tetraura	BSE	60, 96
Order Eunicida, Family Oenonidae	Oenone fulgida (Savigny in Lamarck, 1818) as Aglaurides fulgida in reference 96	BSE	60, 96
	Oenone fulgida (Savigny in Lamarck, 1818) as Aglaurides fulgida in reference 96	PBl	60, 96
Order Eunicida, Family Onuphidae	Diopatra tridentata Hartman, 1944	BSE	60, 96
	Hyalinoecia juvenalis Moore, 1911	BSE	60, 96,189
	Hyalinoecia juvenalis Moore, 1911	PBl	60
Order Phyllodocida, Family Glyceridae	One species	PJu	185
	Glycera tesselata Grube, 1840	PBl	95
Order Phyllodocida, Family Iphionidae	Iphione ovata Kinberg, 1855	BSE	60, 94
Order Phyllodocida, Family Nereididae	One species	PJu	185
Order Phyllodocida, Family Polynoidae	Lepidasthenia varius Treadwell, 1917 as Lepidasthenia picta in reference 189	BSE	60, 189
Order Phyllodocida, Family Sigalionidae	Pelogenia antipoda Schmarda, 1861 as Psammolyce antipoda (Schmarda) anoculata	PBl	60, 94
	Sigalion lewisii Berkeley & Berkeley, 1939 as Thalenessa lewisii	BSE	94
	Sthenelais fusca Johnson 1897 as Stenelais variabilis colorata	BSE	94
Subclass Sedentaria, Order Spionida, Family Magelonidae	Magelona sp.	PJu	185
Order Terebellida, Family Terebellidae	One species	PJu	185
	Lanicola guillermoi Capa & Hutchings, 2006	BEH	185
	Terebella gorgonae Monro, 1933	BSE	60, 189
Order Unassigned, Family Chaetopteridae	One species	PJu	185
Order Unassigned, Family Opheliidae	Armandia maculata (Webster, 1884) as Ammotrypane bermudiensis	BSE	60, 189
Phylum NEMERTEA	One species	PJu	185
Phylum ARTHROPODA, Subphylum Crustacea Class Malacostraca, Order Amphipoda, Family Ampeliscidae	Ampelisca brevisimulata Barnard, 1954	BSE	5
	Ampelisca cristata Holmes, 1908	BSE	5
	Ampelisca hancocki Barnard, 1954	BSEa	5
	Ampelisca lobata Holmes, 1908	BSE	5
	Ampelisca milleri Barnard, 1954	BSE	5
	Ampelisca milleri Barnard, 1954	PBl	5
	Ampelisca pugetica Stimpson, 1864 as Ampelisca pugetica forma macrodentata	BSE	5
	Ampelisca romigi Barnard, 1954 as Ampelisca isocornea	BSE	5
	Ampelisca schellenbergi Shoemaker, 1933	PBl	5
	Ampelisca venetiensis Shoemaker, 1916	BSE	5
Order Amphipoda, Family Aoridae	Paramicrodeutopus schmitti (Shoemaker, 1942) as Microdeutopus schmitti	PBl	152
Order Amphipoda, Family Neomegamphopidae	Neomegamphopus roosevelti Shoemaker, 1942	PBl	152
Order Amphipoda, Family Phoxocephalidae	Microphoxus minimus Barnard, 1954	PBla	5, 6
Order Amphipoda, Family Unciolidae	Acuminodeutopus periculosus Barnard, 1969 as Acuminodetopus heteruropus	PBl	152
Order Amphipoda	Several unidentified species	IMu	75
Order Cumacea, Family Bodotriidae	Cyclaspis vargasae Petrescu & Heard, 2004	IMua	161, 162
Order Decapoda, Family Albuneidae	Lepidopa mearnsi Benedict, 1904	PNa	197
Order Decapoda, Family Alpheidae	Alpheus aequus Kim & Abele, 1988	PBla	126, 204
	Alpheus cylindricus Kingsley, 1878	BSE	126
	Alpheus galapagensis Sivertsen, 1933 as Alpheus canalis Kim & Abele, 1988	BSE	126
	Alpheus galapagensis Sivertsen, 1933 as Alpheus canalis Kim & Abele, 1988	PBl	126
	Alpheus hebes Kim & Abele, 1988	BSE	126
	Alpheus hebes Kim & Abele, 1988	PBl	126
	Alpheus longinquus Kim & Abele, 1988	BSE	126
	Alpheus longinquus Kim & Abele, 1988	PBl	126
	Alpheus normanni Kingsley, 1878	BSE	126
	Alpheus panamensis Kingsley, 1878	CSE	196
	Alpheus paracrinitus Miers, 1881	PBl	126
	Alpheus rostratus Kim & Abele, 1988	BSE	126, 204
	Alpheus sulcatus Kingsley, 1870	CSE	196
	Alpheus umbo Kim & Abele, 1988	BSE	126
	Alpheus sp.	BEH	185
	Pomagnathus corallinus Chace, 1962	BSE	196
	Pomagnathus corallinus Chace, 1962	PBl	202
	Synalpheus digueti Coutière, 1909	BSE	202
Order Decapoda, Family Axiidae	Axiopsis serratifrons (H. Milne-Edwrads, 1873)	IMu	196
Order Decapoda, Family Calappidae	Calappa convexa Saussure, 1853	BSE	81
	Calappula saussurei (Rathbun, 1898) as Calappa saussurei	BSE	198
	Cryptosoma bairdii (Stimpson, 1860) as Cycloes bairdii	BSE	82
	Cryptosoma bairdii (Stimpson, 1860) as Cycloes bairdii	IMu	82
	Platymera gaudichaudii H. Milne-Edwards, 1837 as Mursia gaudichaudii	CSE	129
	Platymera gaudichaudii H. Milne Edwards, 1837	IMu	198
Order Decapoda, Family Callianideidae	Callianidea mariamartae Hernáez & Vargas, 2013	IMua	102
	Paracallianidea laevicauda (Gill, 1859) as Callianidea laevicauda	PBl	196
Order Decapoda, Family Coenobitidae	Coenobita compressus H. Milne Edwards, 1837	IMu	197
Order Decapoda, Family Dairidae	Daira americana Stimpson, 1860	BSE	47
Order Decapoda, Family Daldorfiidae	Daldorfia trigona (A. Milne-Edwards, 1869) as Daldorfia garthi	BSE	47
	Daldorfia trigona (A. Milne-Edwards, 1869) as Daldorfia garthi	PBl	78
Order Decapoda, Family Diogenidae	Aniculus elegans Stimpson, 1858	IMu	197
	Trizopagurus magnificus (Bouvier, 1898)	IMu	197
Order Decapoda, Family Domeciidae	Cherusius triunguiculatus (Borradaile, 1902) as Maldivia galapagensis	BSE	83
Order Decapoda, Family Dromiidae	Hypoconcha panamensis Smith, 1869	BSE	82
	Moreiradromia sarraburei (Rathbun, 1910)	BSE	198
Order Decapoda, Family Dynomenidae	Hirsutodynomene ursula (Stimpson, 1860) as Dynomene ursula Stimpson, 1860	IMu	151
Order Decapoda, Family Epialtidae	Herbstia pubescens Stimpson, 1871	BSE	80
	Herbstia pubescens Stimpson, 1871	PBl	79
	Macrocoeloma maccullochae Garth, 1940	BSE	79, 80
	Macrocoeloma maccullochae Garth, 1940	PBl	78
	Microlissa aurivilliusi (Rathbun, 1898) as Lissa aurivilliusi	BSE	80
	Microlissa aurivilliusi (Rathbun, 1898) as Lissa aurivilliusi	PBl	79
	Pelia tumida (Lockington, 1877)	PBl	79
	Stenocionops ovatus (Bell, 1835) as Stenocionops ovata	IMu	198
Order Decapoda, Family Eriphiidae	Eriphia squamata Stimpson, 1859	BSE	47
	Eriphides hispida (Stimpson, 1860)	BSE	47
Order Decapoda, Family Ethusidae	Ethusa lata Rathbun, 1893	BSE	81
	Ethusa lata Rathbun, 1893	IMu	198
	Ethusa panamensis Finnegan, 1931 as Ethusa mascarpone panamensis	IMu	82
Order Decapoda, Family Gecarcinidae	Johngarthia planata (Stimpson, 1860)	IMu	160
Order Decapoda, Family Grapsidae	Geograpsus lividus (Milne Edwards, 1837)	BSE	47
	Grapsus grapsus (Linnaeus, 1758)	BSE	47
	Grapsus grapsus (Linnaeus, 1758)	PPG	47
	Pachygrapsus transversus (Gibbes, 1850)	BSE	47
Order Decapoda, Family Hippidae	Emerita rathbunae Schmitt, 1935	Jun	197
Order Decapoda, Family Hippolytidae	Hippolyte williamsi Schmitt, 1924	IMu	201
	Lysmata argentopunctata Wicksten, 2000	IMu	201, 204
	Trachycaris restricta (Milne-Edwards, 1878)	BSE	196
Order Decapoda, Family Inachidae	Ericerodes angulatus (Finnegan, 1931) as Podochela angulata	BSE	79
	Ericerodes angulatus (Finnegan, 1931) as Podochela angulata	BSE	80
	Ericerodes veleronis (Garth, 1948) as Podochela veleronis	PBl	79
	Eucinetops panamensis Rathbun, 1923	BSE	47
	Eucinetops panamensis Rathbun, 1923	PBl	79
	Podochela ziesenhennei Garth, 1940	PBl	79
Order Decapoda, Family Inachoididae	Collodes tenuirostris Rathbun, 1894	IMu	198
	Euprognatha bifida Rathbun, 1893	BSE	80
	Inachoides laevis Stimpson, 1860	BSE	79
	Inachoides laevis Stimpson, 1860	BSE	80
	Paradasygyius depressus (Bell, 1835)	BSE	79
	Pyromaia tuberculata (Lockington, 1877)	IMu	198
	Stenorhynchus debilis (Smith, 1871)	BSE	79
	Stenorhynchus debilis (Smith, 1871)	PBl	79
	Stenorhynchus debilis (Smith, 1871)	BSE	80
Order Decapoda, Family Leucosiidae	Ebalia magdalenensis Rathbun, 1933	BSE	82
	Leucosilia jurinii (Saussure, 1853)	BSE	82
	Lithadia cumingii Bell, 1855	BSE	82
	Persephona subovata (Rathbun, 1894) as Iliacantha hancocki	BSE	82
	Randallia agaricias Rathbun, 1898	BSE	82
Order Decapoda, Family Majidae	Maiopsis panamensis Faxon, 1893	IMu	198
Order Decapoda, Family Mithracidae	Ala cornuta (Stimpson, 1860) as Anaptychus cornutus	BSE	47
	Ala cornuta (Stimpson, 1860)	BSE	79, 80
	Ala cornuta (Stimpson, 1860)	PBl	79
	Hemus finneganae Garth, 1958	BSE	80
	Microphrys branchialis Rathbun, 1898	BSE	79
	Microphrys platysoma (Stimpson, 1860)	BSE	47
	Mithraculus denticulatus (Bell, 1835) as Mithrax denticulatus	BSE	47
	Mithraculus denticulatus (Bell, 1835) as Mithrax denticulatus	PBl	79
	Mithrax tuberculatus Stimpson, 1860	PBl	79
	Petramithrax pygmaeus Bell, (1836) as Mithrax pygmaeus	BSE	47, 80
	Pitho picteti (Saussure, 1853)	PBl	79
	Pitho picteti (Saussure, 1853)	BSE	80
	Pitho quinquedentata Bell, 1835	BSE	80
	Pitho sexdentata Bell, 1835	BSE	47
	Teleophrys cristulipes Stimpson, 1860	BSE	47, 79
	Teleophrys cristulipes Stimpson, 1860	PBl	79
	Thoe erosa Bell, 1835 as Thoe sulcata panamensis	BSE	47, 79
	Thoe erosa Bell, 1835 as Thoe sulcata panamensis	PBl	79
Order Decapoda, Family Munididae	Pleuroncodes planipes Stimpson, 1860	CSE	129
Orden Decapoda, Family Ocypodidae	Ocypode gaudichaudii Milne Edwards & Lucas, 1843	BPG	44, 46
	Uca (Leptuca) deichmanni Rathbun, 1935 as Uca deichmanni	BSE	45
	Uca (Leptuca) latimanus (Rathbun, 1894) as Uca latimanus	BSE	45
	Uca (Leptuca) panamensis (Stimpson, 1859) as Uca panamensis	BSE	45
	Uca (Leptuca) stenodactylus (H. Milne Edwards & Lucas, 1843) as Uca stenodactyla	BSE	45
	Uca (Leptuca) terpsichores Crane, 1941 as Uca terpsichores	BSE	45
	Uca (Minuca) brevifrons (Stimpson, 1860) as Uca brevifrons	BSE	45
	Uca sp.	MCa	32
	Uca sp.	MSa	32
	Uca sp.	MPG	32, 208
	Uca sp.	MPN	32
Order Decapoda, Family Oziidae	Epixanthus tenuidactylos (Lockington, 1877) as Ozius tenuidactylus	BSE	47
	Ozius verreauxii Saussure, 1853	BSE	47
Order Decapoda, Family Paguridae	Pagurus vetaultae Harvey & McLaughlin, 1991	BSEb	97
	Pagurus virgulatus Haig & Harvey, 1991	BSEa	92, 197
Order Decapoda, Family Palaemonidae	Ancylomenes lucasi (Chace, 1937) as Periclimenes lucasi	PBl	118
	Brachycarpus biunguiculatus (Lucas, 1846)	BSE	119
	Brachycarpus biunguiculatus (Lucas, 1846)	PBl	119
	Brachycarpus biunguiculatus (Lucas, 1846)	IMu	196
	Fennera chacei Holthuis, 1951	BSE	118
	Periclimenes infraspinis (Rathbun, 1902) as Periclimenaeus infraspinis	BSE	118, 202, 204
	Periclimenes murcielagensis Vargas, 2000	IMua	194, 204
	Pontonia margarita Smith, 1869	BSE	118
	Pontonia margarita Smith, 1869	PBl	118
	Pontonia margarita Smith, 1869	IMu	196
	Pontonia simplex Holthuis, 1951	BSE	196
	Waldola schmitti Holthuis, 1951	IMu	196
Order Decapoda, Family Panopeidae	Eurypanopeus planus (Smith, 1869)	BSE	47
	Eurypanopeus transversus (Stimpson, 1860)	BSE	47
	Hexapanopeus costaricensis Garth, 1940	BSEa	78, 81
	Hexapanopeus orcutti Rathbun, 1930	BSE	81
	Hexapanopeus sinaloensis Rathbun, 1930	BSE	81
	Malacoplax californiensis (Lockington, 1877) as Speocarcinus californiensis	BSE	81
Order Decapoda, Family parthenopidae	Celatopesia hassleri (Rathbun, 1925) as Cryptopodia hassleri	BSE	79, 80
	Celatopesia hassleri (Rathbun, 1925) as Cryptopodia hassleri	PBl	79
	Heterocrypta macrobrachia Stimpson, 1871	BSE	79
	Hypolambrus hyponcus (Stimpson, 1871) as Pathenope (Pathenope) hyponca	PBl	79
	Mesorhoea bellii (A. Milne-Edwards, 1878)	BSE	198
	Solenolambrus arcuatus Stimpson, 1871	BSE	79, 80
	Solenolambrus arcuatus Stimpson, 1871	PBl	79
	Spinolambrus exilipes (Rathbun, 1893) as Parthenope exilipes	IMu	198
Order Decapoda, Family Pasiphaeidae	Leptochela (Leptochela) serratorbita Spence Bate, 1888 as Leptochela serratorbita	BSE	202, 204
Order Decapoda, Family Penaeidae	Metapenaeopsis beebei (Burkenroad, 1938)	BSE	198
	Penaeus sp. as Pennaeus	MCa	32
	Penaeus sp. as Pennaeus	MSa	32
	Penaeus sp. as Pennaeus	MPG	32, 208
	Penaeus sp. as Pennaeus	MPN	32
Order Decapoda, Family Pilumnidae	Pilumnus pygmaeus Boone, 1927	BSE	47, 81
Order Decapoda, Family Pinnotheridae	Glassella costaricana (Wicksten, 1982)	BEH	185
Order Decapoda, Family Porcellanidae	Euceramus transversilineatus (Lockington, 1878)	PMo	197
	Euceramus transversilineatus (Lockington, 1878)	BSE	90
	Megalobrachium pacificum Gore & Abele, 1973	BEH	197
	Megalobrachium pacificum Gore & Abele, 1973	PBl	87
	Pachycheles biocellatus (Lockington, 1878)	BSE	90
	Pachycheles vicarius Nobili, 1901	BSE	90
	Petrolisthes agassizii Faxon, 1893	PJu	197
	Petrolisthes agassizii Faxon, 1893	BSE	91
	Petrolisthes armatus (Gibbes, 1850)	Jun	197
	Petrolisthes armatus (Gibbes, 1850)	BSE	91
	Petrolisthes artifrons Haig, 1960	BSE	90
	Petrolisthes edwardsii (de Saussure, 1853)	Jun	197
	Petrolisthes edwardsii (de Saussure, 1853)	BSE	90
	Petrolisthes edwardsii (de Saussure, 1853)	PBl	90
	Petrolisthes glasselli Haig, 1957	BSE	90, 91
	Petrolisthes haigae Chace, 1962	Jun	197
	Petrolisthes haigae Chace, 1962	BSE	91
	Petrolisthes haigae Chace, 1962	PSE	197
	Petrolisthes hians Nobili, 1902	BSE	90, 91
	Petrolisthes holotrichus Nobili, 1901	BSE	90
	Petrolisthes lewisi Glassell, 1936 as Petrolisthes lewisi austrinus	BSE	90, 100
	Petrolisthes lewisi Glassell, 1936	PBl	197
	Petrolisthes nobilii Haig, 1960	BSE	90
	Petrolisthes nobilii Haig, 1960	PBl	197
	Petrolisthes ortmanni Nobili, 1901	BSE	90, 91
	Petrolisthes ortmanni Nobili, 1901	PBl	90
	Petrolisthes platymerus Haig, 1960	BSE	90, 101
Order Decapoda, Family Porcellanidae	Petrolisthes platymerus Haig, 1960	PBl	197
	Petrolisthes polymitus Glassell, 1937	PBl	90
	Petrolisthes tonsorius Haig, 1960	BSE	90
	Petrolisthes tridentatus Stimpson, 1859	BSE	90, 91
	Pisidia magdalenensis (Glassell, 1936)	Jun	197
	Pisidia magdalenensis (Glassell, 1936)	BSE	90, 91
	Pisidia magdalenensis (Glassell, 1936)	PBl	90
	Polyonyx nitidus Lockington, 1878	BJu	197
	Porcellana cancrisocialis Glassell, 1936	BSE	90, 91
	Porcellana paguriconviva Glassell, 1936	BSE	90, 91
Order Decapoda, Family Portunidae	Achelous asper (A. Milne-Edwards, 1861) as Portunus (Portunus) panamensis	BSE	81
	Achelous tuberculatus Stimpson, 1860 as Portunus (Acheolus) tuberculatus	BSE	81
	Arenaeus mexicanus (Gerstaecker, 1856)	IMu	81
	Arenaeus mexicanus (Gerstaecker, 1856)	BPG	81
	Callinectes arcuatus Ordway, 1863	BSE	81
	Cronius ruber (Lamarck, 1818)	BSE	81
	Portunus (Portunus) acuminatus (Stimpson, 1871)	BSEc	78, 81, 198
	Portunus (Portunus) asper (A. Milne-Edwards, 1861)	BSE	81
	Portunus (Portunus) asper (A. Milne-Edwards, 1861)	IMu	81
Order Decapoda, Family Pseudorhombilidae	Lophoxanthus lamellipes (Stimpson, 1860)	BSE	47
Order Decapoda, Family Sesarmidae	Aratus pisonii (H. Milne Edwards, 1837)	BSE	47
Order Decapoda, Family Sicyoniidae	Sicyonia disdorsalis (Burkenroad, 1934)	PNa	196
	Sicyonia disedwardsi (Burkenroad, 1934)	PMo	196
	Sicyonia laevigata Stimpson, 1871	BSE	196
	Sicyonia laevigata Stimpson, 1871	IMu	196
	Sicyonia martini Pérez-Farfante & Boothe, 1981	IDa	196
	Sicyonia martini Pérez-Farfante & Boothe, 1981	IMu	198
	Sicyonia martini Pérez-Farfante & Boothe, 1981	PNa	196
	Sicyonia picta Faxon, 1893	PNa	196
Order Decapoda, Family Solenoceridae	Solenocera florea Burkenroad, 1938	PBr	196
Order Decapoda, Family Trapeziidae	Trapezia bidentata (Forskål, 1775) as Trapezia cymodoce ferruginea	BSE	47
Order Decapoda, Family Upogebiidae	Upogebia dawsoni Williams, 1986	BPG	205
Order Decapoda, Family Xanthidae	Cataleptodius taboganus (Rathbun, 1912) as Leptodius taboganus	BSE	47
	Cycloxanthops vittatus (Stimpson, 1860)	BSE	47
	Edwardsium lobipes (Rathbun, 1898) as Medaeus lobipes in reference 81	BSE	81, 198
	Heteractaea lunata (Lucas, in H. Milne Edwards & Lucas, 1844)	BSE	47
	Liomera cinctimanus (White, 1847) as Carpilodes cinctimanus	BSE	47
	Microcassiope xantusii (Stimpson, 1871) as Micropanope xantusii	BSE	47
	Paractaea sulcata (Stimpson, 1860) as Actaea sulcata	BSE	47
Order Decapoda, Family Xanthidae	Platyactaea dovii (Stimpson, 1871) as Actaea dovii	BSE	47, 81
	Williamstimpsonia stimpsoni (Milne Edwards, 1879) as Xanthodius stimpsoni	BSE	47
	Xanthodius sternberghii Stimpson, 1859	BSE	47
Order Mysida, Family Mysidae	Chlamydopleon banneri (Bacescu, 1968) as Bowmaniella banneri	IMu	98
	Several species	IMu	171
Order Stomatopoda, Family Gonodactylidae	Neogonodactylus bahiahondensis (Schmitt, 1940)	PSE	195
	Neogonodactylus bahiahondensis (Schmitt, 1940)	IMu	195
	Neogonodactylus costaricensis (Manning & Reaka, 1979)	BSE	136
	Neogonodactylus festae (Nobili, 1901)	BSE	136
	Neogonodactylus zacae (Manning, 1972)	PMo	195
	Neogonodactylus zacae (Manning, 1972)	BSE	195
Order Stomatopoda, Family Nannosquillidae	Nannosquilla canica Manning & Reaka, 1979	PBl	137
	Nannosquilla decemspinosa (Rathbun, 1910) as Lysiosquilla decemspinosa	PBl	135
Order Stomatopoda, Family Pseudosquillidae	Pseudosquillisma adiastalta Manning, 1964	CSE	195
Order Stomatopoda, Family Squillidae	Crenatosquilla oculinova (Glassell, 1942)	CSE	195
	Squilla biformis Bigelow, 1891	CSE	129, 195
	Squilla panamensis Bigelow, 1891	CSE	195
Order Tanaidacea, Family Leptocheliidae	Several species	IMu	99
Order Tanaidacea, Family Parapseudidae	Parapseudes latifrons (Grube, 1864) as Parapseudes pedispinis	BSE	148
	Parapseudes latifrons (Grube, 1864) as Parapseudes pedispinis	PBl	148
	Parapseudes latifrons (Grube, 1864) as Parapseudes pedispinous	IMu	99
Order Tanaidacea, Family Tanaididae as Family Tanaidae	Several species	IMu	99
Class Maxillopoda, Subclass Copepoda, Family Acartiidae	Acartia (Acartia) negligens Dana, 1849 as Acartia (Planktacartia) negligens	IMu	187
Class Maxillopoda, Infraclass Cirripedia, Order Sessilia, Family Balanidae	Amphibalanus inexpectatus (Pilsbry, 1916) as Balanus inexpectatus	Jun	185
Order Sessilia, Family Chthamalidae	Chthamalus panamensis Pilsbry, 1916	Jun	185
	Chthamalus panamensis Pilsbry, 1916	CSE	27, 163
	Microeuraphia imperatrix (Pilsbry, 1916)	CSE	27
Order Sessilia, Family Tetraclitidae	Tetraclita stalactifera (Lamarck, 1818)	Jun	185
Phylum BRYOZOA, Class Gymnolaemata, Order Cheilostomatida, Family Bugulidae	Sessibugula translucens Osburn, 1950	BSE	156
Order Cheilostomatida, Family Exechonellidae	Anexechona ancorata Osburn, 1950	BSE	156
Order Cheilostomatida, Family Phidoloporidae	Rhynchozoon rostratum (Busk, 1855)	PBl	157
Order Cheilostomatida, Family Schizoporellidae	Schizoporella inarmata Hincks, 1884 as Schizoporella linearis var. inarmata	BSE	157
Class Stenolaemata, Order Cyclostomatida, Family Crisiidae	Crisia occidentalis Trask, 1857	BSE	158
Order Cyclostomatida, Family Lichenoporidae	Disporella californica (d’Orbigny, 1853)	BSE	158
Order Cyclostomatida, Family Tubuliporidae	Tubulipora pulchra MacGillivray, 1885	BSE	158
Order Cyclostomatida, Family Unassigned	Diaperoforma californica (d’Orbigny, 1853) as Diaperoecia californica	PBl	158
	Petralia japonica (Busk, 1884)	BSE	157
Phylum ECHINODERMATA, Class Asteroidea, Order Valvatida, Family Oreasteridae	Pentaceraster cumingi (Gray, 1840) as Oreaster occidentalis	BSE	28
Class Echinoidea, Order Camarodonta, Family Echinometridae	Echinometra vanbrunti A. Agassiz, 1863	Jun	185
Class Holothuroidea, Order Aspidochirotida, Family Holothuriidae	Holothuria (Cystipus) rigida (Selenka, 1867) as Fossothuria rigida	PBl	63
	Holothuria (Halodeima) kefersteinii (Selenka, 1867) as Ludwigothuria kefersteini	PBl	63
	Holothuria (Selenkothuria) lubrica Selenka, 1867	BSE	63
	Holothuria (Semperothuria) languens Selenka, 1867 as Semperothuria languens	PBl	63
	Holothuria (Thymiosycia) arenicola Semper, 1868 as Brandtothuria arenicola	BSE	63
	Holothuria (Thymiosycia) arenicola Semper, 1868 as Brandtothuria arenicola	PBl	63
	Holothuria (Thymiosycia) impatiens (Forskål, 1775) as Brandtothuria impatiens	BSE	63
	Holothuria (Thymiosycia) impatiens (Forskål, 1775) as Brandtothuria impatiens	PBl	63
Order Dendrochirotida, Family Cucumariidae	Neocucumis veleronis (Deichmann, 1941)	PBla	62
	Pseudocnus californicus (Semper, 1868) as Cucumaria californica	BSE	62
	Pseudocnus dubiosus dubiosus (Semper, 1868) as Cucumaria dubiosa	BSE	62
Order Dendrochirotida, Family Phyllophoridae	Pentamera chierchiae (Ludwig, 1887)	BSE	61, 62
	Pentamera chierchiae (Ludwig, 1887)	PBl	62
Order Dendrochirotida, Family Sclerodactylidae	Afrocucumis ovulum (Selenka, 1867) as Euthyonidium ovulum	BSE	61
	Neothyone gibber (Selenka, 1867)	BSE	62
	Neothyone gibbosa Deichmann, 1941	PBl	62
Phylum CHORDATA, Subphyllum Tunicata, Class Ascidiacea, Order Aplousobranchia, Family Didemnidae	Didemnum moseleyi (Herdman, 1886) as Didemnum mosseleyi	IDa	154
	Didemnum moseleyi (Herdman, 1886) as Didemnum mosseleyi	BVi	154
	Lissoclinum caulleryi (Ritter & Forsyth, 1917)	IDa	154
	Lissoclinum caulleryi (Ritter & Forsyth, 1917)	BVi	154
Order Phlebobranchia, Family Ascidiidae	Ascidia ceratodes (Huntsman, 1912)	IDa	154
	Ascidia ceratodes (Huntsman, 1912)	BRo	154
	Ascidia ceratodes (Huntsman, 1912)	BVi	154
Order Aplousobranchia, Family Diazonidae	Rhopalaea birkelandi Tokioka, 1971	IDa	154
	Rhopalaea birkelandi Tokioka, 1971	BRo	154
	Rhopalaea birkelandi Tokioka, 1971	BVi	154
Order Stolidobranchia, Family Styelidae	Eusynstyela tincta (Van Name, 1902) as Polyandrocarpa tincta	IDa	154
	Eusynstyela tincta (Van Name, 1902) as Polyandrocarpa tincta	BVi	154
Subphylum Cepahalochordata, Class Leptocardii, Order Unassigned, Family Branchiostomatidae	Branchiostoma californiense Andrews, 1893	PJu	185
Subphylum Vertebrata, Superclass Pisces, Class Elasmobrabranchii, Order Myliobatiformes, Family Dasyatidae	Dasyatis longa (Garman, 1880) as Dasyatis longus	Cua	24, 178
Order Myliobatiformes, Family Urotrygonidae	Urobatis pardalis Del Moral-Flores, Angulo, López & Bussing, 2015	IMub	64
Order Torpediniformes, Family Narcinidae	Narcine entemedor Jordan & Starks, 1895	Cua	40, 138, 178
Class Actinopteri, Order Anguilliformes, Family Muraenidae	Echidna nocturna (Cope, 1872)	Cua	156,178
Order Perciformes, Family Cirrhitidae	Oxycirrhites typus Bleeker, 1857	IMu	7
Order Perciformes, Family Gobiidae	Chriolepis cuneata Bussing, 1990	CSE	19
	Chriolepis cuneata Bussing, 1990	IMua	19
	Elacatinus digueti (Pellegrin, 1901) as Elacatious ioornatus in reference 19	IMua	19, 117
Order Perciformes, Family Haemulidae	Microlepidotus brevipinnis (Steindachner, 1869)	Cua	149
Order Perciformes, Family Labridae	Thalassoma lucasanum (Gill, 1862)	IMu	146
Order Perciformes, Family Labrisomidae	Dialommus fuscus (Gilbert, 1891)	IMu	88
	Paraclinus monophthalmus (Günther, 1861)	BSE	179
Order Perciformes, Family Opistognathidae	Opistognathus fossoris Bussing & Lavenberg, 2003	IMua	21
Order Perciformes, Family Trpterygiidae	Lepidonectes clarkhubbsi Bussing, 1991	BSEb	20
	Lepidonectes clarkhubbsi Bussing, 1991	CSEb	20
	Lepidonectes clarkhubbsi Bussing, 1991	IMua, b	20
Orden Syngnathiformes, Family Syngnathidae	Bryx veleronis Herald, 1940	IMu	88
Class Reptilia, Order Crocodylia, Family Crocodylidae	Crocodylus acutus (Cuvier, 1807)	MPG	128
	Crocodylus acutus (Cuvier, 1807)	MPN	140
Order Testudines, Family Cheloniidae	Chelonia mydas agassizi Bocourt, 1868	PNa	33, 34, 35, 65, 67, 115
	Lepidochelys olivacea (Eschscholtz, 1829)	PNc	1, 10, 25, 33, 35, 36, 37 48, 49, 70, 71, 74, 84, 89, 120, 121, 122, 123, 147, 148, 150, 153, 155, 164, 165, 166, 167, 168, 169, 175, 183, 190, 192, 193, 202
	Lepidochelys olivacea (Eschscholtz, 1829)	PNa	1, 34, 35, 65, 66, 67
Order Testudines, Family Dermochelyidae	Dermochelys coriacea (Vandelli, 1761)	PNa	34, 35, 65
Class Aves, Order Pelecaniformes, Family Fregatidae	Fregata magnificens Mathews, 1914	IBo	2
Order Pelecaniformes, Family Pelecanidae	Pelecanus occidentalis Linnaeus, 1766	PNa	3
Class Mammalia, Order Cetartiodactyla, InfraOrder Cetacea, SuperFamily Mysticeti, Family Balaenopteridae	Megaptera novaeangliae Borowski, 1781	SMa	139
SuperFamily Odontoceti, Family Delphinidae	Globicephala macrorhynchus Gray, 1846	IMu	145
	Orcinus orca Linnaeus, 1758	IMu	145
	Orcinus orca Linnaeus, 1758	SMa	139
	Stenella attenuata graffmani (Lönnberg, 1934)	Cua	145
	Senella attenuata graffmani (Lönnberg, 1934)	IMu	141, 142, 143, 144
	Stenella attenuata (Gray, 1846)	SMa	139
	Tursiops truncatus Montagu, 1821	SMa	139
SuperFamily Odontoceti, Family Kogiidae	Kogia breviceps (de Blainville, 1838)	IMu	145
SuperFamily Odontoceti, Family Physeteridae	Physeter macrocephalus Linnaeus, 1758 as Physeter catodon	PSE	177