Arrival to and location of Bufo woodhousei in the breeding pond: effect on the operational sex ratio.

Individual male and female Bufo woodhousei spent only two or three nights of the several week breeding season in the pond. In spite of the continual arriving and leaving of individuals the operational sex ratio (the number of females available to mate/number of males attempting to mate) varied little, either temporally across the breeding season or spatially between male groups in the pond. Male body size distributions also varied little between nights and between male groups.