Xiang Yu1,2,3, Zilong Qiu4,5, Dai Zhang6,7,8. 1. Institute of Neuroscience, State Key Laboratory of Neuroscience, Center for Excellence in Brain Science and Intelligence Technology, Chinese Academy of Sciences, Shanghai, 200031, China. yuxiang@ion.ac.cn. 2. University of Chinese Academy of Sciences, Beijing, 100049, China. yuxiang@ion.ac.cn. 3. School of Life Science and Technology, ShanghaiTech University, Shanghai, 201210, China. yuxiang@ion.ac.cn. 4. Institute of Neuroscience, State Key Laboratory of Neuroscience, Center for Excellence in Brain Science and Intelligence Technology, Chinese Academy of Sciences, Shanghai, 200031, China. zqiu@ion.ac.cn. 5. University of Chinese Academy of Sciences, Beijing, 100049, China. zqiu@ion.ac.cn. 6. Institute of Mental Health, Peking University Sixth Hospital, Beijing, 100191, China. daizhang@bjmu.edu.cn. 7. Peking-Tsinghua Center for Life Sciences, Peking University, Beijing, 100871, China. daizhang@bjmu.edu.cn. 8. PKU-IDG/McGovern Institute for Brain Research, Peking University, Beijing, 100871, China. daizhang@bjmu.edu.cn.