Mark Westoby1, Georges Kunstler2, Michelle R Leishman3, John Morgan4. 1. Department of Biological Sciences, Macquarie University, Sydney NSW 2109, Australia. Electronic address: mark.westoby@mq.edu.au. 2. Irstea, UR EMGR, 2 rue de la Papeterie BP-76, F-38402, St-Martin-d'Hères, France. 3. Department of Biological Sciences, Macquarie University, Sydney NSW 2109, Australia. 4. Department of Ecology, Environment and Evolution, La Trobe University, Bundoora VIC 3086, Australia.