Literature DB >> 27551230

An updated list of type material of Ephemeroptera Hyatt & Arms, 1890, deposited at the Zoological Museum of Hamburg (ZMH).

Michel Sartori1, Martin Kubiak2, Hossein Rajaei3.   

Abstract

The type specimens of Ephemeroptera (Insecta) housed at the Zoological Museum of Hamburg (ZMH) are compiled in this document. The current nomenclature of all species is given. In total, Ephemeroptera type material of ZMH encompasses 161 species. Fifty-one holotypes and five lectotypes are present. Forty-one species are represented by syntypes, 85 by paratypes and five by paralectotypes. Material of two species (Cinygma asiaticum Ulmer, 1924 and Pseudocloeon klapaleki Müller-Liebenau, 1982) is missing. The present catalogue is an updated version of Weidner (1964a).

Entities:  

Keywords:  Ephemeroptera; Hamburg; Ulmer; ZMH; type specimens

Year:  2016        PMID: 27551230      PMCID: PMC4978026          DOI: 10.3897/zookeys.607.9391

Source DB:  PubMed          Journal:  Zookeys        ISSN: 1313-2970            Impact factor:   1.546


Introduction

The collection of the Zoological Museum of Hamburg (ZMH) contains 433 species and approximately 4,500 specimens. In total, 161 species are represented by type material. In this sense, this is one of the largest collections with the highest number of type specimens of this insect order in Germany. Furthermore, the mayfly collection together with the caddisfly collection comprise some of the oldest voucher material of the entomological collection at the ZMH which was mainly collected at the end of the 19th century and in the first decades of the 20th century by the Hamburgian entomologist Dr. h.c. Georg Ulmer (1877–1963) (e.g., Weidner 1964b). Georg Ulmer was born in Hamburg on 5 March 1877. He was the oldest son of seven siblings. After his father died in 1889, he decided to become a school teacher. During his qualification time in the early 1890s, he began collecting insects in the vicinity of Hamburg (Ulmer 1964). Initially, he started to compile a beetle collection but since he observed a mass emergence of mayflies at the river Fulda near Kassel in 1898, he started to focus his collecting efforts and taxonomic studies on primarily aquatic insect groups like mayflies or caddisflies (Ulmer 1964). In 1899, he finished his teacher training and started his career as a board school teacher in Hamburg for 32 years until 1934. Beside his profession, he studied voluntarily the faunistics, biology, systematics and taxonomy of (e.g., Ulmer 1904a, 1926, 1939) and (e.g., Ulmer 1905, 1951), and other freshwater invertebrates (e.g., Ulmer 1901, 1902a). His first publication (Ulmer 1900) was followed by 175 scientific publications until his death on 15 January 1963 (Illies 1964). In the first years of his scientific career, he mainly studied the faunistics and taxonomy of caddisflies in the vicinity of Hamburg (e.g., Ulmer 1902b), or other localities in Germany (e.g. Ulmer 1904b, 1915). From the year 1905 on, he intensively started to study the material from foreign countries (including tropical regions), which has been lent to him by colleagues or other institutions (e.g., Ulmer 1905). Besides the , Ulmer worked on mayflies (e.g., Ulmer 1908, 1909, 1912). Between 1904 and 1943, he published thirty research papers on the systematics and taxonomy of (Kimmins 1963; Illies 1964) with the description of 111 species (Weidner 1964a). His studies on mayflies resulted in the fundamental revision of the Southeast Asian mayfly fauna with the description of numerous species (Ulmer 1939). During World War II (in 1943), the Zoological Museum Hamburg (that time located near the central railway station) was bombed and nearly all dry preserved material housed in the collections was destroyed and burned in the fire. Only material deposited in ethanol was transferred into the underground networks and survived the war (Weidner 1967). The valuable and collection of Georg Ulmer was kept in his private house outside of Hamburg. In 1964, the insect collection of Georg Ulmer was donated to the ZMH (Weidner 1964b) providing the majority of the recent voucher material of the ZMH in these two insect orders. Georg Ulmer was an outstanding specialist and his comprehensive studies on the faunistics and taxonomy of various groups of freshwater invertebrates formed the foundation of the recent taxonomy and systematics of those groups, especially and . In addition to Ulmer’s collection, the ZMH also stores primary type material described by other authors, such as Klapalek (1905a,b), Lestage (1930, 1935), Malzacher (1976, 2015, 2016), Müller-Liebenau (1971, 1974, 1980a, b, 1981), and Sartori (2008, 2014e). An interesting finding was the discovery of several paratypes of species described by Esben-Petersen (e.g. 1909, 1912a, b, 1913), which were not labelled as such but come from the type series, judging from the dates and places on labels. More information on these mayfly specialists can be found at http://www.ephemeroptera.de/inhaltsverz_deutsch/Galerie____/galerie____.html, a webpage maintained by Udo Jacob and Arne Haybach. The only attempt, prior to this type catalogue, has been made by Weidner (1962, 1964a, 1977), who published a list of type material deposited in ZMH, without any taxonomic remarks. At that time, 121 species were mentioned. This catalogue is accessible online (https://www.cenak.uni-hamburg.de/sammlungen/zoologie/entomologie/typenkatalog.html). Multiple subsequent taxonomic surveys utilizing a variety of different, high-resolution methods (morphological and molecular) led not only to the description of many new taxa, but also several supraspecific changes (e.g. Peters and Edmunds 1972; Puthz 1977; Braasch and Soldán 1986; Campbell and Suter 1988; Gillies 1990; Grant and Peters 1993; McCafferty and Waltz 1995; Lugo-Ortiz and McCafferty 1996, 1999; Sartori et al. 2008; Sartori 2014a, b, c.). The catalogue that follows is arranged alphabetically by species name, followed by the original combination, author(s) name and year of publication. For each entry, an actual nomenclature (valid names or synonymy) is given. The kind of type and the metamorphic stage of specimens are abbreviated as indicated in the table header. The present study is a completely independent and new work; data reported by Weidner were not copied as a base for this catalogue. This study is entirely original work, based on the first author’s specimen-by-specimen examination of the ZMH collections from December 2013 to June 2014. Recent publications by Malzacher (2015, 2016) on Ulmer’s collection were also added. List of type material of Hyatt & Arms, 1890, deposited at the Zoological Museum of Hamburg, with the state and metamorphic stage of materials, current nomenclature, family assignment, related literature and complementary information. Abbreviations in this list are as follows: Type: HT= Holotype; LT= Lectotype; PT= Paratype(s); PLT= Paralectotype(s); ST = Syntypes. (state): (1) Specimen conserved in alcohol; (2) Pinned dry specimen; (3) Specimen or part of it mounted on microscopic slide. Stage: F= Female imago; M= Male imago; N= Nymph; SF= Female subimago; SM= Male subimago. Complementary information: a. Holotype wrongly designated by Weidner (1962, 1964a); b. Holotype designated by Ulmer on the label, not in the publication; c. Holotype apparently not in Stockholm (see http://www.nrm.se/download/18.1cb760b014a762ca801342e9/1421068753427/EPHEMEROPTERA_list.pdf); d. Specimen not labelled, but comes from the type series; e. Not type material! The types are the nymphs and the adults were only described in 1925; f. Only fore- and hindlegs in ZMH; g. Only fore wing in ZMH and remains in Naturkunde Museum, Berlin; h. Only foreleg; i. Only hind wing in ZMH and remains in Naturkunde Museum, Berlin; j. Only one forewing in ZMH and remains apparently not in Stockholm (http://www.nrm.se/download/18.1cb760b014a762ca801342e9/1421068753427/EPHEMEROPTERA_list.pdf); k. Wrongly considered by Ulmer, as a syn. of (Hagen).
Table 1.

List of type material of Hyatt & Arms, 1890, deposited at the Zoological Museum of Hamburg, with the state and metamorphic stage of materials, current nomenclature, family assignment, related literature and complementary information. Abbreviations in this list are as follows: Type: HT= Holotype; LT= Lectotype; PT= Paratype(s); PLT= Paralectotype(s); ST = Syntypes. (state): (1) Specimen conserved in alcohol; (2) Pinned dry specimen; (3) Specimen or part of it mounted on microscopic slide. Stage: F= Female imago; M= Male imago; N= Nymph; SF= Female subimago; SM= Male subimago. Complementary information: a. Holotype wrongly designated by Weidner (1962, 1964a); b. Holotype designated by Ulmer on the label, not in the publication; c. Holotype apparently not in Stockholm (see http://www.nrm.se/download/18.1cb760b014a762ca801342e9/1421068753427/EPHEMEROPTERA_list.pdf); d. Specimen not labelled, but comes from the type series; e. Not type material! The types are the nymphs and the adults were only described in 1925; f. Only fore- and hindlegs in ZMH; g. Only fore wing in ZMH and remains in Naturkunde Museum, Berlin; h. Only foreleg; i. Only hind wing in ZMH and remains in Naturkunde Museum, Berlin; j. Only one forewing in ZMH and remains apparently not in Stockholm (http://www.nrm.se/download/18.1cb760b014a762ca801342e9/1421068753427/EPHEMEROPTERA_list.pdf); k. Wrongly considered by Ulmer, as a syn. of (Hagen).

TaxonOriginal nameAuthor(s), yearType (state)StageCurrent nomenclatureFamilyLiterature & Complementary information
alegrettae Massartella alegrettae Ulmer, 1943HT (2)M Massartella alegrettae Ulmer, 1943 Leptophlebiidae (i.)
ambiguum Pseudocloeon ambiguum Müller-Liebenau, 1982PT (1)N Liebebiella ambigua (Müller-Liebenau, 1982) Baetidae Waltz and McCafferty 1987
ammophila Oligoneuria ammophila Spieth, 1938PT (1)N Homoeoneuria ammophila (Spieth, 1938) Oligoneuriidae Edmunds and Allen 1957
anatolica Rhithrogena anatolica Kazanci, 1985PT (1)M Rhithrogena anatolica Kazanci, 1985 Heptageniidae (d.)
annandalei Polymitarcys annandalei Chopra, 1927ST (1)F Ephoron annandalei (Chopra, 1927) Polymitarcyidae Hubbard and Srivastava 1985
apicatum Cloeon apicatum Navás, 1933ST (2)M Cloeon navasi van Bruggen, 1957 Baetidae van Bruggen 1957; (d.)
balcanicus Metretopus balcanicus Ulmer, 1920HT (3)M Metreletus balcanicus (Ulmer, 1920) Ameletidae Puthz 1977a; (f.)
belgica Torleya belgica Lestage, 1917(PT) (2)M, F Torleya major (Klapalek, 1905a) Ephemerellidae (e.)
bengalensis Ecdyonurus bengalensis Ulmer, 1920HT, PT (2, 3)M, F, SM, SF Ecdyonurus bengalensis Ulmer, 1920 Heptageniidae (b.)
berneri Rheobaetis berneri Müller-Liebenau, 1974HT (1)N Heterocloeon berneri (Müller-Liebenau, 1974) Baetidae McCafferty and Provonsha 1975
bicorne Centroptilum bicorne Ulmer, 1909PT (1)F Afroptilum bicorne (Ulmer, 1909) Baetidae Gillies, 1990
biobionicum Deleatidium biobionicum Ulmer, 1938PT (2)F Meridialaris biobionica (Ulmer, 1938) Leptophlebiidae Peters and Edmunds 1972; (g.)
bishopi Platybaetis bishopi Müller-Liebenau, 1980aHT (1)N Platybaetis bishopi Müller-Liebenau, 1980 Baetidae
boettgeri Pseudocloeon boettgeri Ulmer, 1924ST (1, 2, 3)M, F Pseudocloeon boettgeri Ulmer, 1924 Baetidae
borneonia Epeorella borneonia Ulmer, 1939LT, PLT (1, 2)M, F, SF Epeorella borneonia Ulmer, 1939 Heptageniidae Sartori 2014c; (a.)
braueri Hagenulodes braueri Ulmer, 1920PT (1)M Hagenulodes braueri Ulmer, 1920 Leptophlebiidae
brenneriana Rhithrogena brenneriana Klapalek, 1905aPT (1)M Rhithrogena alpestris Eaton, 1885 (syn.) Heptageniidae Puthz 1975
brunneum Cloeon brunneum Esben-Petersen, 1909ST (1, 2)M, F Americabaetis peterseni (Hubbard, 1974) (syn. obj.) Baetidae Lugo-Ortiz and McCafferty 1999
caenoides Neoephemeropsis caenoides Ulmer, 1939LT, PLT (1, 3)M, F, N Potamanthellus caenoides (Ulmer, 1939) Neoephemeridae Bae and McCafferty 1998
camerunense Pseudocloeon camerunense Ulmer, 1920ST (1, 3)M, F Ophelmatostoma camerunense (Ulmer, 1920) Baetidae Gillies et al. 1990
canariensis Baetis canariensis Müller-Liebenau, 1971HT, PT (1)M, N Baetis canariensis Müller-Liebenau, 1971 Baetidae
cavum Cinygma cavum Ulmer, 1927PT (1)M Cinygmula cava (Ulmer, 1927) Heptageniidae Levanidova 1972
chinensis Baetis chinensis Ulmer, 1936ST (1)M, F Baetis chinensis Ulmer, 1936 Baetidae
chinensis Heptagenia chinensis Ulmer, 1920ST (2)M Heptagenia chinensis Ulmer, 1920 Heptageniidae (d.)
chinensis Siphluriscus chinensis Ulmer, 1920PT (3)M Siphluriscus chinensis Ulmer, 1920 Siphluriscidae (f.)
corsicus Ecdyonurus corsicus Esben-Petersen, 1912aPT (2)M Ecdyonurus corsicus Esben-Petersen, 1912 Heptageniidae Belfiore 1987; (k.)
costaricanus Leptohyphes costaricanus Ulmer, 1920HT (2, 3)F Tricorythodes costaricanus (Ulmer, 1920) Leptohyphidae Bamgardner 2008
crassinervis Polyplocia crassinervis Ulmer, 1939PT (2)SM Polyplocia campylociella Ulmer, 1939 (syn.) Euthyplociidae Demoulin 1953
croaticus Siphlonurus croaticus Ulmer, 1920PT (1, 3)M Siphlonurus croaticus Ulmer, 1920 Siphlonuridae
curvata Paraleptophlebia curvata Ulmer, 1927PT (1, 3)M Paraleptophlebia strandii (Eaton, 1901) (syn.) Leptophlebiidae Kluge 2009
deigma Liebebiella deigma Waltz & McCafferty, 1987HT, PT (1, 3)N Liebebiella vera (Müller-Liebenau, 1981) (syn.) Baetidae Kluge and Novikova 2011
difficilum Pseudocloeon difficilum Müller-Liebenau, 1982PT (1, 3)N Liebebiella difficila (Müller-Liebenau, 1982) Baetidae Waltz and McCafferty 1987
diptera Hagenulopsis diptera Ulmer, 1920ST (1)M, SM, SF Hagenulopsis diptera Ulmer, 1920 Leptophlebiidae
duporti Ephemera duporti Lestage, 1921PT (2)M Ephemera duporti Lestage, 1921 Ephemeridae
eatoni Rhithrogena eatoni Esben-Petersen, 1912aPT (2)F Rhithrogena eatoni Esben-Petersen, 1912 Heptageniidae (d.)
edmundsi Platybaetis edmundsi Müller-Liebenau, 1980bHT (1)N Platybaetis edmundsi Müller-Liebenau, 1980 Baetidae
ehrhardti Thraulus ehrhardti Ulmer, 1920ST (1, 2, 3)M, F Needhamella ehrhardti (Ulmer, 1920) Leptophlebiidae Dominguez and Flowers 1989
eloisae Derlethina eloisae Sartori, 2008PT (1)N Derlethina eloisae Sartori, 2008 Teloganodidae Sartori et al. 2008
feuernborni Pseudoligoneuria feuernborni Ulmer, 1939ST (1, 3)N Chromarcys magnifica Navás, 1932 (syn.) Oligoneuriidae Kluge 2004
flexus Siphlurus flexus Clemens, 1913PT (1)M Siphloplecton basale (Walker, 1853) (syn.) Metretopodidae McDunnough 1924
fluviatile Cloeon fluviatile Ulmer, 1920PT (1, 3)M, F Cloeon fluviatile Ulmer, 1920 Baetidae
formosana Ephemera formosana Ulmer, 1920PT (2, 3)M, F, SM, SF Ephemera formosana Ulmer, 1920 Ephemeridae
formosanus Chirotonetes formosanus Ulmer, 1912ST (1, 2, 3)M, SF Isonychia formosana (Ulmer, 1912) Isonychiidae Ueno 1931
fruhstorfferi Massartella fruhstorfferi Ulmer, 1943PTSF Massartella brieni (Lestage, 1924) (syn.) Leptophlebiidae Pescador and Peters 1990
fuegiensis Ameletus fuegiensis Lestage, 1935HT (1, 3)N Metamonius anceps (Eaton, 1885) (syn.) Nesameletidae Mercado and Elliott 2004
fulmeki Acentrella fulmeki Ulmer, 1939HT, PT (1, 3)M, F Labiobaetis fulmeki (Ulmer, 1939) Baetidae McCafferty and Waltz 1995
fusca Atalophlebia fusca Ulmer, 1920HT (1)M Koorrnonga fusca (Ulmer, 1920) Leptophlebiidae Campbell and Suter 1988; (c.)
gornostajevi Epeorus gornostajevi Tshernova, 1981PT (1)M Epeorus gornostajevi Tshernova, 1981 Heptageniidae
grandis Chirotonetes grandis Ulmer, 1913ST (1, 3)M Isonychia grandis (Ulmer, 1913) Isonychiidae Ulmer 1924
guranica Heptagenia guranica Belov, 1981PT (1)M Heptagenia guranica Belov, 1981 Heptageniidae
haenschi Euthyplocia haenschi Ulmer, 1942PT (2)M Euthyplocia haenschi Ulmer, 1942 Euthyplociidae
horai Potamanthellus horai Lestage, 1930HT (2)SM Potamanthellus amabilis (Eaton, 1892) (syn.) Neoephemeridae Bae and McCafferty 1998
hutanis Dudgeodes hutanis Sartori, 2008PT (1)N Dudgeodes hutanis Sartori, 2008 Teloganodidae Sartori et al. 2008
hyalinus Ecdyonurus hyalinus Ulmer, 1912ST (1, 2, 3)M, F, SM, SF Afronurus hyalinus (Ulmer, 1912) Heptageniidae Kang and Yang 1994
insularis Rhithrogena insularis Esben-Petersen, 1913PT (2)M, F Rhithrogena insularis Esben-Petersen, 1913 Heptageniidae (d.)
irina Cinygmula irina Tshernova & Belov, 1982PT (1)M Cinygmula irina Tshernova & Belov, 1982 Heptageniidae
jacobsoni Tricorythus jacobsoni Ulmer, 1913ST (1, 3)M, F Sparsorythus jacobsoni (Ulmer, 1913) Tricorythidae Sroka and Soldán 2008; (a.)
japonicus Chirotonetes japonicus Ulmer, 1920ST (1, 2, 3)M, F Isonychia japonica (Ulmer, 1920) Isonychiidae Ueno 1931; (a.)
javanicus Afronurus javanicus Ulmer, 1939HT (1)M Afronurus javanicus Ulmer, 1939 Heptageniidae
javanicus Baetis javanicus Ulmer, 1913ST (1, 2, 3)F Baetis javanicus Ulmer, 1913 Baetidae
jorgenseni Campsurus jorgenseni Esben-Petersen, 1912bST (1)M, F Campsurus jorgenseni Esben-Petersen, 1912 Polymitarcyidae
jorgenseni Cloeon jorgenseni Esben-Petersen, 1909ST (1, 3)M, F Americabaetis jorgenseni (Esben-Petersen, 1909) Baetidae Lugo-Ortiz and McCafferty 1999
karnyi Hagenulus karnyi Ulmer, 1939ST (1, 3)SM, SF Choroterpes (Euthraulus) karnyi (Ulmer, 1939) Leptophlebiidae Peters and Edmunds 1970
kraepelini Pseudocloeon kraepelini Klapalek, 1905bLT, PLT (1, 3)M, SM Pseudocloeon kraepelini Klapaleck, 1905 Baetidae Waltz and McCafferty 1985
krieghoffi Chitonophora krieghoffi Ulmer, 1920ST (2)M, S Ephemerella mucronata (Bengtsson, 1909) (syn.) Ephemerellidae Jacob 1974
lacuscoerulei Tasmanophlebia lacuscoerulei Tillyard, 1933PT (1)SF Tasmanophlebia lacuscoerulei Tillyard, 1933 Oniscigastridae (d.)
laminatum Deleatidum laminatum Ulmer, 1920PT (3)M Meridialaris laminata (Ulmer, 1920) Leptophlebiidae Peters and Edmunds 1972; (h.)
lamuriensis Thalerosphyrus lamuriensis Sartori, 2014eHT, PT (1, 3)N Thalerosphyrus lamuriensis Sartori, 2014 Heptageniidae
latifrons Cinygmula latifrons Tshernova & Belov, 1982PT (1)M Cinygmula hirasana (Imanishi, 1935) (syn.) Heptageniidae Kluge 1995
latus Tricorythus latus Ulmer, 1916PT (1, 2, 3)M Tricorythus latus Ulmer, 1916 Tricorythidae
lestagei Atalophlebiodes lestagei Ulmer, 1938ST (1, 3)N Meridialaris lestagei (Ulmer, 1938) Leptophlebiidae Dominguez et al. 2006
lieftincki Heptagenia lieftincki Ulmer, 1939HT, PT (1, 3)M, F Compsoneuria lieftincki (Ulmer, 1939) Heptageniidae Sartori 2014a
lobatus Ecdyonurus lobatus Ulmer, 1924ST (1, 2, 3)M, F, SF Afronurus lobatus (Ulmer, 1924) Heptageniidae Mol 1987
longilobata Leptophlebia longilobata Tshernova, 1928aPT (1, 3)M Paraleptophlebia longilobata (Tshernova, 1928) Leptophlebiidae Tiunova and Kluge 2016
longus Tricorythus longus Ulmer, 1916PT (1, 2, 3)M, F Tricorythus longus Ulmer, 1916 Tricorythidae
lucida Atalophlebia lucida Ulmer, 1920HT (1)M Thraulophlebia lucida (Ulmer, 1920) Leptophlebiidae Demoulin 1955; (c.)
macedonicus Rhoenanthus macedonicus Ulmer, 1920HT (2)M Neoephemera maxima (Joly, 1871) Neoephemeridae Bae and McCafferty 1998
maculipennis Thraulus maculipennis Ulmer, 1920HT (1)M Hermanella maculipennis (Ulmer, 1920) Leptophlebiidae Dominguez and Flowers 1989
magnificus Rhoenanthus magnificus Ulmer, 1920PT (1, 2, 3)M Rhoenanthus magnificus Ulmer, 1920 Potamanthidae
major Choroterpides major Ulmer, 1939HT, PT (1, 3)M, N Dilatognathus major (Ulmer, 1939) Leptophlebiidae Kluge 2012
malaisei Cinygma malaisei Ulmer, 1927PT (1, 3)M Cinygmula malesei (Ulmer, 1927) Heptageniidae Levanidova 1972
marginatus Thraulus marginatus Ulmer, 1913ST (1, 3)M, F Choroterpes (Euthraulus) marginatus (Ulmer, 1913) Leptophlebiidae Peters and Edmunds 1970
media Ephemera media Ulmer, 1936ST (2)M, F, SM, SF Ephemera media Ulmer, 1936 Ephemeridae (a.)
media Teloganopsis media Ulmer, 1939HT, PT (1, 3)M, N Teloganopsis media Ulmer, 1939 Ephemerellidae Ubero-Pascal and Sartori 2009; Sartori 2014f; (d.)
melli Thalerosphyrus melli Ulmer, 1926PT (2, 3)M, F Epeorus melli (Ulmer, 1926) Heptageniidae Zhou et al. 2007
mexicana Heptagenia mexicana Ulmer, 1920PT (3)M Maccaffertium mexicanum (Ulmer, 1920) Heptageniidae Wang and McCafferty 2004; (f.)
mjoebergi Euphyurus mjoebergi Ulmer, 1917PT (1, 3)M Ulmerophlebia mjoebergi (Ulmer, 1917) Leptophlebiidae Demoulin 1955
montium Thraulus montium Ulmer, 1943HT (2)M Traverella montium (Ulmer, 1943) Leptophlebiidae Allen 1973
multus Baetis multus Müller-Liebenau, 1984PT (3)N Labiobaetis multus (Müller-Liebenau, 1984) Baetidae McCafferty and Waltz 1995
nasuta Heptagenia nasuta Ulmer, 1939ST (1, 2)M, F Trichogenia nasuta (Ulmer, 1939) Heptageniidae Webb et al. 2006; (a)
natans Chankagenesia natans Buldowsky, 1935PT (1)M, F Chankagenesia natans Buldowsky, 1935 Palingeniidae
necatii Ecdyonurus necatii Kazanci, 1987PT (1)SM Electrogena necatii (Kazanci, 1987) Heptageniidae Kazanci 2001
necopinatus Baetis necopinatus Müller-Liebenau, 1981HT (1, 3)M Labiobaetis necopinatus (Müller-Liebenau, 1981) Baetidae McCafferty and Waltz 1995
nigrescens Baetis nigrescens Navás, 1932ST (2)M Baetis nigrescens Navás, 1932 Baetidae Müller-Liebenau 1971
nigrescens Tasmanophlebia nigrescens Tillyard, 1933PT (1)F, N Tasmanophlebia nigrescens Tillyard, 1933 Oniscigastridae (d.)
nigropunctata Caenis nigropunctata Klapalek, 1905bST (2, 3)F Caenis nigropunctata Klapalek, 1905 Caenidae Malzacher 2015
nigropunctatula Caenis nigropunctatula Malzacher, 2015HT, PT (1, 3)M, F Caenis nigropunctatula Malzacher, 2015 Caenidae
nitidum Centroptilum nitidum Ulmer, 1916ST (2)F Bugilliesia nitida (Ulmer, 1916) Baetidae Lugo-Ortiz and McCafferty 1996
novatus Baetis novatus Müller-Liebenau, 1981HT (3)N Baetis novatus Müller-Liebenau, 1981 Baetidae
obscurum Pseudocloeon obscurum Ulmer, 1913ST (3)M Pseudocloeon obscurum Ulmer, 1913 Baetidae Müller-Liebenau 1981
olivascens Baetis olivascens Ulmer, 1939HT, PT (1, 3)M, SM, SF Baetis olivascens Ulmer, 1939 Baetidae
operosus Baetis operosus Müller-Liebenau, 1984PT (3)N Labiobaetis operus (Müller-Liebenau, 1984) Baetidae McCafferty and Waltz 1995
orientale Pseudocloeon orientale Müller-Liebenau, 1982PT (1)N Liebebiella orientale (Müller-Liebenau, 1982) Baetidae Waltz and McCafferty 1987
orientalis Palingenia orientalis Chopra, 1927PT (2)M Palingenia orientalis Chopra, 1927 Palingeniidae
orientalis Raptobaetopus orientalis Müller-Liebenau, 1978HT (3)N Baetopus orientalis (Müller-Liebenau, 1978) Baetidae Jacob 1991
ornata Rhithrogeniella ornata Ulmer, 1939HT, PT (1, 3)M, F Rhithrogeniella ornata Ulmer, 1939 Heptageniidae Sartori 2014d
ornatum Procloeon ornatum Tshernova, 1928bLT (1, 3)M Procloeon ornatum Tshernova, 1928 Baetidae Sowa 1975
paradoxa Anagenesia paradoxa Buldowsky, 1935PT (1)M, F Anagenesia paradoxa Buldowsky, 1935 Palingeniidae
parvus Ecdyonurus parvus Ulmer, 1912ST (1, 2, 3)M, F Rhithrogena parva (Ulmer, 1912) Heptageniidae Sartori 2014b
patagonica Atalophlebia patagonica Lestage, 1931HT (1, 3)F Meridialaris patagonica (Lestage, 1931) Leptophlebiidae Peters and Edmunds 1972
pekinensis Caenis pekinensis Malzacher, 2016HT, PT (1,3)N Caenis pekinensis Malzacher, 2016 Caenidae
pekingensis Baetis pekingensis Ulmer, 1936HT, PT (1)M, F, SM, SF Alainites pekingensis (Ulmer, 1936) Baetidae Waltz et al. 1994
peruvianus Baetis peruvianus Ulmer, 1920ST (2)M, SM Andesiops peruvianus (Ulmer, 1920) Baetidae Lugo-Ortiz and McCafferty 1999
pescadori Jubabaetis pescadori Müller-Liebenau, 1980bHT (3)N Jubabaetis pescadori Müller-Liebenau, 1980 Baetidae
peterseni Leptohyphes peterseni Ulmer, 1920ST (1, 3)M, F, SM, SF Leptohyphes peterseni Ulmer, 1920 Leptohyphidae
petersi Rheobaetis petersi Müller-Liebenau, 1974HT (1)M, N Heterocloeon petersi (Müller-Liebenau, 1974) Baetidae McCafferty and Provonsha 1975
philippinensis Caenodes philippinensis Ulmer, 1924ST (1, 2, 3)M Caenis philippinensis (Ulmer, 1924) Caenidae Kluge 1985
pictipennis Ephemera pictipennis Ulmer, 1924ST (2)M, F Ephemera pictipennis Ulmer, 1924 Ephemeridae
powelli Lachlania powelli Edmunds, 1951PT (1)M, F, N Lachlania saskatschevanensis Ide, 1941 (syn.) Oligoneuriidae McCafferty 1996
proba Choroterpes proba Ulmer, 1939ST (1, 3)N Choroterpes proba Ulmer, 1939 Leptophlebiidae
prominens Habrophlebiodes prominens Ulmer, 1939HT (1, 3)M Habrophlebiodes prominens Ulmer, 1939 Leptophlebiidae
pseudorhodani Baetis pseudorhodani Müller-Liebenau, 1971HT, PT (1)M, N Baetis pseudorhodani Müller-Liebenau, 1971 Baetidae
puigae Teloganopsis puigae Ubero-Pascal & Sartori, 2009PT (1)N Teloganopsis puigae Ubero-Pascal & Sartori, 2009 Ephemerellidae
pulcher Afronurus pulcher Ulmer, 1930PT (1)SM, SF Afronurus collarti (Navás, 1930) (syn.) Heptageniidae Demoulin 1956
purpurata Ephemera purpurata Ulmer, 1920PT (2)M Ephemera purpurata Ulmer, 1920 Ephemeridae
ranauensis Caenis ranauensis Malzacher, 2015HT, PT (1, 3)M, F Caenis ranauensis Malzacher, 2015 Caenidae
rhenicola Caenis rhenicola Malzacher, 1976HT, PT (1)M Caenis pusilla Navás, 1913 (syn.) Caenidae Alba-Tercedor and Malzacher 1986
rossicus Ecdyonurus rossicus Tshernova, 1928bPT (1)M Kageronia fuscogrisea (Retzius, 1783) (syn.) Heptageniidae Puthz 1977b
sauteri Ephemera sauteri Ulmer, 1912PT (2)M, F Ephemera sauteri Ulmer, 1912 Ephemeridae
scotti Caenis scotti Ulmer, 1930PLT (1)M, F Caenis scotti Ulmer, 1930 Caenidae
seminiger Simothraulus seminiger Ulmer, 1939HT (1)M Simothraulus seminiger Ulmer, 1939 Leptophlebiidae Grant and Peters 1993
sexfasciata Atalophlebia sexfasciata Ulmer, 1916HT (1, 3)M Atalomicria sexfasciata (Ulmer, 1916) Leptophlebiidae Harker 1954; (j.)
sikorai Euthyplocia sikorai Vayssière, 1895PT (1, 3)N Proboscidoplocia sikorai (Vayssière, 1895) Euthyplociidae Demoulin 1966
sinensis Iron sinensis Ulmer, 1926PT (2, 3)M Epeorus sinensis (Ulmer, 1926) Heptageniidae Edmunds and Traver 1954
soror Baetis soror Ulmer, 1908ST (1)M, F Offadens soror (Ulmer, 1908) Baetidae Lugo-Ortiz and McCafferty 1998
sowai Rhithrogena sowai Puthz, 1972HT (2)M Rhithrogena sowai Puthz, 1972 Heptageniidae
sudanense Centroptilum sudanense Ulmer, 1916PT (1)M, SM Bugilliesia sudanense (Ulmer, 1916) Baetidae Lugo-Ortiz and McCafferty 1996
sumatrana Baetis sumatrana Ulmer, 1939ST (1, 3)M, F Baetis sumatrana Ulmer, 1939 Baetidae Müller-Liebenau 1981
sumatranus Ecdyonurus sumatranus Ulmer, 1939HT (1, 3)F Rhithrogena sumatrana (Ulmer, 1939) Heptageniidae Sartori 2014b
thienemanni Compsoneuriella thienemanni Ulmer, 1939LT, PLT (1, 2, 3)M, F, N Compsoneuriella thienemanni Ulmer, 1939 Heptageniidae Sartori 2014a
tibiale Cinygma tibiale Ulmer, 1920PT (1, 3)M Rhithrogena tibiale (Ulmer, 1920) Heptageniidae Tshernova and Belov 1982
tibialis Atopopus tibialis Ulmer, 1920PT (2, 3)M, F Atopopus tibialis Ulmer, 1920 Heptageniidae
traverae Rheobaetis traverae Müller-Liebenau, 1974HT (1)N Heterocloeon curiosum (McDunnough, 1923) (syn.) Baetidae McCafferty and Provonsha 1975
triangularis Siphlonurus triangularis Clemens, 1915PT (1)M, F, N Siphlonurus quebecensis (Provancher, 1878) (syn.) Siphlonuridae McDunnough 1925
truncatus Campsurus truncatus Ulmer, 1920PT (2)M, F Campsurus truncatus Ulmer, 1920 Polymitarcyidae
ulmeri Asionurus ulmeri Braasch & Soldán, 1986HT, PT (1, 3)N Asionurus ulmeri Braasch & Soldán, 1986 Heptageniidae
ulmeri Baetis ulmeri Müller-Liebenau, 1981HT (1, 3)M Labiobaetis ulmeri (Müller-Liebenau, 1981) Baetidae McCafferty and Waltz 1995
ulmeri Behningia ulmeri Lestage, 1930HT (1)N Behningia ulmeri Lestage, 1930 Behningiidae
ulmeri Dudgeodes ulmeri Sartori, 2008HT, PT (1, 3)M, SM, F, N Dudgeodes ulmeri Sartori, 2008 Teloganodidae Sartori et al. 2008 Sartori 2014f
ulmeri Trichogenia ulmeri Braasch & Webb, 2006HT, PT (1, 3)N Trichogenia ulmeri Braasch & Webb, 2006 Heptageniidae
ulmeriana Caenis ulmeriana Malzacher, 2015HT, PT (1, 3)M, F Caenis ulmeriana Malzacher, 2015 Caenidae
umbrata Teloganella umbrata Ulmer, 1939HT (1)SF Teloganella umbrata Ulmer, 1939 Teloganellidae
unguicularis Euphyurus unguicularis Ulmer, 1917PT (1)M Austrophlebiodes unguicularis (Ulmer, 1917) Leptophlebiidae Campbell and Suter 1988
unguiculatus Campsurus unguiculatus Ulmer, 1920ST (2)M Tortopsis unguiculatus (Ulmer, 1920) Polymitarcyidae Molineri 2010
ussingi Rhithrogena ussingi Esben-Petersen, 1910PT (2)M Rhithrogena germanica Eaton, 1885 (syn.) Heptageniidae Sowa 1971
verum Pseudocloeon verum Müller-Liebenau, 1982PT (1, 3)N Liebebiella vera (Müller-Liebenau, 1981) Baetidae Waltz and McCafferty 1987
virens Cloeon virens Klapalek, 1905bST (2, 3)M, F Cloeon virens Klapalek, 1905 Baetidae
vitellinum Centroptilum vitellinum Ulmer, 1939HT (2)M Centroptilum vitellinum Ulmer, 1939 Baetidae
winckleri Isonychia winckleri Ulmer, 1939ST (2)M, F, SF Isonychia winckleri Ulmer, 1939 Isonychiidae (a.)
wui Leptophlebia wui Ulmer, 1936ST (1)M, F Leptophlebia wui Ulmer, 1936 Leptophlebiidae
wui Caenis wui Malzacher, 2016HT, PT (1,3)N Caenis wui Malzacher, 2016 Caenidae
Missing material
asiaticum Cinygma asiaticum Ulmer, 1924ST (2)M Incertae sedis Heptageniidae Tshernova and Belov 1982
klapaleki Pseudocloeon klapaleki Müller-Liebenau, 1982PT (1)N Liebebiella klapaleki (Müller-Liebenau, 1982) Baetidae Waltz and McCafferty 1987
  6 in total

1.  Redescription of Paraleptophlebia falcula Traver 1934 with notes on status and composition of Paraleptophlebia Lestage 1917 and Neoleptophlebia Kluge 1997 (Ephemeroptera: Leptophlebiidae).

Authors:  Tatiana M Tiunova; Nikita Kluge
Journal:  Zootaxa       Date:  2016-04-06       Impact factor: 1.091

2.  The concept of Compsoneuria Eaton, 1881 revisited in light of historical and new material from the Sunda Islands (Ephemeroptera: Heptageniidae: Ecdyonurinae).

Authors:  Michel Sartori
Journal:  Zootaxa       Date:  2014-07-09       Impact factor: 1.091

3.  What is Ecdyonurus sumatranus Ulmer, 1939? A contribution to the knowledge of the genus Rhithrogena in the Oriental Region (Ephemeroptera, Heptageniidae).

Authors:  Michel Sartori
Journal:  Zootaxa       Date:  2014-05-26       Impact factor: 1.091

4.  The species of Thalerosphyrus Eaton, 1881 (Insecta, Ephemeroptera, Heptageniidae, Ecdyonurinae) in Java and Sumatra, with some comments on the diversity of the genus in the Oriental Realm.

Authors:  Michel Sartori
Journal:  Zookeys       Date:  2014-06-25       Impact factor: 1.546

5.  Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae).

Authors:  Michel Sartori
Journal:  Zookeys       Date:  2014-07-29       Impact factor: 1.546

6.  On the validity of Epeorella Ulmer, 1939 (Ephemeroptera, Heptageniidae) with general considerations on the Heptageniidae of the Sunda Islands.

Authors:  Michel Sartori
Journal:  Zookeys       Date:  2014-10-13       Impact factor: 1.546

  6 in total
  1 in total

1.  A Review of the Genus Cloeon from Chinese Mainland (Ephemeroptera: Baetidae).

Authors:  Xiaoli Ying; Wenjuan Li; Changfa Zhou
Journal:  Insects       Date:  2021-12-07       Impact factor: 2.769

  1 in total

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