Literature DB >> 26696765

Birds from the Azores: An updated list with some comments on species distribution.

Luís Md Barcelos1, Pedro R Rodrigues2, Joël Bried3, Enésima P Mendonça1, Rosalina Gabriel1, Paulo Alexandre Vieira Borges1.   

Abstract

BACKGROUND: An updated checklist of the Birds of the Azores is presented based on information compiled from Rodrigues et al. (2010) and from the websites, Azores Bird Club. (2014), Aves dos Açores (2014) Azores Bird Sightings (2014) and Vittery (2014), since 2010. NEW INFORMATION: The checklist has a total of 414 species, including 38 new species. Almost half of the species and subspecies that occur in the Azores have a Palearctic origin, the remaining ones being essentialy Nearctic and Holarctic species. São Miguel is the island with the highest number of bird species, followed by Terceira, Corvo and Flores islands.

Entities:  

Keywords:  Azores; birds checklist; species distribution

Year:  2015        PMID: 26696765      PMCID: PMC4678808          DOI: 10.3897/BDJ.3.e6604

Source DB:  PubMed          Journal:  Biodivers Data J        ISSN: 1314-2828


Introduction

Birds (: ) are some of the most iconic animals. They play important roles in the ecosystem, and since they are abundant and diverse in most urban and rural areas, humans have established a good long-lasting relationship with them (e.g., birdwatching, photography). However, amongst the ca 10.000 bird species which have been living on Earth since the appearance of modern humans, many species were regionally lost or extinct, or are endangered, especially on islands (Sax et al. 2002, Elphick et al. 2010, Rando et al. 2013, Alcover et al. 2015). Updating a list of birds from remote oceanic islands represents an important step towards an improved knowledge of colonization mechanisms and migratory patterns by animals and a contribution towards the conservation and management of insular taxa and their natural habitats. The Azores Archipelago is located in the North Atlantic Ocean, between 36°55’ and 39°43’ N, and 24°46’ and 31°16’ W, about 1500 km from mainland Europe and 1900 km from North America (Fig. 1). The Azores is a volcanic oceanic archipelago consisting of nine islands and several islets of recent volcanic origin (0.25 to 8.12 My old), which are spread over >600 km along a northwest-southeast transect (França et al. 2003). The oceanic distribution of these islands and, as a consequence, their relative positions, has led to the formation of the following groups: (i), a western group, comprising the islands of Flores and Corvo; (ii), a central group, with the islands of Faial, Pico, São Jorge, Terceira and Graciosa and (iii), an eastern group, made up of the islands of São Miguel and Santa Maria.
Figure 1.

Map of Azores islands, Portugal (Modified from Amorim 2005).

Due to its remoteness, the Azores archipelago was discovered and humans settled only at the beginning of the fifteenth century. The settlers described the islands as covered by dense native vegetation, with countless birds, especially seabirds (Frutuoso 2005a, Frutuoso 2005b, Frutuoso 2005c). Until then the archipelago was definitely a bird paradise, not only for seabirds that could use the vast coastal cliffs of the islands and islets to breed, but also for terrestrial birds which were widespread in the lush Azorean forests (Rodrigues and Cunha 2012). After human colonization started in 1439, several bird species suffered a sudden decrease in population size due to their use as food and oil source for human consumption, as well as to the introduction of predators and the destruction of their habitat (Rodrigues et al. 2010). As a consequence of the human impact, six flightless bird species are confirmed to be extinct in the Azores, namely one species (Rando et al. 2013) and five species (Alcover et al. 2015), but recent findings indicate that many more will be described soon (Josep Alcover and Juan Rando, pers. comm.). Despite the reduction of several bird populations, the Azores remain of great interest for ornithologists and are considered as one of the most popular destinations for European birdwatchers (Pereira 2010, Rodrigues and Michielsen 2010), all the more since they hold two endemic species: the Azores Bullfinch ( Godman, 1866) (Fig. 2), and Monteiro’s Storm Petrel ( Bolton et al. 2008) (Fig. 3), and 11 endemic subspecies. Besides, the Azores archipelago displays colonies of several seabird species of great importance in the European context, and it is also possible to observe pelagic birds and a large variety of migratory birds, some of which are very rare Nearctic vagrants to the western Palearctic (Rodrigues and Michielsen 2010, Barreiros et al. 2014).
Figure 2.

Azores Bullfinch ( Godman, 1866). Photo by PAV Borges (2008).

Figure 3.

Monteiro’s Storm Petrel ( Bolton et al., 2008). Photo by PH Silva (2014).

Rodrigues and Cunha (2011) resumed the ornithological history of the Azores, from the beginning of human settlement until recently, and the latest checklist of birds from the Azores archipelago was published in 2010 by Rodrigues et al., as part of the complete list of Azorean biota published by Borges et al. (2010). This list was based on an exhaustive review of the published literature (e.g., Le Grand 1993, Monteiro et al. 1996, Monteiro et al. 1999, Elias et al. 2003, Elias et al. 2004, Elias et al. 2005, Elias et al. 2006, Bried et al. 2007, Jara et al. 2007, Jara et al. 2008, Bolton et al. 2008, Imber 2008, Vittery 2014) but also on unpublished data following Rodebrand (2010) and Rodebrand S. & The Birding Azores Team (2010). This checklist totalled 384 species, distributed between three categories: breeding, non-breeding and potentially breeding (species reported as introduced and/or escapes from captivity and for which suitable nesting conditions exist in the Azores, but whose numbers are insufficient to allow the maintenance of self-sustaining populations in this archipelago). During the last years, and due to the growing interest of birdwatchers in the Azores avifauna, several dedicated websites have been developed, such as ABC – Azores Bird Club. (2014) (formerly Birding Azores) (http://www.birdingazores.com/), AdA – Aves dos Açores (2014) (http://azores.avesdeportugal.info/) and ABS – Azores Bird Sightings (2014) (http://azoresbs.weebly.com/ from January 2012 to February 2014 and http://azoresbirdsightings.blogspot.co.uk/ from August 2014 onward). These websites are presently used by birdwatchers as a tool to register new information about species occurrence and distribution in the Azores. The present paper updates the previous checklist (Rodrigues et al. 2010), with respect to species distribution within the archipelago and also by including new species to the archipelago. The updating is based on the sightings reports available on the websites mentioned above. We also perform a nomenclatural review of the Orders, Families, Species and Subspecies, following the 2014 IOC World Bird list (v 4.3) (Gill and Donsker 2015) published by the International Ornithologists’ Union (available at http://www.worldbirdnames.org/). Some additional notes on the distribution of the different species among the islands and their biogeographic origin are presented.

Materials and methods

Based on Rodrigues et al. (2010), we completed a full taxonomic revision using the IOC Bird list (Gill and Donsker 2015) as a reference to identify the changes which occurred in the taxonomic hierarchy and synonymies. Then, we updated the list of the species recorded in the Azores archipelago until the end of 2014, as well as their distribution within the archipelago. The species newly reported for the Azores are signalled as NSR (New Species Record). Most of these NSRs were reported after 2010, but there are also a few missing records prior to this date. The species previously reported by Rodrigues et al. (2010), but whose distribution has changed since then, are signalled with an asterisk (*). All the new information about species occurrence and distribution was compiled based on the birdwatching websites mentioned earlier, and relies on a large number of birdwatcher contributions. In order to make the data easier to consult, the main checklist (Checklist 1) includes all the breeding and non-breeding species together, including the escapes and/or introduced species which already have feral populations in the Azores. All the other escapes and/or introduced species were excluded from the main list, and can be consulted in Checklist 2. We considered the species with five our less records, in Azores, as Rare Species. This species are presented in a table (Suppl. material 3) with the number of sightings, number of individual recorded, and some notes. Besides the taxonomical information and the distribution among the Azorean islands (Fig. 4), we include information on: i) biogeographical origin, based on (Lomolino et al. 2010); ii) breeding status; iii) islands where breeding has been documented and iv) current protection status of the species according to the Azorean Decree DLR n.° 15/2012/A Governo dos Açores (2012) from 2nd April 2012, which transposes the European Directives (Birds), Conventions (Bern and OSPAR), Agreements (AEWA) and specific regional protection statuses (TOP100) to the regional legislation. The protection statuses are represented by a letter or code (corresponding to the directive, convention or law) and by a Roman numeral (corresponding to the annex) as follows: P – Priority species for conservation; A – Birds directive; B – Bern Convention; AEWA – African-Eurasian Waterbird Agreement; O – OSPAR Convention; T100 – 100 prioritary threatened species for management purposes in the European biogeographical region of Macaronesia (Martín et al. 2008); (X) – New species disaggregated from Madeiran Storm-petrel (Harcourt, 1851) and which is known to breed only on Praia and Baixo islets (Graciosa Island).
Figure 4.

Global species richness on the nine Azorean islands (COR= Corvo; FLO = Flores; FAI = Faial; PIC = Pico; GRA =Graciosa; SJG = São Jorge; TER = Terceira; SMG = São Miguel; SMR = Santa Maria) (Suppl. material 1).

Checklists

List of the non-breeding and breeding species from Azores

(Linnaeus, 1758)

Ecological interactions

Native status
Nearctic

Distribution

COR; FLO; GRA; TER*; SMG

Notes

Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Holarctic
Conservation status
A-IIA; AEWA COR; FLO; FAI; PIC; SJG; TER; SMG; SMR* Regular Wintering. Rodrigues et al. (2010) Gmelin, 1789 Nearctic COR; FLO; FAI; PIC; SJG; TER; SMG; SMR* Regular Wintering. Rodrigues et al. (2010) Gmelin, 1789 Nearctic COR; FLO; FAI; PIC; TER; SMG; SMR* Regular Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Holarctic A-IIA; AEWA COR*; FLO; FAI; TER; SMG; SMR* Regular Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic A-IIA; AEWA COR; FLO; FAI; PIC; GRA*; SJG; TER; SMG; SMR Regular Wintering. Rodrigues et al. (2010) Linnaeus, 1766 Nearctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic A-IIA; AEWA COR; FLO; FAI*; PIC; SJG*; TER; SMG; SMR Regular Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Holarctic A-IIA; AEWA COR (Occasional Breeder); FLO (Occasional Breeder); FAI; PIC; GRA*; SJG; TER; SMG (Occasional Breeder); SMR Regular Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic A-IIA; AEWA COR; FLO; FAI; PIC; TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) Brewster, 1902 Nearctic COR; FLO (Occasional Breeder); TER; SMG Regular Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Holarctic A-IIA; AEWA COR; FLO; FAI; TER; SMG Occasional Wintering. Rodrigues et al. (2010) (Scopoli, 1769) Holarctic PIC*; TER; SMG; SMR Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; TER*; SMG; SMR Occasional Wintering. Rodrigues et al. (2010) Baillon, 1834 Palearctic FLO; PIC*; TER; SMG; SMR Occasional Wintering. Rodrigues et al. (2010) (Latham, 1787) Palearctic COR; TER Occasional Wintering. Rodrigues et al. (2010) (Eyton, 1838) Nearctic FLO; FAI; PIC*; TER; SMG; SMR* Occasional Wintering. Rodrigues et al. (2010) (Donovan, 1809) Nearctic A COR; FLO; FAI; PIC; SJG; TER; SMG; SMR Regular Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FLO; TER; SMG Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FLO; FAI; PIC; SJG*; TER; SMG; SMR Regular Wintering. Rodrigues et al. (2010) (Linnaeus, 1761) Holarctic COR; FLO; FAI*; PIC*; SJG*; TER; SMG; SMR Occasional Wintering. Rodrigues et al. (2010) (Güldenstädt, 1770) Palearctic SMG Occasional Wintering. Rodrigues et al. (2010) (Müller, 1776) Holarctic COR; FLO; TER; SMG Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR*; FAI*; PIC*; GRA*; SJG; TER; SMG; SMR* Occasional Wintering. Rodrigues et al. (2010) (Bechstein, 1803) Palearctic COR*; FLO; SJG; SMG Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic PIC; GRA; TER Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic COR*; SJG; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic TER; SMG Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic FAI*; PIC*; SJG*; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Palearctic FAI; PIC; GRA; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Vieillot, 1816) Afro-tropical SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; FLO; SMG Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic FLO; FAI; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic FLO; FAI; SJG; TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Holarctic GRA; TER* Occasional Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Holarctic COR; FLO; FAI; PIC; TER; SMG; SMR Occasional Wintering. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic FLO; TER Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic COR; SMG Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic SMG Occasional Wintering. Rodrigues et al. (2010) (Pallas, 1764) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic FLO; TER*; SMG Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Seoane, 1894) Palearctic A-IIA PIC (Breeder); GRA*; TER (Breeder); SMG*; SMR (Breeder) Introduced. Rodrigues et al. (2010) Hartert, 1917 Palearctic A-IIB COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Native. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic SMR Occasional Migrant. New Azores Record (Brünnich, 1764) Holarctic COR*; FLO; FAI; PIC; GRA; TER; SMG; SMR Regular Wintering. Rodrigues et al. (2010) (Pontoppidan, 1763) Holarctic GRA; SMG Occasional Wintering. Rodrigues et al. (2010) (Temminck, 1828) Sub-Antarctic FAI; PIC Occasional Migrant. Rodrigues et al. (2010) (Jardine & Selby, 1828) Pantropical P; A-I; B-II; T100 FLO*; FAI*; PIC*; GRA*; SMG*; SMR (Breeder) Native. Rodrigues et al. (2010) (Cory, 1881) Palearctic P; A-I; B-II; T100 COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Native. Rodrigues et al. (2010) (Linnaeus, 1761) Holarctic COR; FLO; FAI; PIC; GRA*; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Giglioli & Salvadori, 1868) Pantropical COR; FLO*; FAI; PIC; GRA* Occasional Migrant. Rodrigues et al. (2010) (Nichols & Mowbray, 1916) Nearctic SMR Occasional and Non-Breeding. Rodrigues et al. (2010) Mathews, 1934 Palearctic GRA Occasional Migrant. New Azores Record (Mathews, 1934) (Salvadori, 1899) Palearctic P; A-I; B-II COR; FLO; FAI; PIC; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Kuhl, 1820) Nearctic FAI*; PIC; GRA Occasional Migrant. Rodrigues et al. (2010) Mathews, 1934 Palearctic GRA Occasional Migrant. New Azores Record (Schlegel, 1863) Pantropical FAI Occasional Migrant. Rodrigues et al. (2010) (Bonaparte, 1857) Palearctic P; A-I; B-II; O; T100 COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER*; SMG (Breeder); SMR (Breeder) Macaronesian Endemic. Rodrigues et al. (2010) (O'Reilly, 1818) Southern Atlantic A COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Southern hemisphere A COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) Lowe, 1921 Palearctic FLO; FAI; SMG Occasional Migrant. Rodrigues et al. (2010) (Brünnich, 1764) Holarctic P; A; B-II COR (Breeder); FLO (Breeder); FAI*; PIC*; GRA*; SJG*; TER*; SMG (Breeder); SMR (Breeder) Native. Rodrigues et al. (2010) (Kuhl, 1820) Sub-Antarctic and Antarctic COR*; FLO; FAI*; PIC; GRA*; TER*; SMG; SMR* Regular Migrant. Rodrigues et al. (2010) (Harcourt, 1851) Pantropical P; A-I; T100 COR*; GRA (Breeder); SJG*; SMG*; SMR (Breeder) Native. Rodrigues et al. (2010) (Vieillot, 1818) Holarctic COR*; FLO; FAI; PIC; GRA; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) (Swinhoe, 1867) Sub-tropical Pacific GRA Occasional Migrant. New Azores Record Bolton et al., 2008 Palearctic P; A-I (x) COR*; GRA (Breeder); SJG* Azores Endemic. Rodrigues et al. (2010) (Latham, 1790) Palearctic A FLO; FAI*; PIC; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic FLO; TER; SMG; SMR* Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic SMG Occasional Wintering. Rodrigues et al. (2010) (Boddaert, 1783) Holarctic FAI Occasional Wintering. New Azores Record Brehm, 1831 Holarctic FAI; TER; SMG Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; FLO; PIC; TER; SMG; SMR* Occasional Wintering. Rodrigues et al. (2010) (Pallas, 1764) Palearctic FLO; TER*; SMG Occasional Wintering. Rodrigues et al. (2010) Pallas, 1811 Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) Peters, 1930 Pantropical A; AEWA FAI; PIC; GRA (Occasional Breeder); SMG* Occasional Migrant. Rodrigues et al. (2010) Daudin, 1802 Pantropical COR*; FLO; FAI*; SJG*; TER*; SMG* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic PIC*; TER Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR; FLO; PIC; TER; SMG; SMR Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1766) Holarctic COR*; FLO*; FAI*; PIC*; GRA*; SJG*; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic SMR Occasional Migrant. Rodrigues et al. (2010) Gmelin, 1789 Nearctic COR; FLO; FAI; PIC; GRA*; SJG; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic A; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant; Regular Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Nearctic COR; FLO; FAI; PIC; SJG; TER; SMG Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) Linnaeus, 1766 Palearctic COR*; PIC*; TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Scopoli, 1769) Palearctic COR*; FAI*; PIC; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Rackett, 1813) Nearctic COR*; FLO; PIC*; GRA*; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic FLO; PIC; SJG; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic FLO; PIC; GRA*; SJG; SMG* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Palearctic P; A-I; B-II; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant; Regular Wintering. Rodrigues et al. (2010) (Bosc, 1792) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Molina, 1782) Nearctic FLO; FAI; PIC*; GRA*; SJG; TER; SMG Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Müller, 1776) Nearctic PIC; SMG Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Palearctic FLO; PIC; GRA; SJG; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; PIC; TER; SMR Occasional Wintering. New Azores Record (Linnaeus, 1758) Holarctic COR; FLO; FAI; PIC; GRA; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Mathews, 1914 Sub-tropical and Tropical Atlantic; Tropical-eastern Pacific COR*; FLO*; PIC*; TER*; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant; Regular Wintering. Rodrigues et al. (2010) Lesson, 1831 Pantropical FAI; GRA* Occasional Migrant. Rodrigues et al. (2010) (Boddaert, 1783) Pantropical GRA*; SMG Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Lesson, 1831) Nearctic COR; FLO; FAI; PIC; GRA*; TER; SMG; SMR Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic COR; PIC*; GRA*; TER; SMG; SMR Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic COR*; FLO; FAI; PIC; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Swann, 1919) Palearctic P; A; B-II; T100 COR; FLO; FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Azores Endemic. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic GRA; TER Occasional Migrant. New Azores Record (Pontoppidan, 1763) Holarctic COR; FAI; PIC*; GRA*; TER Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR*; FLO; TER* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic FLO; TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Palearctic COR; FLO; PIC*; TER; SMR Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic TER Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic FLO; SMG Occasional Migrant. Rodrigues et al. (2010) (Boddaert, 1783) Palearctic GRA; SMR Occasional Migrant. New Azores Record (Linnaeus, 1758) Palearctic COR; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FAI; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Gmelin, 1789 Nearctic FLO; FAI*; PIC; GRA*; TER; SMG Occasional Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic A-IIA; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER (Breeder); SMG (Breeder); SMR Regular Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A-IIB; AEWA COR*; FLO (Breeder); FAI*; PIC*; GRA*; SJG*; TER (Breeder); SMG (Breeder); SMR (Breeder) Native. Rodrigues et al. (2010) Thomson, 1842 Palearctic GRA*; TER*; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic FLO; FAI*; GRA*; SMG Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic FLO*; SJG; SMR Occasional Migrant. Rodrigues et al. (2010) (Scopoli, 1769) Palearctic FAI; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Palearctic COR; FLO*; PIC*; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Pallas, 1776) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic FLO Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic FAI; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic PIC; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic P; A-I; B-II; AEWA COR; FLO; FAI; PIC; GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Regular Wintering. Rodrigues et al. (2010) Pallas, 1773 Palearctic COR Occasional Migrant. New Azores Record Scopoli, 1786 Palearctic A; B-II; AEWA COR; FLO; TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Holarctic A; B-II; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR; FAI; SMR Occasional Migrant. Rodrigues et al. (2010) Bonaparte, 1825 Nearctic A COR; FLO; FAI; PIC; TER; SMG; SMR Regular Migrant; Regular Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Nearctic COR; FLO; FAI; PIC*; TER; SMG; SMR* (Occasional Breeder) Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR*; FLO; FAI; GRA; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Müller, 1776) Nearctic COR; FLO; FAI; PIC; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Holarctic TER; SMR* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic A-IIB; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant; Regular Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A-IIB; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FLO; FAI; PIC; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic A; B-II; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant; Regular Wintering. Rodrigues et al. (2010) (Bechstein, 1812) Nearctic COR; FLO; SMG Occasional Migrant. Rodrigues et al. (2010) (Horsfield, 1821) Nearctic TER Occasional Migrant. Rodrigues et al. (2010) (Pallas, 1764) Holarctic A; B-II; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant; Regular Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic P; A-I; B-II; AEWA FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Coues, 1861) Holarctic FLO*; SJG; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic A-IIB; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) (Pontoppidan, 1763) Palearctic A; B-II; AEWA COR; FLO; FAI; PIC; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Vieillot, 1819) Nearctic A COR; FLO; FAI; PIC; GRA*; SJG; TER; SMG; SMR* Regular Migrante; Occasional Wintering. Rodrigues et al. (2010) (Bonaparte, 1826) Nearctic PIC; SJG*; SMG Occasional Migrant. Rodrigues et al. (2010) (Brünnich, 1764) Holarctic COR; FLO; FAI; PIC; GRA; TER; SMG Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Cabanis, 1857) Nearctic COR*; PIC; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Vieillot, 1819) Holarctic COR; FLO; FAI; PIC; SJG; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) (Leisler, 1812) Palearctic COR; FLO; FAI; PIC; SJG; TER; SMG; SMR Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Vieillot, 1819) Nearctic COR; FLO; FAI; PIC; SJG; TER; SMG Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR; FLO; FAI; PIC; GRA*; SJG; TER; SMG; SMR* Regular Migrante; Occasional Wintering. Rodrigues et al. (2010) (Leisler, 1812) Palearctic TER; SMG* Occasional Migrant. Rodrigues et al. (2010) (Ord, 1825) Nearctic COR; FLO; FAI*; PIC; GRA; SJG; TER; SMG; SMR* Regular Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A-IIA; AEWA COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA*; SJG (Breeder); TER (Breeder); SMG (Breeder); SMR* Native. Rodrigues et al. (2010) (Latham, 1787) Palearctic SMR Occasional Migrant. New Azores Record (Gmelin, 1789) Nearctic COR; FLO; PIC; SJG; TER; SMG; SMR* Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Say, 1822) Nearctic COR; TER; SMG; SMR* Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic TER Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FLO*; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A-IIB; AEWA SJG; TER; SMG Regular Migrant. Rodrigues et al. (2010) (Brünnich, 1764) Palearctic COR; FAI; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic FLO*; PIC; GRA; TER; SMG; SMR Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) Latham, 1790 Nearctic COR; FLO; FAI*; PIC*; GRA; SJG*; TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A-IIB; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant; Regular Wintering. Rodrigues et al. (2010) Vieillot, 1817 Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic TER*; SMG Occasional Migrant. Rodrigues et al. (2010) (Vieillot, 1819) Nearctic COR; PIC; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic P; A-I;A-IIB; AEWA COR; FLO; FAI; PIC; GRA*; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic A-IIA; AEWA COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA*; SJG (Breeder); TER (Breeder); SMG (Breeder); SMR* Native. Rodrigues et al. (2010) (Pallas, 1764) Palearctic FAI; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR* Regular Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic FLO; PIC*; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic COR; FLO; PIC; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Gunnerus, 1767) Palearctic A-IIB; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic FLO; PIC; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic SJG; TER; SMG Occasional Migrant. Rodrigues et al. (2010) Wilson, 1813 Nearctic COR*; FLO; GRA*; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Bechstein, 1803) Palearctic FLO; PIC*; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic FLO; SJG; TER; SMG; SMR Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Vieillot, 1819) Nearctic COR; FLO; FAI; PIC; TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Güldenstädt, 1775) Holarctic TER Occasional Wintering. New Azores Record (Linnaeus, 1758) Pantropical FLO Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic FLO; TER; SMG* Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic COR; TER; SMG* Occasional Migrant. Rodrigues et al. (2010) (Pallas, 1811) Palearctic TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Temminck, 1815) Palearctic TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Ord, 1815) Nearctic FAI; PIC; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Palearctic A-IIB; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant; Regular Wintering. Rodrigues et al. (2010) (Gmelin, 1789) Holarctic PIC; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Pallas, 1776) Palearctic FAI; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Pallas, 1770) Holarctic FLO; TER; SMG* Occasional Migrant. Rodrigues et al. (2010) (Payraudeau, 1826) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Temminck, 1820) Palearctic FAI; PIC*; TER; SMG Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) Pontoppidan, 1763 Palearctic FLO; FAI; TER; SMG Occasional Migrant. Rodrigues et al. (2010) Richardson, 1831 Nearctic TER Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FLO; PIC; TER; SMG Occasional Wintering. Rodrigues et al. (2010) Ord, 1815 Nearctic A COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant; Regular Wintering. Rodrigues et al. (2010) Meyer, 1822 Holarctic COR; FLO; FAI; PIC; GRA; TER; SMG Occasional Wintering. Rodrigues et al. (2010) Brewster, 1883 Nearctic FLO; TER*; SMG; SMR* Occasional Wintering. Rodrigues et al. (2010) Gunnerus, 1767 Holarctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Holarctic A-IIB; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Wintering. Rodrigues et al. (2010) Dwight, 1922 Palearctic P; A-IIB; AEWA COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Azores Endemic. Rodrigues et al. (2010) Naumann, 1840 Palearctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Coues, 1862 Nearctic COR*; FLO; FAI; PIC; SJG*; TER; SMG Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; FLO; FAI; PIC; GRA; TER; SMG; SMR Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Wagler, 1831) Nearctic FLO; TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Swainson, 1837) Pantropical PIC; GRA (Occasional Breeder); SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Tropical Atlantic A; AEWA FAI*; GRA (Breeder); SJG*; TER*; SMG*; SMR Native. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic A; AEWA COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant; Regular Wintering. Rodrigues et al. (2010) Montagu, 1813 Holarctic P; A-I; B-II; AEWA; O; T100 COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Native. Rodrigues et al. (2010) Nuttall, 1834 Nearctic COR; FLO; TER; SMG* Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Holarctic P; A-I; B-II; AEWA; T100 COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Native. Rodrigues et al. (2010) Pontoppidan, 1763 Holarctic COR; FLO; FAI; TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Pallas, 1764) Palearctic FLO; SMG Occasional Migrant. Rodrigues et al. (2010) (Boddaert, 1783) Holarctic FLO; GRA; TER Occasional Migrant. Rodrigues et al. (2010) (Latham, 1787) Palearctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Sabine, 1819) Holarctic COR*; FLO; FAI; PIC*; GRA; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Vieillot, 1819 Holarctic FAI; PIC; GRA*; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Saunders, 1893 Antarctic FLO*; FAI*; GRA*; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) (Temminck, 1815) Holarctic COR; FLO; FAI; PIC; GRA; SJG*; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) (Brünnich, 1764) Palearctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Regular Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Holarctic FAI; TER; SMG Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FAI; PIC; GRA; SJG; TER; SMG Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic FLO; FAI; PIC; GRA*; TER; SMG; SMR Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic FLO*; FAI; PIC*; TER; SMG Occasional Wintering. Rodrigues et al. (2010) Gmelin, 1789 Palearctic A-IIA COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Introduced. Rodrigues et al. (2010) Hartert, 1905 Palearctic P; A-I COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Azores Endemic. Rodrigues et al. (2010) (Frivaldszky, 1838) Palearctic COR*; FAI*; PIC (Breeder); GRA*; TER (Breeder); SMG; SMR Native. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FLO; TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; FLO Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; FLO; FAI; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Wilson, 1811) Nearctic COR*; FLO; SMG Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic PIC*; SJG; TER*; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Scopoli, 1769) Holarctic SMG Occasional Migrant. Rodrigues et al. (2010) (Pontoppidan, 1763) Holarctic COR; FLO; FAI; PIC; GRA; SJG*; TER; SMG; SMR Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A; B-II COR*; FLO*; FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR* Native. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic COR*; FLO; FAI; PIC*; SJG* Occasional Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR*; GRA*; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Forster, 1771) Nearctic COR; FLO; FAI; PIC; SJG; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Gray, 1830) Afro-tropical SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic FLO; FAI; GRA; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Shelley, 1870) Palearctic FLO*; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; FLO; FAI; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FAI*; TER*; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic FLO; FAI; PIC; GRA; SJG; TER*; SMR* Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic FLO; PIC*; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; FAI Occasional Migrant. New Azores Record (Linnaeus, 1766) Nearctic COR Occasional Migrant. Rodrigues et al. (2010) Radde, 1863 Palearctic PIC Occasional Migrant. New Azores Record Linnaeus, 1758 Holarctic COR; FLO; TER; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) Géné, 1839 Palearctic SMG Occasional Migrant. New Azores Record Fleischer, 1818 Palearctic SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) Tunstall, 1771 Holarctic COR; FLO; FAI; PIC*; GRA; TER; SMG; SMR Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Holarctic AZO Occasional Migrant. New Azores Record Linnaeus, 1758 Nearctic TER; SMG Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR; SMG* Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR; FLO; FAI; PIC; GRA; TER; SMG; SMR Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) Linnaeus, 1766 Palearctic COR*; FLO; FAI; PIC; TER*; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Scopoli, 1769) Afro-tropical FAI*; TER*; SMG (Breeder) Introduced. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic SMG; SMR Occasional Migrant. New Azores Record Vieillot, 1808 Nearctic COR Occasional Migrant. New Azores Record Vieillot, 1808 Nearctic COR; SMG Occasional Migrant. Rodrigues et al. (2010) (Boddaert, 1783) Nearctic COR; FLO* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR; FLO; FAI*; TER* Occasional Migrant. Rodrigues et al. (2010) (Cassin, 1851) Nearctic COR; FLO* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR*; FLO; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic PIC; TER; SMG Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Vieillot, 1808 Nearctic COR; FLO Occasional Migrant. New Azores Record (Linnaeus, 1758) Holarctic FAI Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR; FLO; FAI; TER*; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Leisler, 1814) Palearctic SJG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Laxmann, 1769) Palearctic COR*; FAI; SJG; SMG* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FLO; FAI; PIC; GRA; SJG*; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Boddaert, 1783 Nearctic COR; FLO; TER Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR; FLO; FAI; PIC*; GRA; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Vieillot, 1817) Nearctic COR; TER Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; FLO Occasional Migrant. Rodrigues et al. (2010) (Vieillot, 1817) Palearctic SMR Occasional Migrant. New Azores Record (Linnaeus, 1758) Holarctic COR*; FLO; PIC; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Vieillot, 1808) Nearctic COR; FLO; SMG Occasional Migrant. Rodrigues et al. (2010) (Vieillot, 1817) Palearctic COR; FLO; FAI; PIC; GRA*; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Blyth, 1842) Palearctic PIC*; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FLO; SJG; TER*; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; TER Occasional Migrant. New Azores Record (Jerdon, 1845) Palearctic COR Occasional Migrant. Rodrigues et al. (2010) (Boddaert, 1783) Palearctic FLO Occasional Migrant. New Azores Record (Vieillot, 1817) Palearctic COR Occasional Migrant. New Azores Record Seebohm, 1883 Palearctic P; A; B-II SMG (Breeder) Azores Endemic. Rodrigues et al. (2010) Murphy & Chapin, 1929 Palearctic P; A; B-II FLO; FAI; PIC; GRA; SJG; TER Azores Endemic. Rodrigues et al. (2010) Vaurie, 1954 Palearctic P; A; B-II SMR (Breeder) Azores Endemic. Rodrigues et al. (2010) Alexander, 1898 Palearctic A; B-II COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Azores Endemic. Rodrigues et al. (2010) (Boddaert, 1783) Palearctic COR; FLO* Occasional Migrant. Rodrigues et al. (2010) (Pallas, 1764) Palearctic COR Occasional Migrant. New Azores Record (Bechstein, 1792) Palearctic FLO Occasional Migrant. New Azores Record Latham, 1787 Palearctic TER Occasional Migrant. New Azores Record (Linnaeus, 1758) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic FLO Occasional Migrant. New Azores Record Hartert, 1903 Palearctic A-IIB COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Azores Endemic. Rodrigues et al. (2010) (Pallas, 1811) Nearctic COR; FLO* Occasional Migrant. Rodrigues et al. (2010) (Lafresnaye, 1848) Nearctic COR; FLO Occasional Migrant. Rodrigues et al. (2010) (Nuttall, 1840) Nearctic COR; FLO Occasional Migrant. New Azores Record (Gmelin, 1789) Nearctic COR*; SMG Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Hartert, 1905 Palearctic A-IIB COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Azores Endemic. Rodrigues et al. (2010) Linnaeus, 1766 Nearctic COR Occasional Migrant. New Azores Record Temminck, 1820 Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) Brehm, 1831 Palearctic SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR; FLO; PIC*; GRA; TER; SMG; SMR Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR; SMG*; SMR* Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A; B-II FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Native. Rodrigues et al. (2010) (Pallas, 1764) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Bechstein, 1792) Palearctic TER Occasional Migrant. Rodrigues et al. (2010) (Pallas, 1764) Palearctic COR; FLO; SMR Occasional Migrant. New Azores Record (Gmelin, 1789) Nearctic COR; FLO; FAI; PIC; GRA; SJG; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A; B-II FLO; SMR Occasional Migrant. Rodrigues et al. (2010) (Temminck, 1829) Palearctic FLO Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1774) Palearctic TER*; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR* (Occasional Breeder); FLO; PIC; SMG Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR*; FLO; SJG; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Introduced. Rodrigues et al. (2010) (Linnaeus, 1766) Palearctic TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Afro-tropical TER (Breeder); SMG (Breeder) Introduced. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic SMG Occasional Migrant. Rodrigues et al. (2010) (Pallas, 1811) Palearctic COR; TER*; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic FLO; TER Occasional Migrant. Rodrigues et al. (2010) (Tunstall, 1771) Nearctic COR; FLO*; PIC; TER; SMG; SMR Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; TER Occasional Migrant. New Azores Record Linnaeus, 1758 Palearctic COR*; FLO; GRA; TER; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) Vaurie, 1957 Palearctic A; B-II COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Azores Endemic. Rodrigues et al. (2010) Pallas, 1776 Palearctic COR; SMR* Occasional Migrant. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic COR; PIC; TER*; SMG; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic COR; FLO; SMG Occasional Migrant. Rodrigues et al. (2010) (Holböll, 1843) Holarctic COR Occasional Migrant. Rodrigues et al. (2010) Tschusi, 1901 Palearctic A; B-II COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Macaronesian Endemic. Rodrigues et al. (2010) (Cabanis, 1851) Palearctic A; B-II FAI*; PIC*; TER (Breeder); SMG (Breeder); SMR* Introduced. Rodrigues et al. (2010) Pucheran, 1859 Palearctic A COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Azores Endemic. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic SMR Occasional Wintering. New Azores Record (Linnaeus, 1758) Palearctic TER*; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic FLO Occasional Migrant. New Azores Record Linnaeus, 1758 Holarctic SMG Occasional Migrant. Rodrigues et al. (2010) Godman, 1866 Palearctic P; A-I; T100 SMG (Breeder) Azores Endemic. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A COR (Breeder); FLO (Breeder); FAI (Breeder); PIC (Breeder); GRA (Breeder); SJG (Breeder); TER (Breeder); SMG (Breeder); SMR (Breeder) Macaronesian Endemic. Rodrigues et al. (2010) (Linnaeus, 1766) Palearctic TER Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic COR; FLO; SMG Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR; FLO Occasional Migrant. Rodrigues et al. (2010) (Wilson, 1811) Nearctic COR; FLO* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR; FLO* Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic COR*; SMG*; SMR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; FLO* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR; FLO*; TER Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic COR Occasional Migrant. Rodrigues et al. (2010) (Boddaert, 1783) Nearctic COR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR; FLO; SMG Occasional Migrant. Rodrigues et al. (2010) (Vieillot, 1809) Nearctic COR Occasional Migrant. New Azores Record (Linnaeus, 1766) Nearctic COR Occasional Migrant. New Azores Record (Wilson, 1811) Nearctic COR*; FLO Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR; FLO; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; FLO* Occasional Migrant. Rodrigues et al. (2010) (Forster, 1772) Nearctic COR; FLO; SMG Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic COR Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR Occasional Migrant. New Azores Record Olson & Reveal, 2009 Nearctic COR Occasional Migrant. New Azores Record (Linnaeus, 1758) Nearctic COR; FLO; SMR* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR; FLO; SMR* Occasional Migrant. Rodrigues et al. (2010) Pallas, 1776 Palearctic SMR Occasional Migrant. New Azores Record (Linnaeus, 1758) Nearctic FLO Occasional Migrant. Rodrigues et al. (2010) (Audubon, 1834) Nearctic COR Occasional Migrant. New Azores Record (Gmelin, 1789) Nearctic COR; FLO Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR; FLO; SMR* Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic COR; SMR* Occasional Migrant. New Azores Record (Forster, 1772) Nearctic COR; FLO; SMR* Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic COR; FLO; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Holarctic COR; FLO; FAI; PIC; GRA; TER; SMG; SMR Occasional Migrant; Occasional Wintering. Rodrigues et al. (2010) (Linnaeus, 1766) Nearctic COR; FLO; SMG; SMR* Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic COR; FLO Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic COR Occasional Migrant. Rodrigues et al. (2010) (Gmelin, 1789) Nearctic COR*; FLO; SMR* Occasional Migrant. Rodrigues et al. (2010)

List of potential breeding species from Azores

(Linnaeus, 1758) Palearctic PIC*; GRA; TER; SMG Occasional Migrant. Rodrigues et al. (2010) (Linnaeus, 1758) Afro-tropical SMG Introduced. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic FAI Introduced. Rodrigues et al. (2010) (Linnaeus, 1758) Palearctic A-IIA SMG Introduced. Rodrigues et al. (2010) Linnaeus, 1758 Palearctic AZO Introduced. Rodrigues et al. (2010) (Linnaeus, 1758) Nearctic SMG Introduced. Rodrigues et al. (2010) (Lichtenstein, 1823) Afro-tropical SMG Introduced. Rodrigues et al. (2010) (Müller, 1776) Afro-tropical TER Introduced. Rodrigues et al. (2010)

Analysis

The updated list of the bird species recorded in the Azores comprised 38 newly recorded species, increasing the total number of recorded species and subspecies to 414. These 414 taxa belong to 24 orders, 67 families, 202 genera and 404 species. Some of the species and genus names from the previous list were changed and some families were moved to other orders. The taxonomic revision results in the addition of four new orders to the checklist. São Miguel Island stands out as the richest island (299 spp. and sspp.), followed by Terceira (248 spp. and sspp.), Corvo and Flores (238 and 232 spp. and sspp., respectively) (Fig. 4). The islands of Corvo (western group) and Santa Maria (eastern group) were the ones with the highest increases in species number (plus 14,1% and 12,%, respectively). Almost half of the species and subspecies that occur in the Azores come from the Palearctic (n = 195, that is, 47% of spp and sspp) but the Nearctic is also well represented (n = 125 i.e., 30% of spp and sspp). The species and subspecies of Holarctic origin totalled 68 (i.e., 17% of spp and sspp) (Fig. 5). The (songbirds), (waders, skuas, gulls, terns and auks) and (swans, geese and ducks) are by far the most represented orders with 129, 111 and 42 taxa, respectively.
Figure 5.

Biogeographical origin of the species present in Azores archipelago; “Other” – marine regions (Indo-Pacific; Southern Atlantic; Sub-Antarctic; Antarctic; Tropical Atlantic) (Suppl. material 2).

Discussion

The most striking feature of this checklist is that it shows not only an impressive increase in the number of new species records for the Azores (36, that is, plus 8,7% in less than five years), but also an enlargement of the distribution of several species among the different Azorean islands. A simple explanation for this could be the increase in the number of observers performing birdwatching out of the “high season” (autumn and spring) for vagrant birds, and possibly also the creation of the “ dos Açores” website, whose aim is to provide information about all common species and also about rarities, even though is mostly aimed at a non-birdwatcher public. The situation of the Azores in the middle of the North Atlantic makes these islands a safe place for the birds taken away from their normal migratory routes by strong winds or by changes in wind direction (vagrants) (Liechti 2006). Vagrant birds have been observed using trans-oceanic ships to rest until they detect a landmass at close range and usually fly towards it (Lees and Gilroy 2009). The higher number of Nearctic species reported on Corvo Island could be related to the relative proximity of this island to the American continent, allowing birds to wander into it. Another explanation for this may be the combination of a very small area (17 km2) and the high observation pressure during the autumn, when several tens of birdwatchers visit the island. On the opposite side of the archipelago, Santa Maria Island, the closest island to the European continent, holds a higher proportion of Palearctic species. The comparatively low number of non-breeding species observed on Graciosa and São Jorge islands may be explained, at least partly, by a lower observation pressure than on the other islands. The native birds from the Azores provide good examples of insular speciation, since two endemic species and 11 endemic subspecies are known. Moreover, many more endemic terrestrial species have gone extinct (Rando et al. 2013, Alcover et al. 2015), possibly as a consequence of direct predation by man, the introduction of non-native predatory species (e.g., cats, rats, ferrets and weasels), and habitat degradation or loss. The current endemic species and subspecies are the survivors of almost 600 years of human occupation and major land-use changes. The recent population recovery of the Azores Bullfinch (Fig. 2) in the north-eastern part of São Miguel is considered as a good example of the best practices of LIFE conservation programs (Ceia et al. 2010). Another example of successful conservation measures comes from the rabbit eradication and habitat restoration on Praia islet (Graciosa Island), which resulted in a significant increase in the breeding numbers of Monteiro's Storm-petrels, Madeiran Storm-petrels and terns (Bolton et al. 2004, Bried et al. 2009, Bried and Neves in press). Species lists such as this one are important to draw attention to particular taxonomic groups, to serve as a baseline in long-term systematic monitoring, to elucidate patterns of species diversity and distribution, to elucidate information gaps about distribution, and can serve as a tool in educational projects. The currently ongoing initiatives concerning citizen ornithological science in the Azores will be of high value for the knowledge of the biology and phenology of migratory birds and will hopefully increase the pressure towards the definition and preservation of more Important Bird Areas (IBAs) in the archipelago. Global species richness on the nine Azorean islands Data type: Species distribution Brief description: Global species richness on the nine Azorean islands (COR= Corvo; FLO = Flores; FAI = Faial; PIC = Pico; GRA =Graciosa; SJG = São Jorge; TER = Terceira; SMG = São Miguel; SMR = Santa Maria). File: oo_62386.xlsx Biogeographical origin of the species present in Azores archipelago Data type: occurences Brief description: Biogeographical origin of the species present in Azores archipelago. “Other” refers to marine regions only represented once (Indo-Pacific; Southern Atlantic; Sub-Antarctic; Antarctic; Tropical Atlantic). File: oo_62383.xlsx List of Rare Species Data type: occurences Brief description: Species with five or less records and individuals counts, in the Azores, with some notes. File: oo_62653.xlsx
  2 in total

1.  Species invasions exceed extinctions on islands worldwide: a comparative study of plants and birds.

Authors:  Dov F Sax; Steven D Gaines; James H Brown
Journal:  Am Nat       Date:  2002-12       Impact factor: 3.926

2.  A new species of extinct scops owl (Aves: Strigiformes: Strigidae: Otus) from São Miguel Island (Azores Archipelago, North Atlantic Ocean).

Authors:  Juan Carlos Rando; Josep Antoni Alcover; Storrs L Olson; Harald Pieper
Journal:  Zootaxa       Date:  2013       Impact factor: 1.091

  2 in total
  3 in total

1.  Biota from the coastal wetlands of Praia da Vitória (Terceira, Azores, Portugal): Part 3 - Birds.

Authors:  Sofia Goulart; João Pedro Barreiros; Mariana R Brito; Sónia Santos; César M M Pimentel; Elisabete Nogueira; Paulo Alexandre Vieira Borges
Journal:  Biodivers Data J       Date:  2019-05-28

2.  Biota of coastal wetlands of Praia da Vitória (Terceira Island, Azores): Part 2 - Bryophytes.

Authors:  Rosalina Gabriel; César M M Pimentel; David Claro; Mariana R Brito; Javier Díaz-Castillo; Cecília Sérgio; Manuela Sim-Sim; Paulo Alexandre Vieira Borges
Journal:  Biodivers Data J       Date:  2019-07-01

3.  Insect ectoparasites from wild passerine birds in the Azores Islands.

Authors:  Lucie Oslejskova; Sarka Kounkova; Daniel R Gustafsson; Roberto Resendes; Pedro Rodrigues; Ivan Literak; Oldrich Sychra
Journal:  Parasite       Date:  2020-11-20       Impact factor: 3.000

  3 in total

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