Sarah E Goodwin1, Jeffrey Podos2. 1. Graduate Program in Organismic and Evolutionary Biology, University of Massachusetts Amherst, Amherst, MA, USA. 2. Graduate Program in Organismic and Evolutionary Biology, University of Massachusetts Amherst, Amherst, MA, USA Department of Biology, University of Massachusetts Amherst, Amherst, MA, USA jpodos@bio.umass.edu.