Shucen Wan1,2, Liwen Jiang3,4. 1. Molecular Biotechnology Program, School of Life Sciences, The Chinese University of Hong Kong, Shatin, New Territories, Hong Kong, China. sarabiwan@gmail.com. 2. School of Life Sciences, The Chinese University of Hong Kong, Shatin, New Territories, Hong Kong, China. sarabiwan@gmail.com. 3. Centre for Cell and Developmental Biology and State Key Laboratory of Agrobiotechnology, School of Life Sciences, The Chinese University of Hong Kong, Shatin, New Territories, Hong Kong, China. ljiang@cuhk.edu.hk. 4. School of Life Sciences, The Chinese University of Hong Kong, Shatin, New Territories, Hong Kong, China. ljiang@cuhk.edu.hk.