Maxime Derex1, Marie-Pauline Beugin1, Bernard Godelle1, Michel Raymond2. 1. University of Montpellier II, Place Eugène Bataillon, 34095 Montpellier Cedex 5, France. 2. 1] University of Montpellier II, Place Eugène Bataillon, 34095 Montpellier Cedex 5, France [2] CNRS, Institute of Evolutionary Sciences, CC 065, Place Eugène Bataillon, Montpellier 34095, France.