Analysis of Emerson enhancement under conditions where photosystem II is inhibited - Are the two photosystems indeed separated?

Photoacoustic measurements of photosynthetic oxygen evolution and its enhancement by addition of background far-red light (Emerson enhancement) were made on both intact and inhibited or stressed leaves. The extent of enhancement increased with the dehydration treatment and decreased with a mild heat treatment or with the addition of DCMU. It nevertheless persisted even at a very high degree of inhibition - a result which indicates high population ratio of single pairs of combined photosystem II and photosystem I units, functional in whole chain electron transport. This implies a restriction on the separation between photosystems I and II, in contrast to existing concepts.