Review of the odd chrysidid genus Loboscelidia Westwood, 1874 (Hymenoptera, Chrysididae, Loboscelidiinae).

The chrysidid genus Loboscelidia is reviewed and 11 new species are described, including Loboscelidia cinnamonea (Borneo), Loboscelidia fulgens (Viet Nam), Loboscelidia fulva (Thailand), Loboscelidia incompleta (India), Loboscelidia kafae (Borneo), Loboscelidia laminata (Viet Nam), Loboscelidia meifungae (Borneo), Loboscelidia nasiformis (Thailand), Loboscelidia nitidula (Thailand), Loboscelidia pecki (Viet Nam), and Loboscelidia sisik (Borneo). A key to males of the species of Loboscelidia is given.

Introduction

Loboscelidiinae is one of the smaller subfamilies in the family Chrysididae. The subfamily contains two genera, Westwood, 1874 and Kimsey, 1988. As of the publication of Kimsey and Bohart (1991), contained 30 species and one species. Since then four and two species have been added (Kojima and Ubaidillah 2003, Terayama et al. 1998, PageBreakXu et al. 2006). An additional 11 new species are described below. This study focuses on males and their characteristics as the systematics of the group is focused primarily on this sex due to the rareness of females in collections and the strong sexual dimorphism between males and females.

The subfamily is primarily south Asian with four northern Australian species. Every major south Asian island may have at least one endemic species of , and every new intensive collecting effort using Malaise traps or flight-intercept traps turns up new species. Thus, the loboscelidiine fauna appears to be largely under-sampled.

Loboscelidiines are among the most aberrant-looking and highly modified chrysidids, and as a result their actual family and even superfamily placement has varied considerably over the years. These are small-bodied, non-metallic brown wasps, with a superficial resemblance to members of the family Diapriidae (see Fig. 1). In fact Westwood (1874) originally described as a species of diapriid (Superfamily Proctotrupoidea). Ashmead (1902) then moved the genus to the family Figitidae (Superfamily Cynipoidea). Maa and Yoshimoto (1961) then moved the genus into its own family, Loboscelidiidae (Superfamily Bethyloidea). Finally, after making a detailed analysis of the metasomal morphology Day (1978) concluded that the group actually belonged in the family Chrysididae (Superfamily Chrysidoidea).

Figure 1.

Habitus photograph of male sp. in Queensland, Australia. Photo courtesy of Alex Wild; myrmecos.net.

Loboscelidiines are characterized by a number of unusual features (Figs 1, 2). The antennae insert horizontally on a shelf-like extension in the middle of the face (the shelf-like extension is termed the frontal projection below); the vertex is prolonged posteriorly into a neck-like projection fringed with ribbon-like setae; the pronotum is not freely hinged to the scutum and has a short line of ribbon-like setae along the anterolateral corner; the tegula is very large, covering both wing bases, and is held in place by a ridge on the mesopleuron; the mesopleuron is smooth without sculpturing, except for a shallow, trough-like scrobal sulcus in some species, the propodeum lacks a dorsal surface and has an ear-like lateral projection over the spiracle, and the forewing lacks a stigma, costal and subcostal veins.

Figure 2.

Diagram of lateral views of male . A Head, antenna removed. B Habitus of body. C Hindleg: (a) tubular part of femur width (b) femoral flange width (c) femoral flange length (d) femoral length. Abbreviations: HL = head length HW = head breadth M+Cu = media + cubital veins M = medial vein cu-a = cubital-anal cross vein R = radial vein R1 = first radial branch Rs = radial sector SS = scrobal sulcus.

Distinctions between and have been summarized in Kimsey and Bohart (1991). Briefly can be distinguished from by the forewing venation extending into the basal one-third to one half of the wing (considerably less than one-fourth in ), vertex convex or flat behind the ocelli, not sharply declivitous as in , and cervical expansion continuous with head, without discrete posterior expansion and with well-developed genal and cervical fringe. Cervical expansion basally constricted and shield-like posteriorly, with small discontinuous genal and cervical fringes in .

Members of the genus are strongly sexually dimorphic, which has led to confusion over generic placement and sex associations. The genus Maa & Yoshimoto, 1961 was erected for what turned out to be female (Day 1978). Males have five external metasomal segments and a long slender flagellum. Females are heavier bodied than the males, with a shorter, broader flagellum and an externally four-segmented metasoma. It's not clear how many characteristics are shared between the two sexes as fewer than 15% of specimens in collections are female and more than onePageBreak species may be present in a single locality. However, the sexes do seem to share some modifications of the wing venation (presence and shape, or absence of the medial vein), shape of the frontal projection, and presence or absence of the scrobal sulcus and notauli.

Little is known of the biology of the Loboscelidiinae. Specimens are rare in collections. However, this situation is probably more a reflection of collecting techniques used and sites visited than any indication of abundance. Malaise trapping in Thailand as part of the National Science Foundation funded TIGER project has yielded more than 100 specimens, more than all other museum holdings. The small number of female collected relative to males may be due to their differing habits. Males may be more frequently caught in traps because they tend to frequent low vegetation and the surface of leaf litter searching for females. Females may spend most of their time in cryptic situations, for example under bark or in the leaf litter, searching for hosts.

The morphology of the female ovipositor and mandibles closely resembles that of the Amiseginae, suggesting that loboscelidiines, like amisegines parasitize walking stick eggs. There is one report of an unidentified species of reared from the eggs of the phasmatid sp. (Riek 1970). It is also possible, given the structural modifications of the group, including the leg and antennal flanges, the very large tegula and the tegular clip that females at least may search for walking stick eggs in ant nests. Fouts (1922) suggested that the group is myrmecophilous based on the odd morphology. Walking stick eggs may be collected by ants because of the egg's strong resemblance to seeds.

Distribution map of the genus in south Asia and Australia.

Materials and methods

Specimens were borrowed from the following museums, and type repositories are indicated by the acronyms: AEI – American Entomological Institute, Gainesville, Florida); ANIC – Australian National Insect Collection; BME – Bohart Museum of Entomology, University of California, Davis, USA; BMNH – The Natural History Museum, London, UK; BPBM – Bishop Museum, Honolulu, Hawaii, USA; CAS – California Academy of Sciences, San Francisco, USA, CNC – Canadian National Insect Collection, Ottawa, Ontario, Canada; CSIRO, Canberra, Australia, Australian National Insect Collection; MNHN – Museum National d'Histoire Naturelle, Paris; QSBG – Chiang Mai Royal Botanical Garden, Chiang Mai, Thailand; ROM – Royal Ontario Museum, Toronto, Canada; UCR – Entomological Research Museum, University of California, Riverside, USA, and USNM – U.S. National Museum, Washington, D.C., USA.

Additional type repositories include: CASB - Institute of Zoology, etc.; Institute of Zoology, Beijing, China; MZB – Museum Zoologicum Bogoriense Cibinong, Indonesia; NMNS – National Museum of Natural Science, Taichung, Taiwan; OUMNH – Oxford University Museum of Natural History, Oxford, UK; QDPI – CSIRO Long Pocket Laboratories, Indooroopily, Queensland, Australia; SCAC – Hymenoptera Collection, SouthPageBreak China Agricultural University, Guangzhou, and ZFCL – Hymenoptera Collection, Zhejiang University, Hangzhou, China.

Morphological terminology follows that used by Kimsey and Bohart (1991) and is further described in Fig. 2. The hindwing lacks venation, so wing vein characters are only for the forewing. Wing veins are given in the text as abbreviations: Cu = cubital vein, cu-a = cubital-anal cross vein, M = medial vein, R = radial vein, Rs = radial sector, R1 = first radial branch. Scrobal sulcus refers to the transverse trough on the mesopleuron below the forewing ending in the scrobe adjacent to the metapleuron. The shape of the frontal projection is determined viewed in front view. It is considered triangular if the ventral angle of the projection ends in a point or the flat surface is less than one-tenth the length of the upper surface. The projection is considered rectangular if it is a true rectangle or rhomboid. Head length versus width is measured from the apex of the cervical extension to the furthermost point of the frontal projection and across the widest part of the head in lateral view. Antennal articles are measured at the point of greatest breadth and compared with the total length of the article. Wing veins are compared relative to the length of R1. Pronotal dimensions are measured from the medial length of the pronotum in dorsal view to the distance between the apices of the posterolateral angles. The length of a leg flange is measured from the basal joint to the apex of the segment along the ventral margin. The relative width of leg flanges are measured across the broadest part of the flange relative to the tubular part of the segment at the same point.

Species treatments

Fouts

http://species-id.net/wiki/Loboscelidia_antennata

Loboscelidia antennata

Material studied.

Singapore (USNM); Indonesia: West Kalimantan, Gunung Palung National Park (1 female, ROM); 2 female specimens were examined including the holotype.

Diagnosis.

The male of this species is unknown, but may very well prove to be the female of Fouts, based on the triangular frontal projection, flattened cervical expansion, curved medial vein and lack of a scrobal sulcus.

Kimsey

http://species-id.net/wiki/Loboscelidia_asiana

Loboscelidia asiana

Only the holotype was seen.

The most distinctive feature of is the presence of spatulate or leaf-like setae on the gena, a character shared only with (as in fig. 16). However, can be distinguished from by the submedially curved medial vein (nearly flat in ), scape striate and more than 3.5× as long as broad (smooth and less than 3× as long as broad in ) and no scrobal sulcus (present in ).

Figures 14–26

. Lateral view of male head, with basal antennal segments. 17–19. Front view of face with antennae removed 20 Dorsal view of thorax, with wings removed 21, 22 Dorsal view of pronotum scutum and tegulae 23–26 Forewings.

Krombein

http://species-id.net/wiki/Loboscelidia_atra

Loboscelidia atra

Only the holotype was seen.

This is one of several species with a well-developed, complete scrobal sulcus. A combination of features will separate from these other species, including the rectangular frontal projection (in lateral view), scape more than 3× as long as broad, cu-a vein less than half as long as R, Rs twice as long as R, and metanotum half as long as the scutellum.

Kimsey

http://species-id.net/wiki/Loboscelidia_australis

Figure 22

Loboscelidia australis

Australia: New South Wales, Queensland; two specimens were seen including the holotype.

This is one of three species (including and ), all Australian, that lack notauli (as in Fig. 22). can be distinguished from these by the submedially curved medial vein, rectangular frontal projection, pronotum with sharp lateral fold or ridge, flagellomere XI less than 3× as long as broad, and fore and midtibial flanges less than 0.5× as long as their respective tibial lengths.PageBreak

Fouts

http://species-id.net/wiki/Loboscelidia_bakeri

Figure 24

Loboscelidia bakeri

Malaysian Borneo, Sabah, Sandakan (2 males, USNM), Kinabalu National Park Poring Hot Springs (2 males including two syntypes, CNC, USNM).

Diagnosis. can be immediately distinguished from all other species by the distinctively dorsomedially up-domed propodeum. It is also one of four species, including , and that lack a medial vein (as in Fig. 24).

Fouts

http://species-id.net/wiki/Loboscelidia_brunnea

Loboscelidia brunnea

Malaysian Borneo, Sabah; only the holotype was seen.

Four species, , , and , have a strongly flattened cervical expansion. can be distinguished from theseby the extreme reduction of cu-a, Rs vein less than 3.5× as long as R, the legs coarsely striate, and hindtibial posterior margin essentially ecarinate.

Krombein

http://species-id.net/wiki/Loboscelidia_castanea

Loboscelidia castanea

Sri Lanka, Sabaragamuwa Prov.; only the holotype was seen.

This is one of the species with a complete scrobal sulcus and triangular frontal projection. It shares a long scape (more than 3× as long as broad) with one of these, . can be distinguished from these species and by a combination of characters, including cu-a less than 0.5× as long as R, A 0.6× as long as Cu+M, flagellomere I shorter than II, flagellomere XI more than 4× as long as broad, and the fore, mid and hindfemoral flanges as broad as the tubular part of the respective femora.

Maa & Yoshimoto

http://species-id.net/wiki/Loboscelidia_cervix

Figure 23

Loboscelidia cervix

New Britain: near Keravat only the holotype was seen.

This is one of two species, including , known from New Britain. Both have the notauli not reaching the posterior margin of the scutum (Fig. 23) and the frontal projection sublinear in front view. can be distinguished from by the shorter scape (2.6–2.8× as long as broad in , 3.0–3.1× in ), Rs more than 3× as long as R (less than 3× in ), cu-a longer than R (shorter in ) and partial scrobal sulcus (absent in ). The Australian species is the only other with long cu-a longer than R.

sp. n.

urn:lsid:zoobank.org:act:E5A2B8FA-4264-468B-B3A5-B52456903906

http://species-id.net/wiki/Loboscelidia_cinnamonea

Figures 4 25 37

Figures 4–13.

Lateral view of male head, with basal antennal segments.

Figures 36–46.

Lateral view of male fore (a), mid (b) and hind (c) legs.

Type material.

Holotype male: Thailand: Chiang Mai Pr., Doi Chiangdao NP, Pha Tang substation, 526 m, 19°24.978"N, 98°54.886"E, Malaise trap, 3-9/v/2008, Jugsu & Watwanich, T5802 (QSBG).

Paratypes (25 males): 3 males, same data as type; 1 male: Doi Chiangdao NP, 491 m, 19°24.278'N, 98°55.311'E, Malaise trap, 15–21/v/2008, Jugsu & Watwanich, T5815; 1 male: Doi Chiangdao NP, Pha Tang substation, 491 m, 19°24.278'N, 98°55.311'E, Malaise trap, 9–15/v/2008, Jugsu & Watwanich, T5812; 2 males: Doi Chiangdao NP, Huai Na Lao, 500m, 19°24.731'N, 98°55.315'E, YPT 5-6/v/2008, Jugsu & Watwanich, T5806; 1 male: Doi Chiangdao NP, Huai Na Lao, 500m, 19°24.731'N, 98°55.315'E, YPT 9-10/v/2008, Jugsu & Watwanich, T5811; 1 male: Doi Chiangdao NP, Huai Na Lao, 500m, 19°24.731'N, 98°55.315'E, YPT 4-5/v/2008, Jugsu & Watwanich, T5805; 2 males: Lampang Pr., Chae Son NP, Doi Laan, 18°51.815'N, 99°22.122'E, 1413 m, Malaise trap, 9-15/v/2008, Kwannui & Sukpeng, T5292; 1 male: Chae Son NP, 18°49.894'N, 99°28.354'E, 467 m, Malaise trap, 23–30/v/2008, Kwannui & Sukpeng, T5305; 1 male: Chae Son NP, 18°50.012'N, 98°28.656'E, 419 m, pan trap, 7-8/v/2008, Kwannui & Sukpeng, T5304; 3 males: Chae Son NP, 18°49.894'N, 99°28.354'E, 467 m, Malaise trap, 1-7/v/2008, Kwannui & Sukpeng, T5309; 1 male: Chanthaburi Pr., Khao Khitchakut NP, Khao Prabaht peak, 12°50.45'N, 102°09.81'E, 875 m, Malaise trap, 20–27/ii/2009, Suthida & Charoenchai, T4045; 1 male: Khao Khitchakut NP, Khao Prabaht peak, 12°50.45'N, 102°09.81'E, 875 m, Malaise trap, 6-13/ii/2009, SuthidaPageBreak & Charoenchai, T4039; 1 male: Trang Prov., Khaeochong Mt, 75 m, 7°33.038'N, 99°47.369'E, Malaise, 28/iv-2/c/2005; 2 males: near Nam Tock Ton Prov., Khoa Chong Mt.,140 m, 7°32.015'N, 99°47.036'E, iv/2005 and ii/2005; 1 male: Phetchabun Pr., Nam Nao NP, 16°43.695'N, 101°33.797'E, 921 m, Malaise trap, 5-12/v/2007, L. Janteab, T2657; 1 male: Kanchanaburi Pr., Khuean Srinagarindra NP, PageBreak14°38.136'N, 98°59.837'E, 210 m, pan trap, 21-22/viii/2008, Chatchawan, T3438; 1 male: Sakon Pr., Nakhon Phu Phan NP, 17°03.543'N, 103°58.452'E, 8-14/vii/2006, MT, W. Kongnara, T197; 1 male: Suphan Buri Pr., Khao Yai NP, Kong Geo waterfalls, 900 m, 30/vi/1990, J. Heraty, H90/108. Paratypes are deposited in QSBG and BME.

Additional non-type specimens (27) were seen from: Borneo: north, Tawa, Quoin Hill (1 male, BPB); Sabah: Kinabalu Nat. Park, Poring Hot Springs (4 males, CNC); Sarawak: sw Gunung Buda, 64 km s Limbang (BME); W. Kalimantan: Gunung Palung Nat. Pk. (3 males, ROM, BME); E. Kalimantan: Kac. Pujungan, Kayan-Mantarang Nat. Res. (1 male, ROM); West Java: Gede-Pangrango Nat. Park, Situ Gunung (2 males, ROM, BME); Sumatra: Aceh, Gunung Leuser Nat. Pk. (1 male, ROM); Malaysia: Selangor (1 male, UCR);Pahang: Kuala Tahan, Taman Negara Nat. Park (1 male, UCR); Malaya: 10 mi e Gombak (1 male, UCR); Thailand: Mae Hong Son, Namtok Mae Surin Nat. Pk (1 male, QSBG); Nakon Si Thammarat:Namtok Yong Nat. Pk. (1 male, QSBG); Phang Na: Khuraburi Dist. south end of Koh Res. (1 male, UCR); Trang: Forest Res. Sta. Khao Chong (1 male, UCR); Singapore (7 males, BPBM, UCR).

is most similar to , as both share an arched medial vein, rectangular frontal projection, complete notauli, without a scrobal sulcus and the cu-a vein reduced to a tiny stub or absent. It can be distinguished from by the more typical frontal projection, fore and midtibiae without discrete, measureable flanges, R1 obsolescent and Rs 3× or more as long as R.

Male description.

Body length 2.0–3.0 mm; forewing length 2.5–3.5 mm. Head (Fig. 4): length twice breadth in side view; eye asetose; frontal projection rectangular in front view; frons smooth, not microstriate; frons with low ridge extending from vertex along inner eye margin; vertex without transverse fovea, cervical expansion strongly curved in profile; gena without scale-like setae; scape smooth, length 3.9 breadth; flagellomere I length 2× breadth; flagellomere II length 2.3× breadth; flagellomere XI length 5× breadth. Mesosoma: pronotal length 0.9× breadth, without lateral carina, pronotum narrower than head width; scutum with notauli reaching posterior margin; scutellum with sublateral carina, without fine dense striae laterally; metanotum without medial ridge, impunctate laterally, 0.4× as long as scutellum; mesopleuron without scrobal sulcus; propodeum without transverse dorsal carina; legs (Fig. 37) smooth, polished; forefemoral flange 0.4 x femur length, flange maximum width equal to width of tubular part of femur; foretibial flange absent; midfemoral flange 0.6× femur length, flange maximum width 0.6× width of tubular part of femur; midtibial flange absent; hindfemoral flange 0.9× femur length, flange maximum width 0.7× width of tubular part of femur; hindtibial flange as long as tibia, flange maximum width 0.8× width of tubular part of tibia; hindtibia with two longitudinal carinae on posterior margin; hindcoxa without longitudinal carina on inner medial surface; forewing (Fig. 25) R1 length 0-0.2× R length; cu-a length 0.1× R length; Rs length twice R length; Cu+M length 0.4-0.6× A length; medial vein curved submedially. Color: body reddish brown to dark orange; wing membrane brown-tinted, with untinted areas adjacent to vein remnants; veins brown.

Female.

Unknown.PageBreak

Etymology.

The species name is Latin for brown as in the spice, cinnamon.

Fouts

http://species-id.net/wiki/Loboscelidia_collaris

Loboscelidia collaris

Indonesia: W. Kalimantan: Gunung Palung Nat. Pk (14 males, ROM; E. Kalimantan: Kac. Pujungan, Kayan-Matanrang Nat. Res. (3 males, ROM, BME); 38 km n alikpapan, Sambojal2 (1 male, ROM); Sumatra: Aceh, Gunung Leuser Nat. Park, Ketambe Res. Sta. (7 males, ROM, BME); Malaysia: Sabah, Mt. Kinabalu N.P., Poring Hot Spgs (2 males, CNC); Sarawak: Gunung Mulu National Park (4 males, BME, ROM); Selangor: 16 mi e Gombak, Univ. Malaya Forest (1 male, UCR); Singapore: (1 male, USNM), Timah Nat. Res. (1 male, CNC);; Thailand: Chaiyaphum,Tat Tone NP (1 male, QSBG); Trang: Near Nam Tock Tjon Prov., Khoa Chong Mt. (3 males, CNC); Phattalung Nam Tok Phrai Wan (1 male, UCR); 40 specimens were examined including the holotype.

This is another species with a complete scrobal sulcus and triangular frontal projection. Male can be distinguished from species with these traits by the combination of the pronotum with a sharp crease or ridge dorsolaterally, scape less than 3× as long as broad, flagellomeres I and II more than twice as long as broad, flagellomere XI 3.5–4.0× as long as broad, and the fore, mid and hindfemoral flanges as long as the femora.

Kieffer

http://species-id.net/wiki/Loboscelidia_defecta

Loboscelidia defecta

Viet Nam: Karyu Danar (1 male, BPBM), Thailand: Mae Hong Son Pr., Namtok Mae Surin NP (1 male, BME); Nakhon Si Pr., Thammarat Namtok Yong (1 male, QSBG); Surat Thani Pr., Khao Sok Np, Klong Morg unit (1 male, BME); Chiang Mai Pr., Doi Chiangdao NP (1 male, QSBG); Malaysia: Sarawak, Gunung Lulu National Park (1 male ROM); 6 specimens were seen that appear to fit the original description.

The types of are apparently lost. However, based on Kieffer's (1916a) illustration it is one of the species that lacks a cu-a vein. In the same paper Kieffer attributed and to a 1915 paper he gives in the 1916a paper as “Philippine J. Sci. v. 10 p?”, but there was evidently no paper published by Kieffer in 1915 in volume 10 of this journal. Instead, Kieffer and Kieffer were PageBreakpublished as new species one month after the 1916a paper (1916b). The 1915 date may have been a mistake on his part caused by delays in publication of the description paper in the Philippine Journal of Science.

urn:lsid:zoobank.org:act:229B3296-7FD3-49E4-8626-590CD8CDC23E

http://species-id.net/wiki/Loboscelidia_fulgens

Figs 5 26 38

Holotype male: Viet Nam: Tuyen Quang Prov., 360 m, Na Hang Reserve, 16–20 May 1997, FIT, S. B. Peck, 97-10 (CNC). Paratypes: 3 males same data as holotype; 1 male: 20-24 May 1997, 97-13; 1 male: 300 m, 97-17; 1 male: Ha Tinh, Huong Son, 450 m, 18°22'N, 105°13'E, 22 April-1 May 1998, L. Herman, LT (BME, CNC).

This is one of four species, including , and that completely lack a medial vein. can be separated from in males by the shorter Rs vein, 1.5× as long as R, versus twice as long in , and having well-developed tibial flanges, which are lacking in . can be immediately distinguished from by lacking the uniquely up-domed propodeum characteristic of .

Body length 1.5–2.0 mm; forewing length 2.0–2.5 mm. Head (Fig. 5): length 1.8× height in side view; eye asetose; frontal projection rectangular in front view; frons smooth, not microstriate; frons with low ridge extending from vertex along inner eye margin; vertex with transverse fovea, posterior expansion strongly curved in profile; gena without scale-like setae; scape striate, length 2.9× breadth; flagellomere I length 2× breadth; flagellomere II length 1.8× breadth; flagellomere XI length 3× breadth. Mesosoma: pronotal length 0.8× breadth, without lateral carina, narrower than head in dorsal view; scutum with notauli reaching posterior margin; scutellum with fine dense striae laterally; metanotum with medial ridge, impunctate laterally, 0.4× as long as scutellum; mesopleuron without scrobal sulcus; propodeum without transverse dorsal carina; legs (Fig. 38) smooth, polished; forefemoral flange 0.5× femur length, flange maximum width 0.6× width of tubular part of femur; foretibial flange 0.5× tibial length, flange maximum width 0.4 x width of tubular part of tibia; midfemoral flange absent; midtibial flange 0.6× femur length, flange maximum width 0.5× width of tubular part of tibia; hindfemoral flange 0.8× femur length, flange maximum width 0.6× width of tubular part of femur; hindtibial flange 0.8× femur length, flange maximum width 0.7× width of tubular part of tibia; hindtibia with two longitudinal carinae on posterior margin; hindcoxa without longitudinal cPageBreakarina on inner medial surface; forewing (Fig. 26) R1 length 0.4× R length; cu-a absent; Rs length 1.4× R length; Cu+M length 0.6× A length; medial vein present, flat medially. Color: body brown to reddish brown; wing membrane brown-tinted, untinted along vein remnants; veins brown.

Etymology. The species name, , refers to the shining integument (Latin, adj).

urn:lsid:zoobank.org:act:4719E8B5-6A56-4325-AEE0-4DD9DB50BC1D

http://species-id.net/wiki/Loboscelidia_fulva

Figs 6 27 39

Figures 27–35.

Male forewings.

Holotype male: Thailand: Nan Prov., Doi Phu Kha NP, 19°12'418"N, 101°4'809"E, 1326 m, MT, 15-22 Sept. 2007, Charoen & Nikom, T3217 (QSBG).

is one of five species with a straight medial vein, including , , and . It can be distinguished from and by having notauli, from by having the cu-a vein one-half or more as long as R and Cu+M as long as A, and from by the rectangular frontal projection, Rs about 3× as long as R and the scutellum coarsely areolate (smooth to longitudinally striate in ).

Body length 2.5 mm; forewing length 3.0 mm. Head (Fig. 6): length 1.6× height in side view; eye asetose; frontal projection rectangular in front view; frons with lateral ridge adjacent to eye margin; vertex without transverse fovea, posterior expansion strongly curved in profile; frons without carina or ridge extending from vertex along inner eye margin; gena without scale-like setae; scape smooth, length 2.7× breadth; flagellomere I length 1.6× breadth; flagellomere II length 1.7× breadth; flagellomere XI length 3.5–4.0× breadth. Mesosoma: pronotal length 0.8× breadth, with lateral carina, as wide as head in dorsal view; scutum with notauli reaching posterior margin; scutellum posteriorly coarsely rugose; metanotum with three medial ridges, impunctate laterally, 0.4× as long as scutellum; mesopleuron with scrobal sulcus; propodeum without transverse dorsal carina; legs (Fig. 39) smooth, polished; forefemoral flange 0.5× femur length, flange maximum width 0.5× width of tubular part of femur; foretibial flange 0.6× tibial length, flange maximum width 0.8× width of tubular part of tibia; midfemoral flange 0.6× femur length, flange maximum width 0.6× of tubular part of femur; midtibial flange 0.8× femur length, flange maximum width 0.7 of tubular part of tibia; hindfemoral flange 0.8× femur length, flange maximum width 0.6× of tubular part of femur; hindtibial flange as long as tibia, flange maximum width 1.2× of PageBreaktubular part of tibia; hindtibia with two longitudinal carinae on posterior margin; hindcoxa with longitudinal carina on inner medial surface; forewing (Fig. 27) R1 length 0.6× R length; cu-a length 0.6× R length; Rs length 3.1× R length; Cu+M as long as A ; medial vein flat. Color: body dark reddish brown; wing membrane brown-tinted, untinted along vein remnants.

Unknown.

The species name, , refers to the brown body color (Latin, f.).

Xu

http://species-id.net/wiki/Loboscelidia_guangxiensis

Loboscelidia guangxiensis

None; published distribution: China: Guangxi, Guangdong.

This is one of five species, including , , and , which have the medial vein partial or absent and cu-a less than 0.2× R or absent. It can be distinguished from these species by Rs more than twice as long as R and R1 more than 0.5× as long as R, flagellomeres I and II twice as long as broad, flagellomere XI less than 3× as long as broad and the hindtibial flange less than half as wide as the tubular part of the tibia.

Kojima

http://species-id.net/wiki/Loboscelidia_halimunensis

Loboscelidia halimunensis Kojima (in

None

Diagnosis. This is another of the species with a flat medial vein. and both lack a cu-a vein. The two species can be separated by the longer Rs vein in (more than 2× as long as R, versus less than 2× in ) and pronotum as long as broad or broader (longer than broad in ). Despite contacting the authors the type could not be located.

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http://species-id.net/wiki/Loboscelidia_incompleta

Figures 8 28

Holotype male: India: Tamil Nadu, Nilgiri Hills, v/1961, P. S. Nathan (CNC).PageBreak

The most distinctive and unique feature of this species is the medially incomplete medial vein. Among the species that lack a medial vein entirely, including , , and , can be distinguished by the Rs vein twice as long as R (1.5× or less in the other species). It does share the fore and midtibial flanges lacking as in .

Body length 2.5 mm; forewing length 3 mm. Head (Fig. 8): length 2× height in side view; eye asetose; frontal projection rectangular in front view; frons with lateral ridge adjacent to eye margin; vertex without transverse fovea, posterior expansion strongly curved in profile; frons without carina or ridge extending from vertex along inner eye margin; gena without scale-like setae; scape longitudinally striate, length 4× breadth; flagellomere I length 2.4× breadth; flagellomere II length 2.2× breadth; flagellomere XI length 3.2× breadth. Mesosoma: pronotal length 1.1× breadth, with lateral carina, nearly as broad as head; scutum with notauli reaching posterior margin; scutellum and metanotum smooth, polished, impunctate; metanotum one-third as long as scutellum propodeum without transverse dorsal carina; mesopleuron without scrobal sulcus; legs polished; forefemoral flange 0.2× femur length, flange maximum width 0.9× width of tubular part of femur; foretibial flange 0.6× femur length, flange maximum width 0.3× width of tubular part of tibia; midfemur without flange; midtibial flange 0.7× tibia length, flange maximum width 0.3× width of tubular part of tibia; hindfemoral flange 0.3× femur length, flange maximum width 0.7× width of tubular part of femur; hindtibial flange 0.7× as long as tibia, flange maximum width 0.5× width of tubular part of tibia; hindtibia with two longitudinal carinae on posterior margin; hindcoxa with/without longitudinal carina on inner medial surface; forewing (Fig. 28) R1 length 0.3× R length; cu-a length absent; Rs length 2.2× R length; Cu+M 0.5× as long as A; medial vein submedially curved, incomplete medially. Color: reddish brown; wing membrane brown-tinted, paler along vein remnants, veins brown.

The name refers to the medially interrupted medial vein of the forewing (Latin)

Kimsey

http://species-id.net/wiki/Loboscelidia_indica

Loboscelidia indica

India: Nilgiri; only the holotype was seen.

is one of two species described from India, including . It is also one of the dozen or so species with a scrobal sulcus and rectangular frontal projection. It can be distinguished from them by the combination of the Rs less than 3× as long as R, A as long or longer than Cu+M, scape less than 3× as long as broad, flagellomeres I and II twice or more as long as broad, and fore, mid and hindtibial flanges 1.5× or more as wide as the tibiae.

Kieffer

http://species-id.net/wiki/Loboscelidia_inermis

Loboscelidia inermis Kieffer 1916: 15. Syntype females (males?); Philippines: Mindanao, Butuan (MNHN, lost?).

No reliably identified specimens have been seen. However, according to Kieffer's (1916) illustration has a well-developed cu-a vein, unlike , which lacks cu-a, or cu-a is represented by a very short stub.

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http://species-id.net/wiki/Loboscelidia_kafae

Figures 9 29 40

Holotype male: Thailand: Chiang Mai Pr., Doi Phahompok NP, Mae Fang Hot spring, 569m, 19°57.961'N, 99°09.355'E, Malaise trap, 7–14/iv/2008, K. Seesom, T6085 (QSBG).

Paratypes (52 males): 2 males: same data as holotype; 1 male: 14-21/ix.2007, P. Wongchai, T6168; 2 males:, 7–14/viii/2007, P. Wongchai, T6144, 6111; 1 male: Doi Phaluang, 1449 m, 20°1'06N, 99°09.581'E, 21–28/ix/2007, P. Wongchai, T6165; 1 male: 28/iv-7/v/2008, K. Seesom, T6084; 1 male: Doi Chiangdao NP, 19°24.278'N, 98°55.311'E, 491 m, 18–25/ix/2007, Jugsu & Watwanich, T5696; 1 male: Doi Chiangdao NP, 19°24.419'N, 98°55.237'E, 549 m, MT, 21–28/viii/2007, Jugsu & Watwanich, T5676; 1 male: Doi Chiangdao NP, Pha Tang, 19°24.978'N, 98°54.886'E, 526 m, Malaise trap, 4–11/ix/2007, Jugsu & Watwanich, T5682; 1 male: Doi Chiangdao NP, 549 m, 19°42.419'N, 98°55.237'E, Malaise trap, 10–17/xii/2007, Jugsu & Watwanich, T5723; 1 male: Haui Na Lao, 500 m, 19°24.731'N, 98°55.315'E, Malaise trap, 15–21/v/2008, Jugsu & Watwanich, T5817; 1 male: Huai Nam Dang NP, 19°18.803'N, 98°36.396'E, Malaise trap, 21–28/ix/2007, Anuchart & Thawatchai, T5507; 1 male: Thung Buatong viewpoint, 19°17.6'N, 93°36.0'E Malaise trap, Anuchart & Thawatchai, 14–21/viii/2007, T5472; 1 male: Chiang Pr., Huai Nam Dang NP, Thung Buatong, 19°17.056'N, 98°36.029'E, Malaise trap, 21–28/viii/2007, Anuchart & Thawatchai, T5471; 1 male: Doi Chiangdao NP, 19°24.419'N, 98°55.237'E, 549 m, malaise trap, 14–21/viii/2007, Jugsu & Watwanich, T5673; 1 male: Kamphaeng Pr., Phet Mae Wong NP, 306 m, 16°02.233'N, 99°13.096'E, pan trap, 9–10/viii/2007, Srilopien & Phumirate, T3769; 1 male: Lampang Pr., Chae Son NP, 18°49.894'N, 99°28.354'E, 467 m, Malaise trap, 1–7/v/2008, Kwannui & Sukpeng, T5309; 1 male: 21–30/v/2008, T5305; 1 male: Chae Son NP, Doi Laan, 18°51.815'N, 99°22.122'E, 1413 m, Malaise trap, 9–15/v/2008, Kwannui & Sukpeng, T5292; 1 male: Kanchanaburi Pr., Khuean Srinagarinda NP, 14°38.123'N, 98°59.657'E, Malaise trap, Somboon & Daorueng, T3462; 1 male: 7–14/v/2009, T4747; 1 male: 201 m, 23–30/iv/2009, T4744; 1 male: 13–20/xi/2008,PageBreak Somboon & Daorueng, T4423; 1 male: 6–13/xi/2008, Somboon & Daorueng, T4420; 1 male; 14°38.312'N, 98°59.643'E, 210 m, Malaise trap, Somboon & Daorueng, T3465; 1 male: Huay Mae Kamint, 14°38.441'N, 98°58.889'E, 240 m, Malaise trap, 7–14/v/2009, Somboon & Daorueng, T4740; 1 male: Nakhon Si Thammarat Pr., Namtok Yong NP, 8°10.434'N, 99°44.508'E, Malaise trap, 8–15/vii/2008, 80 m, U. prai, KT3083; .1 male: 8°14.262'N, 99°48.289'E, Malaise trap, 21–28/vii/2008, 966m, Palboon, T3108; 1 male: 8°16.959'N, 99°39.149'E, Malaise trap, 22–29/vii/2008; 1 male: road to Khao Mhen, 150 m from Nern466, 8°16.959'N, 99°39.149'E, 499 m, Malaise trap, 8–15/vi/2008, S. Samnaokan, T3095; 1 male: Chaiyaphum Pr., Tat Tone NP, 16°0.792'N, 101°58.472'E, Malaise trap, 19–26/v/2007, Jaruphan & Budsawong, 648 m, 2575; 2 males: Petchaburi Pr., Kaeng Krachan NP, 12°47.831'N, 99°27.369'E, Malaise trap, 970 m, 8–15/viii/2008, Sirichai & Chusak, T4346; 1 male: 12°47.963'N, 99°27.188'E, Malaise trap, 5–12/ix/2008, Sirichai & Prasit, T4375; 1 male: 12°50.177'N, 99°28.098'E, Malaise trap, 735 m, 18–25/i/2009, Sirichai, T4406; 1 male: 12°48.107'N, 99°26.669'E, Malaise trap, 3–10/iv/2009, Sirichai, T4687; 1 male: 12°49.302'N, 99°22.263'E, Malaise trap, 254/iii-3/iv/2009, Sirichai, T4739; 1 male: 12°50.177'N, 99°20.688'E, Malaise trap, 735 m, 25/v-1/vi/2009, Sirichai, T5259; 2 males: Pa La-U waterfall, 12°32.154'N, 99°28.098'E, Malaise trap, 26/ix-3/x/2008, Akaradate & Thongbai, T4518; 1 male: 12°32.154'N, 99°28.098'E, Malaise trap, 4–11/xii/2008, Thongbai, T4553; 1 male: Pa La-U/Huai Palao Forest Unit 3, 12°32.149'N, 99°28.265'E, Malaise trap, 18–25/i/2009, Thongbai, T4566; 1 male: 12°32.149'N, 99°28.265'E, Malaise trap, 4–11/i/2009, Thongbai, T4562; 2 males: Phetchabun Pr., Nam Nao NP, 16°43.695'N, 101°33.797'E, 921 m, Malaise trap, 5–12/v/2007, L. Janteab, T2657; 2 males: 16°43.687'N, 101°33.797'E, 754 m, Malaise trap, 19–26/v/2007, N. Hongyothi, T2662; 1 male: Mae Hong Son Pr., Namtok Mae Surin NP, 228 m, 19°21.593'N, 97°59.254'E, Malaise trap, 11–18/xi/2007, M. Namadkum, T5930; 1 male: 19°20.616'N, 97°59.003'E, Malaise trap, 11–18/xi/2007, 334 m, A. Kamkhun, T5934; 1 male: Sakon Nakhon Pr., Phu Phan NP, 17°03.488'N, 103°58.497'E, Malaise trap, 8–14/vii/2006, S. Tongboonchai, T199; 1 male: Prachuab Khiri Khan Pr., Khao Sam Roi Yot NP, 12°13.417'N, 99°56.153'E, Malaise trap, 31/viii-7/ix/2008, Sorat, Yai & Amnad, T4078; 1 male: Bar Hua Tan Thaeo, 12°13.059'N, 99°58.384'E, Malaise trap, 2–9/xi/2008, Yai & Amnad, T4128; 1 male: Phitsanulok Pr., Thung Salaeng Luang NP, 16°52.046'N, 100°49.067'E, Malaise trap, 501 m, 16–23/iv/2007, Pongpitak, T5207 (BME, QSBG).

Additional non-type specimens were seen from Laos (Phongsaly Prov., Ban Sano Mai) (22 males, CNC, BME); Vientiane Prov., Ban Van Eue (1 male, BPBM); Malaysia: Malaya, 13 mi e Gombak (1 male, UCR); Sarawak: Gunung Mulu NP (1 male, ROM) and Borneo: West Kalimantan Gunung Palung Nat Pk. (14 males, BME, ROM) E. Kalimantan: Kac. Plujungan, Kayan Metarang Nat. Res. (1 male, ROM); Viet Nam: Tuyen Quang Prov., Na Hang Res. (2 males, CNC); Thailand: Phitsanulok Pr., Thyng Salaeng Luang (1 males, BME, QSBG); Kanchanaburi: Khuean Srinagarinda NP (1 male, QSBG); Suphanburi Pro., Pu Toei NP (1 male, QSBG).

L. kafae is one of the many species that have a submedially curved medial vein. Males have a short flagellomere I (less than twice as long as broad), which is PageBreakalso found in and . It can be distinguished from these two species by flagellomere XI 4× as long as broad (shorter in the other species), the fore and midtibial flanges as broad as the tibiae and the hindtibial flange twice as broad (narrower in various combinations in the other species).

Body length 2.0–2.5 mm; forewing length 2.5–3.0 mm. Head (Fig. 9): length 1.9× height in side view; eye asetose; frontal projection rectangular in front view; frons smooth; vertex without transverse fovea, posterior expansion convex in profile; frons with low ridge extending from vertex along inner eye margin; gena without scale-like setae; scape smooth, length 3× breadth; flagellomere I length 1.6× breadth; flagellomere II length 2× breadth; flagellomere XI length 4.5× breadth. Mesosoma: pronotal length 0.8× breadth, with sharp lateral fold; scutum with notauli reaching posterior margin; scutellum with fine dense striae sublaterally; metanotum with medial ridge, densely, finely striate on either side, one-third as long as scutellum; mesopleuron with scrobal sulcus; propodeum without transverse dorsal carina; legs (Fig. 40) smooth, polished; forefemoral flange 0.7× femur length, flange maximum width 0.9× width of tubular part of femur; foretibial flange 0.7× femur length, flange maximum width 1.2× width of tubular part of tibia; midfemoral flange 0.8× femur length, flange as wide as tubular part of femur; midtibial flange 0.7× tibial length, flange as wide as tubular part of tibia; hindfemoral flange 0.9× femur length, flange maximum width as wide as tubular part of femur; hindtibial flange as long as femur, flange maximum width 2× width of tubular part of tibia; hindtibia with two longitudinal carinae on posterior margin; hindcoxa without longitudinal carina on inner medial surface; forewing (Fig. 29) R1 length 0.5× R length; cu-a 0.5× R length; Rs length 2.6× R length; Cu+M length 0.5× A length; medial vein submedially curved. Color: body brown; wing membrane lightly brown-tinted along veins and vein remnants, veins brown.

Unknown.

The species name refers to the coffee brown coloration (Thai for coffee, noun).

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http://species-id.net/wiki/Loboscelidia_laminata

Figures 10 30 41

Holotype male: Viet Nam: Tuyen Quang Prov., 360 m, Na Hang Reserve, 16–20 May 1997, FIT, S. B. Peck, 97-10 (CNC). Paratypes (17 males): 6 males, same data as holotype; 6 males, 20-24 May 1997, rainforest, FIT 97-13; 5 males, 97-12 (BME, CNC).

most closely resembles as discussed under that species. However, can be distinguished by flagellomere II less than twice as long as broad, flagellomere XI less than 3.5× as long as broad, and the fore and midtibial flanges narrower than the respective tibiae.PageBreak

Body length 2.0–2.5 mm; forewing length 2.5–3.0 mm. Head (Fig. 10): length 1.8× height in side view; eye asetose; frontal projection rectangular in front view; frons with lateral ridge adjacent to eye margin; vertex without transverse fovea, posterior expansion shallowly curved in profile; frons with low ridge extending from vertex along inner eye margin; gena without scale-like setae; scape smooth, length 2.6× breadth; flagellomere I length 1.7× breadth; flagellomere II length 1.8× breadth; flagellomere XI length 4× breadth. Mesosoma: pronotal length 0.8× breadth, with/out lateral carina, nearly as wide as head in dorsal view; scutum with notauli reaching posterior margin; scutellum with fine dense striae; metanotum with three medial ridges, impunctate laterally; mesopleuron with scrobal sulcus; propodeum without transverse dorsal carina; legs (Fig. 41) coarsely/smooth, polished; forefemoral flange 0.6× femur length, flange maximum width 0.8× width of tubular part of femur; foretibial flange 0.6× femur length, flange maximum width 0.7× width of tubular part of tibia; midfemoral flange 0.6× femur length, flange maximum width 0.8× width of tubular part of femur; midtibial flange 0.8× femur length, flange maximum width 0.4× width of tubular part of tibia; hindfemoral flange 0.8× femur length, flange maximum width 0.9× width of tubular part of femur; hindtibial flange 0.9× as long as tibia, flange maximum width 1.1× width of tubular part of tibia; hindtibia with two longitudinal carinae on posterior margin; hindcoxa with longitudinal carina on inner medial surface; forewing (Fig. 30) R1 length 0.8× R length; cu-a length 0.5× R length; Rs length 3.2× R length; Cu+M 0.5× as long as A ; medial vein submedially curved. Color: dark brown to yellowish brown; wing membrane brown-tinted, untinted along vein remnants; veins brown.

The name refers to the large lamellae or flanges on the legs (Latin).

Kimsey

http://species-id.net/wiki/Loboscelidia_laotiana

Figure 19

Loboscelidia laotiana

Laos: Vientiane Prov, Ban Van Eue (2 males, BPBM, BME); Viet Nam: Fyan (1 male, BME); Malaysia: Sabah: Kinabalu Nat. Pk. (3 males, USNM); Indonesia: Sumatra, Aceh: Mt. Leuser Nat. Pk., Ketambe Res. Sta (1 male, ROM); 7 specimens were seen including the holotype.

is one of the species with a scrobal sulcus and a triangular frontal projection (Fig. 19). It can be distinguished from the others by the combination of the Rs 3× or more as long as R, scape striate and more than 3× as long as broad, flagellomeres I and II less than twice as long as broad, flagellomere XI less than 3× as long as broad, fore and midfemoral flanges as wide as the tubular part of the respective femora and the hindtibial flange twice as wide as the tubular part of the tibia.

Yao, Liu & Xu

http://species-id.net/wiki/Loboscelidia_levigata

Loboscelidia levigata

None.

is one of three species described from southeastern China, including and . It can be distinguished from these by the rectangular frontal projection, and in males R1 as long as R (shorter in and ) and Rs 3× as long as R, as opposed to 2.5× or shorter in and . It can be distinguished from other species by R1 reaching R at a right angle.

(Lin)

http://species-id.net/wiki/Loboscelidia_maai

Scelidoloba maai

Loboscelidia artigena

Loboscelidia latigena

None.

This is one of four species, including , and , with the cervical expansion of the vertex flat in profile. males can be distinguished from these species by having the scape less than 2.5× as long as broad, the presence of a scrobal sulcus, and the tibial flanges wider than the tubular part of the respective tibiae.

Kimsey

http://species-id.net/wiki/Loboscelidia_maculata

Loboscelidia maculata

Australia: Queensland: 7 km sw Bellenden (1 male, ANIC); Mossman Gorge (2 males, CNC); 3 specimens were seen including the holotype.

This is one of the five species with a medially flat medial vein as discussed under . Of these, only and have been described from Australia.PageBreak can be distinguished from by the lack of notauli (shared with ), and the fore and hindtibial flanges twice as wide as the tubular part of the respective tibiae (narrower in and ).

Fouts

http://species-id.net/wiki/Loboscelidia_maculipennis

Figures 11 17, 20 41

Loboscelidia maculipennis

Loboscelidia carinata

Singapore: coll. Baker (1 make, BME), Sungei Bulch (1 male, BME); Indonesia: W. Kalimantan: Gunung Palung Nat. Pk. (6 males, ROM, BME); E. Kalimantan: Kac. Pujungan, Kayen-Mentarang Nat. Res (1 male, ROM), Sumatra: Aceh, Mt. Leuser (1 male, ROM); 12 males were seen including the holotypes of and .

Diagnosis. This is one of four species with a strongly flattened cervical expansion (Fig. 11) as discussed under . males can be distinguished from the other three by cu-a as long as R, Rs vein 4× or longer than R, leg integument smooth (Fig. 41), and hindtibial posterior margin with 2 parallel carinae.

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http://species-id.net/wiki/Loboscelidia_meifungae

Figures 12 30 42

Holotype male: Borneo: Sarawak, sw Gunung Buda, 64 km s Linbang, 4°13'N, 114°56'E, 8–15 Nov. 1996, MT, Heydon & Fung (BME). Paratypes (44): 10 males, same data as holotype; 10 males: 16–28 Nov. 1996; 11 males: 22-28 Nov. 1996, MT, Heydon & Fung; 1 male: November 1996, Heydon & Fung; 1 male: 18 Nov. 1996, Heydon & Fung; 1 male: 23 Nov. 1996; Heydon & Fung; 1 male: Buda Camp, sw Gunung Buda, 64 km s Linbang, 4°11'N, 114°56'E, 4 Nov. 1996, MT, Heydon & Fung; 4 males: Malaysia: Sabah, Kinabalu NP, 800m, Poring Hot Springs Langanan Creek, 22/viii/1988, A. Smetana, B-138; 1 male: Poring Hot Springs, 520 m, 9/v/1987, A. Smetana; 1 male: 480–510 m, 30.viii/1988, A. Smetana, B163; 1 male: 510 m, 13/v/1987; 1 male: Kipungit Creek, 550 m, 26/viii/1988, A. Smetana; 1 male: Liwagu River Trail, 1550 m, 12/viii/1988, A. Smetana, B107 (BME, CNC).

This species belongs in the group of species having a flat medial vein and notauli, including and . It can be distinguished from other members PageBreakof the group by the triangular frontal projection, presence of a scrobal sulcus, cu-a present (shared with ) and midtibial flange absent.

Body length 2.0–4.0 mm; forewing length 2.5–4.5 mm. Head (Fig. 12): length 1.8–2.0× height in side view; eye asetose; frontal projection triangular in front view; frons smooth to microstriate; vertex without transverse fovea, posterior expansion strongly curved in profile; frons without discrete carina or ridge extending from vertex along inner eye margin; gena without scale-like setae; scape with some striae, length 2.1–2.5× breadth; flagellomeres I and II length twice breadth; flagellomere XI length 4× breadth. Mesosoma: pronotal length 0.7–0.8× breadth, with lateral carina; scutum with notauli reaching posterior margin; scutellum with sublateral carina, with fine dense striae laterally; scrobal sulcus represented by series of pits; metanotum with medial ridge, impunctate laterally; propodeum without transverse dorsal carina; legs (Fig. 42) smooth, polished; forefemoral flange 0.5–0.7× femur length, flange maximum width 0.8-1.0× width of tubular part of femur; foretibial flange 0.6–0.9× femur length, flange maximum width 1.0–1.5 x width of tubular part of tibia; midfemoral flange 0.7–0.9× femur length, flange maximum width as wide as tubular part of femur; midtibial flange 0.7× femur length, flange maximum width 1.2× width of tubular part of tibia; hindfemoral flange 0.9× femur length, flange maximum width as wide as tubular part of femur; hindtibial flange 0.9× femur length, flange maximum width 1.7× width of tubular part of tibia; hindtibia with two longitudinal carinae on posterior margin; hindcoxa without longitudinal carina on inner medial surface; forewing (Fig. 30) R1 length 0.5–0.7× R length; cu-a length 0.4–0.5× R length; Rs length 2.5–3.0× R length; Cu+M length 0.7–0.9× A length; medial vein submedially curved. Color: body brown to reddish brown; wing membrane brown-tinted, paler along vein remnants.

Unknown.

This species is named after Mei Lin “Stella” Fung one of the collectors.

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http://species-id.net/wiki/Loboscelidia_nasiformis

Figures 13 31 43

Holotype male: Thailand: Petchaburi Prov., Kaeng Krachan NP, Pa La-U/Huai Palao Forest Unit 3, 12°32'149"N, 99°28'265"E, Malaise trap, 4-11/i2009, Thongbai, T4562 (QSBG).

The most distinctive and unusual feature of this species is the greatly elongate and nose-like frontal projection, which makes the head nearly 3× as long as broad in lateral view. Otherwise, is closest to , with an arched medial vein, rectangular frontal projection (albeit greatly elongate in ), complete notauli, cu-a reduced to a tiny stub or absent, and no scrobal sulcus. Other than the elongate frontal projection, can be separated from by the presence of fore and midtibial flanges (absent in ).PageBreak

Body length 2 mm; forewing length 2.5 mm. Head (Fig. 13): length 2.9× height in side view; eye asetose; frontal projection nasiform; frons smooth; vertex without transverse fovea, posterior expansion strongly curved in profile; frons without carina or ridge extending from vertex along inner eye margin; gena without scale-like setae; scape smooth, without striae, length 3.7× breadth; flagellomeres I and II length 2.2× breadth; flagellomere XI length 3.6× breadth. Mesosoma: pronotal length 0.9× breadth, without lateral carina; scutum with notauli reaching posterior margin; scutellum without sublateral carina, smooth laterally; metanotum without medial ridge, impunctate laterally; propodeum without transverse dorsal carina; legs (Fig. 43) smooth, polished; forefemoral flange 0.5× femur length, flange as wide as tubular part of femur; foretibial flange 0.6× femur length, flange maximum width 0.4× width of tubular part of tibia; midfemoral flange 0.5× femur length, flange maximum width 0.4× width of tubular part of femur; midtibial flange 0.6× femur length, flange maximum width 0.6× width of tubular part of tibia; hindfemoral flange 0.8× femur length, flange maximum width 0.5× width of tubular part of femur; hindtibial flange 0.8× femur length, flange maximum width 0.6× width of tubular part of tibia; hindtibia with two longitudinal carinae on posterior margin; hindcoxa without longitudinal carina on inner medial surface; forewing (Fig. 31) R1 length 0.3× R length; cu-a absent; Rs length 2.6× R length; Cu+M length 0.5× A length; medial vein submedially curved. Color: body brown to reddish brown; wing membrane brown-tinted, paler along vein remnants.

The species is named for the long, nose-like frontal projection (Latin)

Fouts

http://species-id.net/wiki/Loboscelidia_nigra

Loboscelidia nigra

Philippines: Mindanao; only the two syntypes were seen.

As discussed under and , is one of seven species with a triangular frontal projection, complete scrobal sulcus and complete notauli. Dimensions of the antennal articles will separate from these species; the scape is less than 3× as long as broad, flagellomeres I and II are 2.5× as long as broad or longer and flagellomere XI is 4.5× as long as broad.

Kimsey

http://species-id.net/wiki/Loboscelidia_nigricephala

Loboscelidia nigricephala

Australia: Queensland: Mt. Lewis (1 male, CNC); 21 km s Atherton (1 male, QDPI); Hugh Nelson Range, s Atherton (1 male, BME); 3 males were seen, including the holotype.

This is one of five species, including , , and , where the frontal projection is broadly flattened and nearly linear in front view. It can be distinguished from these species by the arched medial vein, cu-a as long as or longer than R, foretibial flange as wide as tubular part of tibia, and the mid and hindtibial flanges 1.5× as wide as tubular part of the tibiae or wider.

Fouts

http://species-id.net/wiki/Loboscelidia_nigricornis

Loboscelidia nigricornis

Philippines: Mindanao I., Agusan, Esperanza Bagugan, Matibog Creek (1 male, BPBM); 6 specimens were seen including the otype series.

This is one of several species with a flat medial vein and rectangular frontal projection. In males, the absence of cu-a and the scape more than 3.3× as long as broad are characteristics shares with . can be distinguished from by Rs less than twice as long as broad (longer in ) and longer pronotum (1.2× as long as broad, versus as long as broad or broader in ).

urn:lsid:zoobank.org:act:B9D54654-B75A-4659-A223-901AFCCDB5BD

http://species-id.net/wiki/Loboscelidia_nitidula

Figures 32 44

Holotype male: Thailand, Petchaburi Prov., Nam Nao NP, 16°43'687"N, 101°33'754"E, 924 m, MT, 5-12/v/2007, N. Hongyothi, T2656 (QSBG). Paratypes (21 males): 1 male: Kaeng Krachan NP, 16/road/stream, 12°48'189"N, 99°26'62"E, MT, 11–18/iii/2009, Sirichai & Prasit, T4685; 1 male: 12°50'177"N, 99°20'688"E, MT, 735 m, 27/xi-4/xii/2008, Sirichai, T4395; 2 males: Chang Mai Prov., Doi Inthanon NP, 7–12/v/1990, E. Fuller, MT; 1 male: Chiangdao NP, Huai Na Lao, 19°24'731"N, 98°55'315"E, 500 m, YPT, 6-7/v/2008, Jugsu & Watwanich, T5808; 1 male: Sakon Nakhon Prov., Phu Phan NP, 14/vii/2006, 17°03'543"N, 103°58'452"E, MT 8-W, Kongnara, T197; 2 males: 17°03'543"N, 103°58'452"E, MT, 15–21/vii/2006, MT, S. Tongboonchai, T200; 3 males: 17°03'488"N, 103°58'497"E, MT, 15–21/vii/2006, MT, S. Tongboonchai,PageBreak T205; 1 male: Nong Bua Prov., Lam Poo Phu Kao Phu, Phan Kham Nat. Pk., 16°49'N, 102°37'E, 208 m, 27/vii-2/viii/2006, MT, R. Singhatip, T85; 1 male: Nakhon Si Prov., Thammarat, Namtok Yong NP, 17°10'434"N, 99°44'508"E, 80 m, MT, 16–23/viii/2008, K. Uprai, T3548; 2 males: Kanchanaburi Prov., Khuean Srinagarindra NP, Huay Mae Kamint, 14°38'312"N, 98°5'643"E, 210 m, MT, 13–20/xi/2008, Somboon & Daorueng, T4424/4423; 1 male: Erawan NP, 100 m, 5/vii/1990, J. Heraty, 90/115; 1 male: Loei Prov., Phu Kradeung NP, 16°49'01"N, 101°47'62"E, 276 m, MT, 14–21/v/2008, T. Phatai, T5011; 1 male: Trang Prov., Nayong, 7 m, 20/ii/2005, 7°33'04"N, 99°49'37"E, MT, D. Lohman; 1 male: Khao Chong Mt. 75 m, 28/iv-2/v/2005, 7°33'38"N, 99°47'369"E, MT; 1 male: Khao Chong Mt. 75 m, x/2005, 7°33'38"N, 99°47'369"E, MT; 1 male: near Nam Tock Ton Prov., Khoa Chong Mt., 140 m, ii/2005, 7°32'15"N, 99°47'36"E, MT, D. Lohman (QSBG, BME, CNC).

Four species have a flattened cervical extension, including brunnea, maculipennis and . Of these four can be distinguished by presence of a scrobal sulcus, a medial metanotal ridge and a large foretibial flange (flange absent in the other species).

Body length 2.0-2.5 mm; forewing length 2.5-3.0 mm. Head: length 1.6× height in side view; eye asetose; frontal projection rectangular in front view; frons smooth, not microstriate; vertex without transverse fovea, posterior expansion convex in profile; frons with ridge extending from vertex along inner eye margin; gena without scale-like setae; scape striate, length 2.7 breadth; flagellomere I length 1.7× breadth; flagellomere II length 1.8× breadth; flagellomere XI length 5× breadth. Mesosoma: pronotal length 0.8× breadth, with fold between dorsal and lateral surfaces, as wide as head width in dorsal view; scutum with notauli reaching posterior margin; scutellum with fine dense striae laterally; metanotum with medial longitudinal striae, impunctate laterally, 0.5× as long as scutellum; mesopleuron with scrobal sulcus; propodeum without transverse dorsal carina; legs (Fig. 44) smooth polished; forefemoral flange 0.7× femur length, flange maximum width 0.8× width of tubular part of femur; foretibial flange 0.9× femur length, flange maximum width as wide as tubular part of tibia; midfemoral flange 0.7× femur length, flange maximum width 0.7× width of tubular part of femur; midtibial flange 0.8× femur length, flange maximum width 1.2× width of tubular part of tibia; hindfemoral flange as long as femur, flange maximum width 1.2× width of tubular part of femur; hindtibial flange as long as femur, flange maximum width 1.6× width of tubular part of tibia; hindtibia with two longitudinal carinae on posterior margin; hindcoxa without longitudinal carina on inner medial surface; forewing (Fig. 32) R1 length 0.7× R length; cu-a length 0.6× R length; Rs length 2.9× R length; Cu+M length 0.6× A length; medial vein submedially curved. Color: body brown to reddish brown; wing membrane brown-tinted, darkest medially, lightest along vein remnants.

The species name, , is Latin for shiny/polished (f.).

Day

http://species-id.net/wiki/Loboscelidia_nixoni

Laccomerista rufescens

Loboscelidia nixoni Day 1978: 29. Replacement name for

Borneo; only the holotype of (Cameron) was seen.

is another of the species characterized by having a curved medial vein, rectangular frontal projection, and no scrobal sulcus, as discussed under . In this group differs from and in having cu-a well-developed and half as long as R. It can be separated from , and by the combination of the scape and flagellomere XI less than twice as long as broad, flagellomeres I and II less than 1.7× as long as broad and hindtibial flange less than 0.7× as wide as tubular part of tibia.

Kimsey

http://species-id.net/wiki/Loboscelidia_novoguineana

Loboscelidia novoguineana

Papua New Guinea: Mt. Suckling (1 male, CNC); Ivimka Res. Station, Lakekamu Basin (3 males, BME); 5 males were seen, including the holotype.

As discussed under , is one of five species with a wide flattened frontal projection. It can be distinguished from these species by the partial notauli, scrobal sulcus indicated by a scrobal pit or several pits, and the fore, mid and hindtibial flanges present and narrower than the respective tibiae. This is the only species described from New Guinea but there surely must be more.

Kimsey

http://species-id.net/wiki/Loboscelidia_ora

Figure 18

Loboscelidia ora

Australia: Queensland: Cape Tribulation (1 male, CNC); Paluma (2 males, CNC, BME); Lacey's Creek, Mission Beach (1 male, CNC); 5 males were examined, including the holotype.PageBreak

can be distinguished from the other species with an apically broad, flattened frontal projection (Fig. 18) by the nearly straight medial vein, cu-a longer than R, Rs more than twice as long as R, A as long or longer than Cu+M, and no notauli.

Maa & Yoshimoto

http://species-id.net/wiki/Loboscelidia_parva

Loboscelidia parva

New Britain; only the holotype was seen.

As discussed under and , is another of the species with a wide, broadly flattened frontal projection. can be distinguished from these species by the arched medial vein, scape 3× as long as broad, partial notauli, pronotum broader than long, and foretibia without a flange.

Kimsey

http://species-id.net/wiki/Loboscelidia_pasohana

Loboscelidia pasohana

Malaysia: Negri Sembilan, Pasho Forest Reserve (1 male, AEI); Sarawak: Gunung Mulu NP (4 males, ROM, BME); Sabah: Mt. Kinabalu (2 males, BMNH); NP, Liwagu Rv. Tr. (1 male, CNC); Thailand: Petchaburi, Kaeng Krachan NP (4 males, QSBC); Chiang Mai: Doi Phahompok NP, Mae Fang Hotspring (1 male, QSBC); 12 specimens were seen including the type series.

This a member of the large group of species with a rectangular frontal projection, submedially curved medial vein, complete scrobal sulcus and complete notauli. can be distinguished from the rest by the following combination of features: Rs nearly as long as R, cu-a half as long as R, flagellomeres I and II less than twice as long as broad, and fore, mid and hindtibial flanges 1.3–1.7× as wide as tubular part of respective tibiae.

urn:lsid:zoobank.org:act:1E7C3500-EE85-481C-AFA6-1DCB27E97A33

http://species-id.net/wiki/Loboscelidia_pecki

Figures 2 14 33

Holotype male: Viet Nam: Tuyen Quang Prov., 360 m, Na Hang Reserve, 16–20 May 1997, FIT, S. B. Peck, 97-10 (CNC).

This species is characterized by the absence of the cu-a vein and having a submedially curved medial vein, characters shared with . It can be distinguished from by Rs 3× as long or longer than R, scape 3× or shorter as long as broad, scrobal sulcus present and the fore and midtibiae without flanges.

Body (Fig. 2) length 2 mm; forewing length 2.5 mm. Head: length 2× height in side view (Fig. 14); eye asetose; frontal projection rectangular in front view; frons smooth, not microstriate; vertex without transverse fovea, posterior expansion strongly curved in profile; frons with ridge extending from vertex along inner eye margin; gena without scale-like setae; scape striate, length 2.9× breadth; flagellomere I length 2.2× breadth; flagellomere II length 2× breadth; flagellomere XI length 4× breadth. Mesosoma: pronotal length 0.9× breadth, with lateral fold separating dorsal from lateral surface, about as wide as head in dorsal view; scutum with notauli reaching posterior margin; scutellum with fine dense striae laterally; metanotum with three medial ridges enclosing roughened medial area, smooth laterally, 0.4-0.5× as long as scutellum; mesopleuron with scrobal sulcus; propodeum without transverse dorsal carina; legs (Fig. 2) smooth, polished; forefemoral flange 0.6 x femur length, flange maximum width 0.8× width of tubular part of femur; foretibial flange 0.8× tibia length, flange maximum width 0.7 x width of tubular part of tibia; midfemoral flange 0.7× femur length, flange maximum width 0.7× width of tubular part of femur; midtibial flange 0.9× femur length, flange maximum width 0.7× width of tubular part of tibia; hindfemoral flange 0.8× femur length, flange maximum width 0.7× width of tubular part of femur; hindtibial flange equal to femur length, flange maximum width 1.3× width of tubular part of tibia; hindtibia with two longitudinal carinae on posterior margin; hindcoxa without longitudinal carina on inner medial surface; forewing (Fig. 33) R1 length as long as R; cu-a length absent; Rs length 3.2× R length; Cu+M length 0.8× A length; medial vein present, submedially curved. Color: Body dark brown; wing membrane brown-tinted, paler along vein remnants, veins brown.

The species is named after the collector, Stuart Peck.

Fouts

http://species-id.net/wiki/Loboscelidia_philippinensis

Loboscelidia philippinensis Fouts, 1922: 623. Syntype males (not females); Philippines: Mindanao, Iligan (USNM).

Philippines: Mindanao (3 males, USNM, BME); the 2 syntypes were also seen.

is one of the group of species characterized by having a submedially curved medial vein, rectangular frontal projection, no scrobal sulcus, and cu-a vein present. It can be distinguished from the rest of the group by the short, broad head in side view (1.2-1.4× as long as high), flagellomere I is more than twice as long as broad and longer than flagellomere II, partial notauli, metanotum half as long or longer than scutellum, A shorter than Cu+M, and hindtibial flange as long as tibia and twice as wide as tubular part of tibiaPageBreak.

Maa & Yoshimoto

http://species-id.net/wiki/Loboscelidia_reducta

Figures 15 34 45

Loboscelidia reducta

Dai Lanh, Nha Trang (1 male, BPBM); Thailand: Loei: Phu Kradueng NP (3 males, QSBC, BME); Phetchabun: Nam Nao NP (4 males, QSBC, PageBreakBME); Prachuab Khiri Khan: Khao Sam Roi Yot NP, Laem Sala Beach (2 males, QSBC); Khonkaen: Nam Pong NP (1 male, QSBC); Sakon Nakon, Phu Phan NP (2 males, QSBC, BME), Mae Hong Son: Namtok Mae Surin NP (3 males, QSBC, BME); Chiang Mai: Huai Nam Dang NP (1 male, BME); Kanchanaburi: Khuean Srinagarindra NP, Tha Thung-na/Chong Kraborg (1 male, QSBC); 22 specimens were seen including the holotype.

is one of the species, including , , and , that have a rectangular frontal projection (Fig. 15), complete notauli, greatly reduced or absent cu-a vein and no medial vein (Fig. 34). It can be distinguished from them by the absence of fore, mid and hindtibial flanges (Fig. 45). This species bears a superficial resemblance to species of .

Fouts

http://species-id.net/wiki/Loboscelidia_rufa

Loboscelidia rufa

Philippines: Misamis Or., Mt. Empagatao (1 male, BPBM); Sibuyan (2 males, USNM); Three specimens were seen including the syntypes.

This is another species in the group with complete notauli, scrobal sulcus and rectangular frontal projection. can be separated from other members of the group by the combination of the hindtibial flange nearly twice as wide as the tubular part of the respective tibiae (shared with ), flagellomeres I and II twice as long as broad or longer, and midtibial flange as long and as wide as the tubular part of the tibia.

Westwood

http://species-id.net/wiki/Loboscelidia_rufescens

Loboscelidia rufescens

Indonesia: Sula Island, Malaysia: Sarawak; only the 2 syntype of Westwood were seen.

is another of the species characterized by having a curved medial vein, rectangular frontal projection and no scrobal sulcus, as discussed under . In this group differs from and in having cu-a well-developed and half as long as R. It can be separated from , and by the combination of the scape and flagellomere XI less than twice as long as broad, flagellomeres I and II less than 1.7× as long as broad and hindtibial flange less than 0.7× as wide as tubular part of tibia.PageBreakPageBreak

Kimsey

http://species-id.net/wiki/Loboscelidia_sarawakensis

Loboscelidia sarawakensis

Malaysia: Sarawak: Gunung Mulu NP (3 males, ROM, BME); Mentawai Range (1 male, ROM); 4th div., Gunung Lulu (1 male, BMNH); 5 males were seen including the holotype.

As discussed under and , is one of seven species with a triangular frontal projection, complete scrobal sulcus and complete notauli. can be separated from other members of the group by the combination of scape less than 2.5× as long as broad, flagellomeres I and II twice as long as broad, flagellomere XI 3.3× as long as broad, metanotum 0.3× as long as scutellum, and fore, mid and hindtibial flanges as long as and at least as wide as tubular part of respective tibiae.

Fouts

http://species-id.net/wiki/Loboscelidia_scutellata

Loboscelidia scutellata

Only the 2 syntypes were seen.

is another of the species with a complete scrobal sulcus and notauli, and a triangular frontal projection. Characteristics that separate this species from the rest include the scape striate and 2.5–2.7× as long as broad, flagellomeres I and II twice as long as broad, flagellomere XI 2.4× as long as broad, fore and midfemoral flanges less than half as long as femora, hindtibial flange as long as tibia and 0.6× as wide as tubular part of tibia.

Kimsey

http://species-id.net/wiki/Loboscelidia_sinensis

Loboscelidia sinensis

Only the holotype was seen.

This is the last of the species group discussed under . can be distinguished from the rest by the short R1 vein (0.2× as long as R), A as long as Cu+M, scape twice as long as broad, flagellomeres I and II 2.5× as long as broad, and metanotum 0.3× as long as scutellum.

urn:lsid:zoobank.org:act:10FCFF3D-8DE8-4511-8671-85B5934C1A1D

http://species-id.net/wiki/Loboscelidia_sisik

Figures 16 35 46

Holotype male: Borneo, W. Kalimantan, Gunung Palung Nat. Pk., 15 June-15 Aug. 1991, Darling, Ubaidillah (Rosichon), Sutrisno, 11S 910131 (MBBJ).

Paratype: 1 male, same data as holotype, 11S 910125 (BME).

This species has a number of distinctive features that in combination will distinguish it from other , including the scale-like setae on the head and legs and the scrobal sulcus reduced to a series of foveae.

Body length 3–4 mm; forewing length 3.5–4.5 mm. Head (Fig. 16): length 1.7× height in side view; eye asetose; frontal projection triangular in front view; frons smooth, not microstriate; vertex without transverse fovea, posterior expansion strongly convex in profile; frons with ridge extending from vertex along inner eye margin; gena without scale-like setae; scape smooth, not striate, length 1.8 breadth; flagellomere I length 1.8× breadth; flagellomere II length 2.8× breadth; flagellomere XI length 3.5× breadth. Mesosoma: pronotal length 0.8× breadth, with lateral carina, as wide as head in dorsal view; scutum with notauli reaching posterior margin; scutellum with scattered large punctures and fine dense striae posteriorly; metanotum medially finely, densely striate/punctate impunctate laterally. 0.3× as long as scutellum; mesopleuron with scrobal sulcus consisting of 3–4 large pits or foveae; propodeum without transverse dorsal carina; legs (Fig. 46) smooth, polished; forefemoral flange 0.7 x femur length, flange maximum width 0.6× width of tubular part of femur; foretibial flange 0.4× tibial length, flange maximum width 0.4 x width of tubular part of tibia; midfemoral flange 0.5× femur length, flange maximum width as wide as tubular part of femur; midtibial flange absent; hindfemoral flange 0.9× femur length, flange maximum width as wide as tubular part of femur; hindtibial flange 0.9× tibial length, flange maximum width 0.6× width of tubular part of tibia; hindtibia with two longitudinal carinae on posterior margin; hindcoxa with longitudinal carina on inner medial surface; forewing (Fig. 35) R1 length 0.5× R length; cu-a length 0.7× R length; Rs length 2.3× R length; Cu+M as long as A; medial vein submedially curved. Color: body dark brown; wing membrane brown-tinted, darker along vein remnants.

Unknown.

The species name is Indonesian for scale, referring to the scale-like setae on the head and legs (noun).

Yao, Liu & Xu

http://species-id.net/wiki/Loboscelidia_striolata

Loboscelidia striolata Yao, Liu & Xu, 2010: 528. Holotype male; China: Guangdong Prov., Nanking National Nature Reserve (SCAC).

None.PageBreak

may very well be part of the species group discussed under and , characterized by a triangular frontal projection, complete scrobal sulcus and complete notauli. However, the published description and images do not show the mesopleuron clearly enough to determine whether the scrobal sulcus is present or not. If it does have a scrobal sulcus then may be synonymous with . Both and share similar head, wing vein, flagellar and leg flange dimensions. They appear to differ in the dimensions of the scape, which 3× as long as broad in and twice as long as broad in and possibly in the presence of the scrobal sulcus in .

Unknown.

1	M vein incomplete medially or absent (as in Figs 24, 26, 28, 35)	2
–	M vein complete	6
2	M vein incomplete medially, Rs twice as long as R (Fig. 28); India	Loboscelidia incompleta sp. n.
–	M vein absent, Rs less than twice as long (as in Fig. 26) or 2.5× as long as R	3
3	Propodeum broadly angulate dorsomedially in posterior view; Borneo	Loboscelidia bakeri Fouts
–	Propodeum flat to gently convex dorsally in posterior view	4
4	Fore, mid and hindtibiae without measurable flanges (Fig. 46); Laos, Viet Nam; Thailand	Loboscelidia reducta Maa & Yoshimoto
–	Fore, mid and hindtibiae with flanges 0.9× as long and 0.3-1.0× as wide as tubular part of respective tibia	5
5	Rs less than 1.5× as long as R, A less than 0.5× as long as Cu+M (Fig. 27); Viet Nam	Loboscelidia fulgens sp. n.
–	Rs more than twice as long as R, A 0.9–1.1× as long as Cu+M; China	Loboscelidia guangxiensis Xu
6	Gena and often legs with scattered scale-like setae (as in Fig. 16)	7
–	Gena and legs without scale-like setae	8
7	M straight medially (Fig. 36); scape less than 3× as long as broad; Borneo	Loboscelidia sisik Kimsey sp. n.
–	M curved submedially; scape more than 3× as long as broad; Viet Nam	Loboscelidia asiana Kimsey
8	Vertex extension flattened in lateral view, not depressed behind ocelli (as in Fig. 11); foretibia without transparent flange, except in Loboscelidia nitidula (as in Fig. 45)	9
–	Vertex extension convex in lateral view, depressed behind ocelli (as in Fig. 4); foretibial flange usually present	12
9	Tibial flanges well-developed (as in Fig. 45); scrobal sulcus present	10
–	Tibial flanges represented by posterior ridge or absent (as in Fig. 42); scrobal sulcus absent	11
10	Rs 3.2–4.0× as long as R; R1 and cu-a shorter than R (Fig. 33); Thailand	Loboscelidia nitidula sp. n.
–	Rs 2.5–3.0× as long as R or shorter; R1 and cu-a as long as R; Taiwan	Loboscelidia latigena Lin
11	Propodeum without transverse subapical carina; cu-a less than 0.3× as long as R; legs smooth, not striate; Borneo, Sumatra	Loboscelidia brunnea Fouts
–	Propodeum with transverse subapical carina; cu-a more than 0.5× as long as R; legs extensively longitudinally striate (Fig. 42); Borneo, Malaysia, Singapore, Sumatra	Loboscelidia maculipennis Fouts
12	M straight medially (as in Fig. 27)	13
–	M curved submedially	18
13	Scutum without notauli (as in Fig. 22)	14
–	Scutum with notauli (as in Figs 21, 23)	15
14	Hindfemoral flange 2.5× as wide as tubular part of femur; hindtibial flange twice as wide as tubular part of tibia; Australia	Loboscelidia maculata Kimsey
–	Hindfemoral flange twice as wide as tubular part of femur; hindtibal flange as wide as tubular part of tibia; Australia	Loboscelidia ora Kimsey
15	Scrobal sulcus present at least as a series of pits or foveae (as in Fig. 2); scape 3.0× as long as broad or shorter; cu-a 0.3× as long or longer than R (as in Fig. 27)	16
–	Scrobal sulcus absent; scape 3.5× as long or longer as broad; cu-a absent	17
16	Face frontal projection rhomboid or rectangular in front view; Rs 3.0× as long as R (Fig. 27); midtibial flange more than half as long and wide as tubular part of tibia (Fig. 39); Thailand, Sumatra	Loboscelidia fulva sp. n.
–	Face frontal projection triangular in front view; Rs 2.5× as long as broad or shorter (Fig. 31); midtibial flange absent or less than half as long and wide as tubular part of tibia (Fig. 43); Borneo	Loboscelidia meifungae sp. n.
17	Rs more than twice as long as R, more than 0.8× as long as M+Cu; Java	Loboscelidia halimunensis Kojima
–	Rs less than twice as long as R, A 0.5–0.7× as long as M+Cu; Philippines	Loboscelidia defecta Kieffer
18	Scutum without notauli or notauli about half as long as scutum (as in Figs 22, 23)	19
–	Scutum with notauli 0.7–1.0× scutal length	21
19	Scutum without notauli; face with frontal projection rhomboid in front view (as in Fig. 20); flagellomeres I-II each less than twice as long as broad; Australia	Loboscelidia australis Kimsey
–	Scutum with notauli about half as long as scutum; face with frontal projection linear to broadly triangular or V-shaped in front view (as in Fig. 19); flagellomeres I-II each twice or more as long as broad	20
20	Foretibia without transparent flange; hindfemoral flange half as wide as femur; Rs more than 3× as long as R; New Britain	Loboscelidia cervix Maa & Yoshimoto
–	Foretibia with transparent flange; hindfemoral flange as wide as femur; Rs less than 3× as long as R; New Britain	Loboscelidia parva Maa & Yoshimoto
21	Frontal projection nearly linear in front view (as in Fig. 18); cu-a as long as R	22
–	Frontal projection rectangular, rhomboid (as in Fig. 20) (extremely elongate in Loboscelidia nasiformis) or triangular (as in Fig. 19); cu-a shorter than R or absent	23
22	Foretibial flange half as wide as tubular part of tibia; midtibial flange half as long and half as wide as tubular part of tibia; New Guinea	Loboscelidia novoguineana Kimsey
–	Foretibial flange as wide as tubular part of tibia; midtibial flange 1.5× as long and as wide as tibia tubular part of; Australia	Loboscelidia nigricephala Kimsey
23	Face with frontal projection elongate and nasiform; head nearly 3× as long as broad (Fig. 13); Thailand	Loboscelidia nasiformis sp. n.
–	Face with frontal projection rectangular to triangular; head twice or less as long as broad	24
24	cu-a less than 0.2× as long as R or absent	25
–	cu-a 0.2–0.4× as long as R	26
25	R1 as long as R, Rs 3× as long as R (Fig. 34); Viet Nam	Loboscelidia pecki sp. n.
–	R1 absent or less than 0.4× as long as R, Rs less than 2.2× as long as R (Fig. 25); Thailand, Borneo, Singapore, Malaya	Loboscelidia cinnamonea sp. n.
26	Midfemoral flange 0.3× as long as femur; R1 less than 0.3× as long as R and A vein as long as Cu+M; China	Loboscelidia sinensis Kimsey
–	Midfemoral flange 0.4-1.0× as long as femur; R1 0.4–1.0× as long as R and A vein shorter than Cu+M, except in Loboscelidia indica	27
27	R1 reaching R at nearly right angle; pronotal length 0.4–0.6× width across posterolateral angles or shorter; China	Loboscelidia levigata Yao, Liu & Xu
–	R1 reaching R at obtuse angle; pronotal length greater than 0.6× width across posterolateral angles	28
28	Scrobal sulcus absent	29
–	Scrobal sulcus present (as in Fig. 2)	31
29	Propodeum with transverse subapical carina; metanotum less than 0.3× as long as scutellum; Borneo, Sula Is	Loboscelidia nixoni Day
–	Propodeum without transverse subapical carina; metanotum more than 0.3× as long as scutellum	30
30	Scape more than 3.0× as long as broad; hindtibial flange wider than tubular part of tibia; Philippines	Loboscelidia philippinensis Fouts
–	Scape less than 3.0× as long as broad; hindtibial flange narrower than tubular part of tibia; Borneo, Sula Is	Loboscelidia rufescens Westwood
31	Frontal projection triangular (as in Fig. 19)	32
–	Frontal projection rhomboid or rectangular (as in Fig. 20)	37
32	Rs more than 3.0× as long as R; flagellomere I less than twice as long as broad; Laos, Sumatra	Loboscelidia laotiana Kimsey
–	Rs 2.5–3.0× or less as long as R; flagellomere I twice or more as long as broad	33
33	Flagellomere XI more than 4.0× as long as broad	34
–	Flagellomere XI 4.0× or less as long as broad	35
34	Scape less than 3× as long as broad; forefemoral flange half as wide as tubular part of femur; hindtibial flange as wide as tubular part of tibia or narrower; Philippines	Loboscelidia nigra Fouts
–	Scape more than 3× as long as broad; forefemoral flange as wide as tubular part of femur; hindtibial flange twice as wide as tubular part of tibia; Sri Lanka	Loboscelidia castanea Krombein
35	Hindtibial flange less than 1.5× as wide as tubular part of tibia; flagellomere XI less than 3× as long as broad; Philippines	Loboscelidia scutellata Fouts
–	Hindtibial flange more than 1.5× as long as wide as tubular part of tibia; flagellomere XI more than 3× as long as broad	36
36	Hindtibial flange 2.0–2.5× as wide as tubular part of tibia (as in Fig. 41); Singapore	Loboscelidia collaris Fouts
–	Hindtibial flange less than twice as wide as tubular part of tibia (as in Fig. 40); Borneo, Sulawesi	Loboscelidia sarawakensis Kimsey
37	Scape 3.9–4.1× as long as broad, flagellomere XI 3.9–4.1× as long as broad; Philippines	Loboscelidia rufa Fouts
–	Scape less than 3.8× as long as broad; flagellomere XI less than 3.8× as long as broad	38
38	Foretibial flange narrower than tubular part of tibia (as in Fig. 40)	39
–	Foretibial flange as wide or wider than tubular part of tibia (as in Fig. 41)	40
39	Rs twice as long as R; scape 3× as long as broad; flagellomere I twice as long as broad; Sri Lanka	Loboscelidia atra Krombein
–	Rs 3× as long as R; scape less than 3× as long as broad; flagellomere I less than twice as long as broad; Viet Nam, Thailand	Loboscelidia laminata sp. n.
40	Fore and midtibial flanges as wide as or narrower than tubular part of respective tibiae; Thailand, Laos, Viet Nam, Malaya, Borneo	Loboscelidia kafae sp. n.
–	Fore and midtibial flanges more than 1.2× as wide as tubular part of respective tibiae	41
41	A longer than Cu-M; Rs less than 3.0× as long as R; pronotum rounded laterally; India	Loboscelidia indica Kimsey
–	A shorter than Cu-M; Rs 3.4× as long as R; pronotum with carinate lateral edge; Borneo, Thailand	Loboscelidia pasohana Kimsey