Liesje Mommer1,2, Monique Weemstra3. 1. Nature Conservation and Plant Ecology, Wageningen University, PO Box 47, 6700 AA Wageningen, the Netherlands. 2. (Author for correspondence: tel +31 317 486944; email liesje.mommer@wur.nl). 3. Forest Ecology and Management, Wageningen University, the Netherlands.
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