Review of the Nearctic genus Lacconotus LeConte (Coleoptera, Mycteridae, Eurypinae).

Lacconotus LeConte, the sole Nearctic representative of the eurypine Mycteridae, is revised, based on morphological features of adults. The following syn. n. is proposed: Lacconotus pallidus Van Dyke, 1928 = Lacconotus pinicola Horn, 1879. The former is a light-colored form with a southern California distribution. A subgen. n.,Alcconotus, is described for Lacconotus pinicola, producing the following comb. n.: Lacconotus (Alcconotus) pinicola (Horn). A lectotype is designated for Lacconotus pinicola. A key separating the two subgenera and species is provided, as are photographs and illustrations of salient structures of adults, and maps showing collection localities. Lacconotus punctatus is newly recorded in Alabama, Arkansas, Massachusetts, Oklahoma, Texas, and Wisconsin; Lacconotus pinicola is newly recorded in Arizona and Utah in the USA, and Baja California Norte in Mexico. Phenology information shows a north-to-south gradation in occurrence time.

Introduction

Among the three subfamilies of Mycteridae, the Eurypinae (= Lacconotinae) are the most diverse with 26 genera and 160 species recognized worldwide, the greatest diversityPageBreak being in New and Old World tropical regions (Pollock 2010). The Nearctic fauna of Mycteridae is not large, but represents all three subfamilies: six species of Clairville (Mycterinae), three species of Latreille (Hemipeplinae), and two species of Eurypinae. The latter are represented by LeConte and Horn (= Van Dyke). The distribution of the family in North America is decidedly either western or eastern, i.e., there are no species represented in the interior of the continent, and no single species bridges this distributional gap (Pollock 2002).

According to Horn (1879: 339), “The Mycteridae seem to have been cast about from place to place by the various students who have had occasion to write about them.” Indeed, the constituents of the ‘modern’ concept of Mycteridae have been placed in multiple families, ranging from Cucujidae (for ) (e.g., LeConte 1854), to Melandryidae (e.g., LeConte and Horn 1883; Van Dyke 1928), and Pythidae (e.g., Seidlitz 1917; Blair 1928). In a phylogenetically based analysis of families related to Mycteridae, Beutel and Friedrich (2005) elucidated the following relationship: (Prostomidae + (Mycteridae + Boridae)); however, they stated that the relationships among the Mycteridae and related families are far from settled.

This study of was undertaken for several reasons. For some time, it was recognized (Pollock, personal observation) that the eastern and western species of the genus were rather dissimilar structurally; recent collections of specimens, especially through Michael Caterino’s “California Beetle Project”, have added much more material for study; and, it is the first author’s goal to revise all genera of world Mycteridae, including the many presently poorly known genera of Eurypinae.

Natural History

As with many other groups of Tenebrionoidea, and Coleoptera generally, relatively little is known of the specific habits of eurypine Mycteridae. Larvae have been described for only a few species; these descriptions (see references in Pollock 2010) indicate that larvae occur under loosened tree bark (e.g., Pic, spp., Abdullah) or in palm leaf axils or dead foliage (e.g. spp.). With the exception of spp., eurypine larvae have flattened bodies and well-developed, complex urogomphal plates, typical of larvae that move subcortically. Lawrence (1991) indicated that mycterid larvae are phytophagous, and that plant-derived material has been found in gut contents of several species.

The biology of seems typical of eurypines: larvae occur under bark of various tree species. Lawrence (1991) illustrated the larva of (Apache Co., Arizona) and indicated that larvae of this species occur under bark of poplar (s) and fir (). Crowson and de Viedma (1964) mentioned a larva, presumed to be that of (= ), from under bark of dead oak ( sp.).

Other specific details pertinent to natural history, derived from label data or other sources, are given under and , below.

Methods and conventions

Standard taxonomic methods were used in this study. Habitus photographs were taken with a Nikon Coolpix 5000® digital camera fitted to a Leica MZ95 stereoscope. Approximately 30 separate photographs were taken for each specimen/structure; these were imported into Combine ZP (Hadley 2010), which stacked and aligned the individual images to create a final photograph completely in focus.

Several measurements were used: HL = length of head from anterior margin of pronotum to labrum; PL = length of pronotum along middle; EL = length of elytron from anterior to posterior extent; GHW = maximum width of head, across eyes; GPW = maximum pronotal width; GEW = maximum width of both elytra; TL = HL + PL + EL.

Label data on type specimens are recorded verbatim, with all label data enclosed in quotes and individual labels separated by a slash (/). Information added by the authors for clarity is enclosed in square brackets ([ ]).

Abbreviations of collections (largely following Evenhuis 2011) consulted and referred to in this study are:

AAAC

Albert A. Allen Collection, Boise, Idaho, USA

AMNH

American Museum of Natural History, New York City, New York, USA

CAS

California Academy of Sciences, San Francisco, California, USA

CMNH

Carnegie Museum of Natural History, Pittsburgh, Pennsylvania, USA

CNC

Canadian National Collection of Insects, Arachnids, and Nematodes, Ottawa, Ontario, Canada

CSCA

California State Collection of Arthropods, Sacramento California, USA

CUIC

Cornell University Insect Collection, Ithaca, New York, USA

DAPC

Darren A. Pollock Collection, Eastern New Mexico University, Portales, New Mexico, USA

EMEC

Essig Museum of Entomology, University of California, Berkeley, California, USA

FMNH

Field Museum of Natural History, Chicago, Illinois, USA

MCZ

Museum of Comparative Zoology, Harvard University, Cambridge, USA

NMNH

National Museum of Natural History, Washington, District of Columbia, USA

NSMC

Nova Scotia Museum, Halifax, Nova Scotia, Canada

QMOR

Collection Entomologique Ouellet-Robert, Université de Montréal, Montréal, Québec, Canada

RBC

Rick Buss Collection, Albuquerque, New Mexico, USA

SBMN

Santa Barbara Museum of Natural History, Santa Barbara, California, USA

TAMU

Texas A & M University, College Station, Texas, USA

UAIC

University of Arizona Insect Collection, Tucson, Arizona, USA

UBC

University of British Columbia, Vancouver, British Columbia, Canada

UCR

University of California-Riverside, Riverside, California, USA

UNHC

University of New Hampshire, Durham, New Hampshire, USA

WFBM

W.F. Barr Entomological Collection, University of Idaho, Moscow, Idaho, USA

WIRC

WisconsinInsect Research Collection, University of Wisconsin, Madison, Wisconsin, USA

Identification

Adults of can be differentiated with the following key:

LeConte

http://species-id.net/wiki/Lacconotus

Lacconotus

Description.

[note: “” indicates the character states for subgenus , while “” refers to the corresponding states in subgenus (see below)].

Body elongate oval (TL/GEW 2.8-3.4), parallel-sided to widened posterior of middle, slightly () to moderately () flattened dorsally. TL 4.6–7.5 mm.

Head relatively short, narrowed slightly posterior of eyes; eyes moderately large, distinctly convex, inner margins convergent anteriorly; facets moderately coarse, withPageBreak intrafacetal setae (especially noticeable in ); frontoclypeal suture indistinct, indicated by slight furrow only; antennal insertions slightly concealed dorsally by raised lateral margins of frons; labrum transverse, anterior margin straight to shallowly sinuate.

Antennae relatively short (Fig. 1) () to moderately elongate (Figs 2–3) (), not exhibiting distinct sexual dimorphism; scape and pedicel moniliform () to slightly elongate (); antennomere 3 elongate, antennomeres 4–10 wider than long, submoniliform () to elongate, triangular to subserrate (); antennomere 11 narrowed distally; sensilla present on distal surface of antennomeres 5–10.

Figure 1.

Dorsal habitus photograph of ; female, New Hampshire. TL = 5.1 mm. Photo credit: Darren Pollock, Eastern New Mexico University.

Figure 2.

Dorsal habitus photograph of ; female, Utah. TL = 6.3 mm.
Photo credit: Darren Pollock, Eastern New Mexico University.

Figure 3.

Dorsal habitus photograph of (‘pallidus'); female, California. TL = 6.5 mm. Photo credit: Darren Pollock, Eastern New Mexico University.

Mandibles relatively short, stout, slightly asymmetrical, apically bidentate; terebral teeth absent, or represented by several minute crenulae; molae approximately equal in size, subquadrate, with slightly developed surface texture; both mandibles with abrupt incision distal of mola; ventral row of microtrichia absent; prostheca distinct, about half length of mandible, inserted near distal edge of mola; maxilla with galea slightly longer than lacinia; galea bluntly rounded distally, relatively densely pubescent; maxillary palpi elongate, apical palpomere securiform () to slightly cultriform (); inner margins of palpomeres 1 and 2 straight () to slightly sinuate (); mentum short, about 2 × wider than long, posterior suture straight () or distinctly arcuate ().

Thorax. Prothorax subquadrate (Figs 1–3), slightly wider than long (GPW/PL = 0.90–1.29); pronotal margins straight and convergent anteriorly, to slightly arcuate and widest near midlength; pronotal disc flat to slightly convex, with variously developed shallow, paired depressions; lateral pronotal carinae absent, margin smooth; posterior margin with pair of small, deep, punctiform pits; prosternum anterior of procoxae short () to moderately elongate (), surface flat to slightly PageBreaksunken medially; intercoxal process short, knife-like, extended to about half length of procoxae; procoxae rounded () to elongate (); procoxal cavities open externally and internally; protrochantin concealed.

Elytra elongate, subovate, disc flat () to slightly convex (), upper surface uniformly and moderately coarsely punctate and setose (slightly more coarse in ), setae closely appressed to elytral surface; apical elytral patch present, but not conspicuous dorsally, not contrasting in color with respect to remainder of elytron; epipleuron narrow, traceable only to abdominal ventrite 3 or 4; mesosternum with posterior intercoxal process parallel-sided, extended posteriorly to near posteriorPageBreak extent of mesocoxae; mesocoxae narrowly but completely separated, trochantins exposed; coxal cavities partly closed laterally by mesepimera; metasternum large, convex, anterior margin with indistinct () to distinct () process, in contact with posterior mesosternal process; median impressed line distinct to at least half distance to anterior margin of metasternum; metendosternite with long, relatively wide stalk; anterior tendons inserted on anterior margin of metendosternite body; laminae large, produced and somewhat angulate laterally.

Wing (Figs 7–8) fully developed, membrane beyond distinct radial cell moderately short (esp. in ); venation similar in both species, but wing membrane and veins relatively darkly pigmented in (Fig. 8), very pale in (Fig. 7); wedge cell present; 3 MP veins reaching wing margin, proximal to CuA+AA; pigmented patches (flecks) present near junction of RP and MP, and near radial cell (), indistinct in .

Figures 7–8.

Wing of species of 7 , wing length = 4.5 mm 8 , wing length = 5.1 mm. Photo credit: Darren Pollock, Eastern New Mexico University.

Legs well developed, similar in relative shape and size on all thoracic segments; middle and hind femora slightly more expanded than front femora; femora relatively slender, but distinctly widened toward midlength; tibiae straight, about same length PageBreakPageBreakas femora, tibial spurs very short, equal in length; tarsomeres slender, 5–5-4; all tarsomeres simple ventrally, except for penultimate tarsomere with large ventral lobe; basal tarsomere on hind tarsus subequal in length to other tarsomeres combined; tarsal claws with large blunt tooth.

Abdomen with all ventrites freely articulated, uniformly punctate and setose, except for male sex patch; sex patch of two forms: small, longitudinally oval, setose patch on ventrite 2, not contrasting in color with ventrite (Fig. 6) (); or large, somewhat bulbous, glabrous area occupying and somewhat distorting the shape of ventrite 2, distinctly contrasting in color to dark ventrite surface (Fig. 5) ().

Figures 5–6.

Male sex patch of species of 5 , scale bar = 0.75 mm; 6 , scale bar = 0.25 mm. Photo credit: Darren Pollock, Eastern New Mexico University.

Male genitalia with median lobe dorsal to tegmen; sternite 9 forming ring-like sclerite, U-shaped in (Fig. 11), Y-shaped in (Fig. 9); tegmen relatively short, stout; basale broader than long, proximal margin deeply emarginate; length of apicale subequal to that of basale (along lateral margins); parameres of apicale short (Fig. 11) () to slightly elongated (Fig. 9) (), with distal, inwardly-directed hook; median lobe (Figs 10, 12) stout, longer than tegmen; ventral side deeply emarginate, dorsal side proximally subquadrate, laterally produced, explanate; apex of median lobe triangular, relatively blunt.

Figures 11–12.

Male genitalia of 11 tegmen 12 median lobe. Scale bar = 0.25 mm. Photo credit: Darren Pollock, Eastern New Mexico University.

Figures 9–10.

Male genitalia of 9 tegmen 10 median lobe. Scale bar = 0.25 mm. Photo credit: Darren Pollock, Eastern New Mexico University.

Female genitalia with elongate, flexible, and only slightly sclerotized ovipositor; coxites 4-segmented, sparsely setose; distal segment short, distinctly more sclerotized than remainder of coxite; styli short, setose, with several very long distal setae; dorsal and ventral baculi well developed, extended to base of coxites; spiculum long, far exceeding length of segment 8; bursa copulatrix small (Fig. 13) () to very large (Fig. 14) (), separated from vagina by narrow constriction, without conspicuous surface texture; spermatheca present, inserted near or at base of bursa, with elongate spermathecal gland.

Figures 13–14.

Internal female genitalia of 13 14 . Scale bar = 0.25 mm. Photo credit: Darren Pollock, Eastern New Mexico University.

LeConte

http://species-id.net/wiki/Lacconotus_punctatus

Figs 1 4 –5 7 9–10 13 15 –16

Figure 4.

Forebody of , dark form. Photo credit: Darren Pollock, Eastern New Mexico University.

Figure 15.

Distribution of and in North America and Mexico.

Figure 16.

Phenology of and in North America and Mexico.

Lacconotus punctatus

Type.

holotype, male, labeled: “[pink circle] / ♂ / Type 4760 / Lacconotus punctatus Lec. / HOLOTYPE ♂ Lacconotus punctatus LeC. exam. Pollock 2000”, in MCZC.

Diagnosis.

This species is easily diagnosed by the following characteristics: body color dark piceous to near black, pronotum with reddish margins and black center (Figs 1, 4); antennae relatively short, antennomeres submoniliform; male sex patch onPageBreak ventrite 2 bulging, glabrous, yellow-orange, contrasting with dark color of venter (Fig. 5); distribution in eastern North America (Fig. 15).

Re-description.

To general features of (see description, above) the following can be added: TL 4.4–5.8 mm; GEW 1.5–2.0 mm; TL/GEW 2.9–3.3. Dorsal body surface uniformly piceous to near black, except for lateral areas of pronotum red-orange (Fig. 1); extent of light area varying, from extreme posterolateral corners to fully 2/3 of pronotal disc; ventral surface and legs dark, piceous to near black; antennomeres 5–10 short, distinctly wider than long, submoniliform; antennal sensilla completely annular, covering entire distal antennal surface, around insertion point of next antennomere; wing very pale, veins present, but inconspicuous; male sex patch (Fig. 5) very large, occupying entire length of second ventrite, prolonged onto ventrite one, glabrous and bulging ventrally; color of sex patch yellow-orange, distinctly contrasting background color of ventrite; tegmen of male genitalia (Fig. 9) moderately elongate, parameres of apicale relatively slender; bursa copulatrix (Fig. 13) spherical, small.

is newly recorded in Alabama, Arkansas, Massachusetts, Oklahoma, Texas, and Wisconsin (see Appendix A). Published records of PageBreakare from Ontario (Campbell 1991)1, Québec (Campbell 1991)2, and Nova Scotia (Majka and Selig 2006) in Canada [Horn (1879) first reported it from “Canada”], and Georgia (Ulyshen et al. 2010), Michigan (Hubbard et al. 1878; Horn 1879), New Hampshire (Chandler 2001), Ohio (Dury 1902; Blatchley 1910), and Pennsylvania (LeConte 1862) in the United States.

Although listed from Ontario in Campbell (1991), we have not been able to find any published record, or any vouchers specimen in any North American collection that would substantiate this report. Consequently, pending verification of its occurrence in this jurisdiction, we remove Ontario from the known distribution of this species.

In addition to the specimen from Montreal in the CUIC (Appendix 1) a second specimen from Québec was formerly in the Ouellet-Robert collection of the Université de Montréal, however, the specimen was stolen before being databased so its collection date and locality in the province are unknown (pers. com., Louise Cloutier)

The new records above make clear that the distribution of in North America is much wider than previously known (Fig. 15). Less than a decade ago, Pollock (2002) reported the species from only four jurisdictions in North America (Québec, Pennsylvania, Ohio, and Michigan). There are now records from 14 states and provinces on the continent. The records from Wisconsin constitute a northwestern range extension of 650 km; and those from Oklahoma a southwestern range extension of 1,000 km. The present distribution indicates that is found over much of eastern North America, from a latitude of 33.6° to 44.3°N, and between longitudes of 64.5° and 95.3°W, much of the continent west of the prairies.

There is also much more information on the range of habitats that occupies. In Nova Scotia a specimen was found in a mixed forest of white pine ( L.), balsam fir ( (L.) Mill), eastern hemlock ( (L.) Carr.), and maple ( spp.) (Majka and Selig 2006). In New Hampshire, W.J. Morse and D.S. Chandler collected 26 specimens at a water tower in a mixed hardwood forest with eastern hemlocks. In Oklahoma a specimen was collected on a dead oak and in Wisconsin a specimen was found in an oak savanna. In Arkansas a specimen was found in a mixed forest/old field.

In Georgia, specimens were collected in mature bottomland hardwood forests in April with flight intercept traps in the forest canopy (Ulyshen et al. 2010). Dominant trees included box elder ( L.), oak ( spp.), ash ( spp.), eastern cottonwood ( (Bartr.) ex. Marsh.), and sweetgum ( L.) with some loblolly pine () (M. Ulyshen, pers. comm.). Five of six specimens were found 15 m above the forest floor (Ulyshen et al. 2010). Ulyshen et al. (2010) proposed that may be an early-seasonal canopy specialist, a reason why it has been so infrequently collected.

The phenology information that is available (Fig. 16) indicates that adults can be found between 31 March and 16 June. Specimens from southern areas (i.e., Georgia) were found in mid April (Ulyshen et al. 2010), whereas those from northern latitudes (i.e., New Hampshire) occur mainly during the last two weeks of May and first weekPageBreak of June (D.S. Chandler, pers. comm.), indicating a north-to-south gradation in occurrence period.

Pollock & Majka subgen. n.

urn:lsid:zoobank.org:act:7F51E3D0-3408-4469-BC83-EFD22E47750F

http://species-id.net/wiki/Alcconotus

Type species.

Horn, by present designation.

Derivation of name.

a partial anagram of , in which its species was formerly placed.

Taxonomic notes.

Pollock (2002) stated that it might be necessary to establish a new genus for the two western species of , although no details were given to justify this possibility. There are many differences between the eastern and western PageBreakspecies of (see description above, for ), but the most significant reason for proposal of this new subgeneric name is the structure of the male sex patch, which differs greatly between and . It could be argued that this might justify separation into two genera; however, within the related genus Clairville (Mycterinae) there are also significant differences in this structure. Also, there are significant differences in the structure of the bursa copulatrix and spermatheca between the two groups (see Figs 13–14); it is impossible to compare these intrageneric differences with other eurypine or mycterid taxa, since the internal female genitalia have yet to be studied in detail in most groups.

See description above, for ; characteristics unique to are indicated with the alternatives for (s. str.).

Horn comb. n.

http://species-id.net/wiki/Lacconotus_pinicola

Figs 2 –3 6 8 11–12 14 15 –16

Lacconotus pinicola

Lacconotus pallidus

Types.

(, all in MCZC).—lectotype (here designated), female, labeled: “Veta Pass 27.6 Col / 592 / [red] Type 7976 / [handwritten] Lacconotus pinicola (Schwz) / J.L. LeConte Collection / LECTOTYPE ♀ Horn 1879; design. D.A. Pollock 1994”. paralectotype. female, labeled: “Col / [blue] Para-Type 8047. / G.H. Horn Collection”, in MCZC.

(, all in CAS).—holotype, male (CAS type # 2585), labeled “Mt. Wilson Cal. 6.13.3 / 7701 / Van Dyke Collection / Holotype [along left margin of label covered in red ink] ♂ Lacconotus pallidus Van Dyke". ALLOTYPE, female, labeled "Carmel, Monterey Co VI-4-1916 Cal. / Van Dyke Collection / Allotype [along left margin of label covered in red ink] ♀ Lacconotus pallidus Van Dyke". Four PARATYPES. Male, labeled "Carmel, Monterey Co VI-4-1916 Cal. / Van Dyke Collection / Paratype [along left margin of label] ♂ Lacconotus pallidus Van Dyke". Male, labeled "ParaisoSpgsCal V.28 1924 L.S. Slevin / L.S. Slevin Collection / Paratype [along left margin of label] Lacconotus pallidus Van Dyke". Female, labeled "ParaisoSprings V.31 1916 Cal. / L.S. Slevin Collection / Paratype [along left margin of label] Lacconotus pallidusPageBreak Van Dyke". Female, labeled "Paraiso Springs V.29 1916 Cal. / CHAMISAL / L.S. Slevin Collection / Paratype [along left margin of label] Lacconotus pallidus Van Dyke".

may be distinguished from by the following features: body color ranging from testaceous to dark brown, uniform dorsally (Figs 2–3); antennae relatively long, subserrate; male sex patch on ventrite 2 small, oval, densely pubescent (Fig. 6); distribution in western North America (Fig. 15).

(see Horn 1879 and Van Dyke 1928) – With general features of subgenus (as described above) with the following: TL 4.8–7.5 mm; GEW 1.5–2.3 mm; TL/GEW 2.8–3.4. Dorsal body surface uniformly testaceous to dark brown or piceous (Figs 2–3), without any color contrast; antennomeres 5–10 relatively elongate, subserrate; antennal sensilla not completely surrounding opening of antennomere, restricted to triangular side of antennomeres; wing membrane distinctly pigmented, veins very conspicuous (Fig. 8); male sex patch longitudinally oval, occupying about 2/3 length of ventrite 2, densely covered with short setae (Fig. 6), not bulbous or contrasting in color; tegmen of male genitalia (Fig. 11) short, parameres stout; bursa copulatrix (Fig. 14) very large, spherical.

Notes.

Van Dyke (1928) established (as distinct from ) based on its lighter color, relatively narrower pronotum, shorter relative length of the elytra, and deeper punctation. However, upon examination of the type series and other specimens, we have determined that the only feature of Van Dyke’s that withstands scrutiny is the habitus color. As well, more detailed examination has revealed that the male and female genitalia and male sex patch are virtually identical between and . One feature, mentioned by Van Dyke (1928) that does seem noteworthy is the somewhat restricted distribution of in southern California. We herein consider a pale “form” of .

is newly recorded from Arizona, Utah, and Baja California Norte in Mexico (see Appendix A). Published records of are from British Columbia (Hatch 1965), California (Fall 1901; Van Dyke 1928), Colorado (Horn 1879; Cockerell 1893; Van Dyke 1928), western Nevada (Horn 1879), and New Mexico (Snow 1882, 1906; Knaus 1907). The range of the species (Fig. 15) shows it to be widely distributed in the southwestern United States (Arizona, California, Colorado, Nevada, New Mexico, and Utah) extending south to Baja California Norte in Mexico, and in southeastern British Columbia. Specimens should be sought in intervening areas in Idaho, Oregon, and Washington to ascertain if these populations are actually disjunct.

A number of specimens examined were found on ponderosa pine ( Douglas ex. C. Lawson), including one specimen which was recorded as emerging from a dead branch. It has also been found on Engelmann spruce ( Parry ex Engelm.), scrub oak ( Greene), and cherry ( sp.). Specimens have been collected at UV lights, with malaise and flight-intercept traps, and by beating vegetation. Fall (1901: 177) wrote “...rare during May and June; found always on oaks, notwithstanding its name.” The larva of has been illustrated by Lawrence (1991), but not described in detail. The phenology information that is available (Fig. 16) indicates that adults can be found between 13 May and 29 August with two specimens having been found in the autumn (6 October and 6 November). The peak in adult numbers appears to be in the first half of July.

1	Dorsal color dark brown to near black, except for variously developed lateral reddish areas on pronotum (Fig. 1) (in some specimens reduced to the extreme posterolateral corners, e.g., Fig. 4); elytral punctation coarse, punctures obvious; antennae short, antennomeres submoniliform; male sex patch large, bulbous, glabrous, and with contrasting pink-red color (Fig. 5); known from eastern North America, west to Texas (Fig. 15)	Lacconotus (Lacconotus) punctatus LeConte
–	Dorsal color uniform, light brown (Fig. 3) to nearly black (Fig. 2); punctation of elytra fine, punctures not conspicuous; antennae relatively long, antennomeres longer than wide; male sex patch small, oval, setose, and not distinctly contrasting in color to venter (Fig. 6); known from western North America, east to Colorado and New Mexico (Fig. 15)	Lacconotus (Alcconotus) pinicola Horn