Zachary Kayler1, Arthur Gessler1,2, Nina Buchmann3,4. 1. Institute for Landscape Biogeochemistry, Leibniz-Centre for Agricultural Landscape Research (ZALF), Eberswalderstr. 84, 15374 Müncheberg, Germany. 2. Professorship for Landscape Biogeochemistry, Humboldt-University at Berlin, Lentze-Allee 75, 14195 Berlin, Germany. 3. Institute of Plant, Animal and Agroecosystem Sciences, LFW C56, ETH Zurich, Universitaetsstrasse 2, 8092 Zurich, Switzerland. 4. (Author for correspondence: tel +41 44 632 39 59; email nina.buchmann@ipw.agrl.ethz.ch).