Multiscale control of flooding and riparian-forest composition in Lower Michigan, USA.

Despite general agreement that river-valley hydrology shapes riparian ecosystems, relevant processes are difficult to distinguish and often inadequately specified in riparian studies. We hypothesize that physical constraints imposed by broad-scale watershed characteristics and river valleys modify local site conditions in a predictable and probabilistic fashion. To test this hypothesis, we employ a series of structural equations that decompose occurrence of riparian ecotypes into regional temperature, catchment storm response, valley hydraulics, and local site wetness via a priori specification of factor structure and ask (1) Is there evidence for multiscale hydrologic control of riparian diversity across Lower Michigan? (2) Do representations of key constraints on flood dynamics distinguish regional patterns of riparian vegetation? (3) How important are these effects? Cross-correlation among geospatial predictors initially obscured much of the variation revealed through analysis of semipartial variance. Causal relationships implied by our model fit with observed variation in riparian conditions (chi-square P = 0.43) and accounted for between 84% and 99% of the occurrence probability of five riparian ecotypes at 94 locations. Results suggest strong variation in the effects of regional climate, and both the relative importance and spatial scale of hydrologic factors influencing riparian vegetation through explicit quantification of relative flood frequency, duration, intensity, and relative overall inundation. Although climate and hydrology are not the only determinants of riparian conditions, interactions of hydrologic sourcing and flood dynamics described by our spatial models drive a significant portion of the variation in riparian ecosystem character throughout Lower Michigan, USA.